MD09G1109100.v1.1

Binds to the signal sequence of presecretory protein when they emerge from the ribosomes and transfers them to TRAM (translocating chain-associating membrane protein)

Design Primers CRISPR Design

Basic Information

Type: gene
Biological Identity
malus_domestica
Chr09
OG0024313
Physical Location & Seq
Forward (+)
8074851 .. 8076311
1461 bp
Loading structure...
UTR
Exon/CDS
Intron
MD09G1109100.v1.1.491

Sequence Viewer

Length: 1461 bp
ATGGTGTTGGCGGAATTAGGAGGGAGCATATCCCGTGCCCTCCAGCAGATGATCAATGCGACGGTGGTCGACGAGAAGGTCCTGAACGACTGCCTCAACGAAATCAGTCGCGCTCTGCTACAGTCCGACGTCCAATTCAGTCTCGTTCGCGACATGCAGATCAACGTTAAAAAGAATGTCAATCTGTCCGATCTCGCTGCCGGTCACAACAAGCGCAAGATCATTCAGCAAGCCGTGTTCGATCAACTCTGCCAAATGCTGCAGGATCCATCGGGGAAACCCGATCAGCTTGTGCCGAAAAAAGGGAAGAAACCGAGCCAAGTTGTAATGTTTGTTGGTTTACAAGGGTCTGGAAAAACCACAACGTGTACAAAATACGCGTATTATCATAAGAACAAAGGCTGGAAACCTGCCTTAGTTTGTGCAGACACATTCCGAGCTGGCGCTTTCGATCAGCTCAAGCAAAACGCCGCCAAGGCTAAGGTTCCGTTTTATGGAAGTTACACAGAGACGGACCCTGTGAAAATTGCGGTCGAAGGTGTGGAGAGGTTAAAAAACGATCAGAGCAATAGTTTTGATCTCATAATTGTTGATACTAGTGGGAGGCACAAGCAAGAAGCTGCTCTTTCCGAAGAGATGCGTCAACTTTCCGAAGCAATCAGTCCCGACCTTGTTGTGTTTGTTATGGATGCAAGTATCGGACAATCTGCATTTCATCAAGCTCAAGCGTTTAAGCAAAGCGTTGCGGTTGGAGCTGTTATTGTAACGAAAATGGACGGTCACGCAAAGGGCGGCGGTACACTCAGTGCTGTTGCAGCGACAAAGAGTCCTGTTATATTTATCGGAACAGGAGAGCACACGGATCAGTTGGAAGCCTTCGATGTGAAACCCTTTGTCCGGCGTCTGTTAGGCATGGGGGACTTGCCTGGACTTGCGAACAAGATTCGCGAAGCTGTTCCGATGGATCAACACGAAGACATATTGGAAAATATTTTACAGGGCGGATTTACGATAAGGACCATGCAAGTGTTGTTTCAGACCAAGCTAAAACTCGGCACACCTAACCAGCTCGCTTCGATGCTCCCTGCAGAATGTACGAGCCGGTTATTTCCTGCAGGACTCGGCGATGATCAGGAAAATCAGAACCAGATGAAGCAGTTCTTGGTCATGATGGATTCAATGACTGCTCAAGAGTTGGACAGCCCGAACCTAAAGGTAAGCGAGTTGCGGAGACAACGGATAGCAAGGGGTTCTGGTCATGAAGTTAGACAGGTAGAAGAGCTGATTGAAGCGTGCAAAGCTGTTAGCAAGAATTTTAGCAAGGGGAGAGCGCTGCTCCGGAGTTTGAACAGCGGCGACATGCGAGCGGTAACCCGGAATATGAGTGGTAATCCGCAGCTGCTCAACCAGGTTGGGGGCATGGGTGGTTTGAGAAGCTTGATGAATCTGGGGAGGTTATAA
Functional Annotation
Gene Ontology
Molecular Function
Biological Process
Cellular Component
GO:0005575 GO:0005622 GO:0005623 GO:0005737 GO:0044424 GO:0044464
KEGG Pathways
Metabolic & Signaling
ko02024 ko03060 ko03070 map02024 map03060 map03070
Pfam Domains
Protein Families
SRP54 SRP54_N SRP_SPB

Protein Analysis

487

Amino Acids

53.13

Weight (kDa)

9.04

Isoelectric Point (pI)

47.09

Instability Index
Protein Domains (Pfam)
Domain Name Pfam ID Position E-value Description
SRP54_N PF02881 6 - 83 4.5e-12 SRP54-type protein, helical bundle domain
SRP54 PF00448 107 - 304 2.4e-73 SRP54-type protein, GTPase domain
cobW PF02492 108 - 261 4.3e-07 CobW/HypB/UreG, nucleotide-binding domain
SRP_SPB PF02978 336 - 437 1.5e-20 Signal peptide binding domain
Hydrophobicity Profile (Kyte-Doolittle)
AI Structure Prediction Report
Calculating structure properties...

Genomic Context

Gene Family Tree

Species Filter
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Publication-ready
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Orthologous Genes (Group: OG0024313)

Species Orthologous Gene IDs
malus_domestica MD09G1109100.v1.1 MD17G1097400.v1.1
pyrus_communis pycom09g03160

Restriction Enzyme Sites

1 / 10
Enzyme Recognition Site Cut Count Positions (bp)
AanI TTATAA 1 cut(s) 1459
AasI GACNNNNNNGTC 1 cut(s) 77
AatII GACGTC 1 cut(s) 132
Acc36I ACCTGC 1 cut(s) 418
AccBSI CCGCTC 1 cut(s) 1367
AccI GTMKAC 1 cut(s) 69
AccII CGCG 4 cut(s) 111, 150, 380, 948
AccIII TCCGGA 1 cut(s) 1338
AclI AACGTT 1 cut(s) 165
AclWI GGATC 4 cut(s) 260, 273, 870, 972
AcsI RAATTY 1 cut(s) 1312
AcyI GRCGYC 2 cut(s) 129, 901
AdeI CACNNNGTG 2 cut(s) 366, 806
AfaI GTAC 3 cut(s) 370, 799, 1096
AfeI AGCGCT 1 cut(s) 1332
AfiI CCNNNNNNNGG 4 cut(s) 302, 494, 897, 1414
AflIII ACRYGT 2 cut(s) 365, 378
AgsI TTSAA 3 cut(s) 1179, 1289, 1348
AhdI GACNNNNNGTC 1 cut(s) 825
AhlI ACTAGT 1 cut(s) 596
AjnI CCWGG 2 cut(s) 925, 1407
AjuI GAANNNNNNNTTGG 2 cut(s) 1145, 1177
Alw21I GWGCWC 1 cut(s) 858
Alw26I GTCTC 3 cut(s) 146, 503, 1225
AlwI GGATC 4 cut(s) 260, 273, 870, 972
Aor13HI TCCGGA 1 cut(s) 1338
Aor51HI AGCGCT 1 cut(s) 1332
ApeKI GCWGC 7 cut(s) 197, 259, 620, 815, 1333, 1396, 1399
ApoI RAATTY 1 cut(s) 1312
Asp700I GAANNNNTTC 2 cut(s) 954, 1157
AspLEI GCGC 4 cut(s) 113, 216, 446, 1333
AspS9I GGNCC 3 cut(s) 79, 514, 1017
AsuC2I CCSGG 1 cut(s) 1375
AvaII GGWCC 3 cut(s) 79, 514, 1017
BaeGI GKGCMC 1 cut(s) 40
BamHI GGATCC 1 cut(s) 265
BbsI GAAGAC 1 cut(s) 981
Bbv12I GWGCWC 1 cut(s) 858
BbvI GCAGC 7 cut(s) 184, 246, 607, 827, 1320, 1386, 1408
BccI CCATC 3 cut(s) 277, 955, 1165
BceAI ACGGC 1 cut(s) 218
BcgI CGANNNNNNTGC 2 cut(s) 179, 213
BciT130I CCWGG 2 cut(s) 927, 1409
BclI TGATCA 2 cut(s) 51, 1129
BcnI CCSGG 1 cut(s) 1375
BcoDI GTCTC 3 cut(s) 146, 503, 1225
BcuI ACTAGT 1 cut(s) 596
BfaI CTAG 1 cut(s) 597
BfmI CTRYAG 4 cut(s) 119, 260, 1086, 1113
BfoI RGCGCY 2 cut(s) 447, 1334
BfuAI ACCTGC 1 cut(s) 418
BglI GCCNNNNNGGC 1 cut(s) 476
Bme1390I CCNGG 3 cut(s) 927, 1375, 1409
Bme18I GGWCC 3 cut(s) 79, 514, 1017
BmeRI GACNNNNNGTC 1 cut(s) 825
BmgT120I GGNCC 3 cut(s) 79, 514, 1017
BmiI GGNNCC 3 cut(s) 267, 486, 516
BmrFI CCNGG 3 cut(s) 927, 1375, 1409
BmsI GCATC 3 cut(s) 627, 679, 1068
BoxI GACNNNNGTC 1 cut(s) 65
BpiI GAAGAC 1 cut(s) 981
BplI GAGNNNNNCTC 2 cut(s) 1320, 1352
BpmI CTGGAG 1 cut(s) 26
Bpu10I CCTNAGC 1 cut(s) 480
BpuEI CTTGAG 3 cut(s) 443, 708, 1173
BpuMI CCSGG 1 cut(s) 1375
BsaHI GRCGYC 2 cut(s) 129, 901
BsaJI CCNNGG 1 cut(s) 474
BsaWI WCCGGW 1 cut(s) 1338
Bsc4I CCNNNNNNNGG 4 cut(s) 302, 494, 897, 1414
Bse118I RCCGGY 2 cut(s) 200, 1101
BseAI TCCGGA 1 cut(s) 1338
BseBI CCWGG 2 cut(s) 927, 1409
BseDI CCNNGG 1 cut(s) 474
BseGI GGATG 1 cut(s) 694
BseLI CCNNNNNNNGG 4 cut(s) 302, 494, 897, 1414
BseMII CTCAG 1 cut(s) 817
BseSI GKGCMC 1 cut(s) 40
BseXI GCAGC 7 cut(s) 184, 246, 607, 827, 1320, 1386, 1408
BsgI GTGCAG 1 cut(s) 444
Bsh1236I CGCG 4 cut(s) 111, 150, 380, 948
Bsh1285I CGRYCG 1 cut(s) 534
BsiEI CGRYCG 1 cut(s) 534
BsiHKAI GWGCWC 1 cut(s) 858
BsiSI CCGG 5 cut(s) 201, 898, 1102, 1339, 1375
BslFI GGGAC 2 cut(s) 648, 932
BslI CCNNNNNNNGG 4 cut(s) 302, 494, 897, 1414
BsmAI GTCTC 3 cut(s) 146, 503, 1225
BsmBI CGTCTC 1 cut(s) 503
BsmFI GGGAC 2 cut(s) 648, 932
Bsp1286I GDGCHC 2 cut(s) 40, 858
Bsp13I TCCGGA 1 cut(s) 1338
Bsp1407I TGTACA 1 cut(s) 368
Bsp68I TCGCGA 2 cut(s) 150, 948
BspCNI CTCAG 1 cut(s) 816
BspEI TCCGGA 1 cut(s) 1338
BspFNI CGCG 4 cut(s) 111, 150, 380, 948
BspHI TCATGA 2 cut(s) 1167, 1258
BspLI GGNNCC 3 cut(s) 267, 486, 516
BspMAI CTGCAG 3 cut(s) 264, 1090, 1117
BspMI ACCTGC 1 cut(s) 418
BspPI GGATC 4 cut(s) 260, 273, 870, 972
BspQI GCTCTTC 1 cut(s) 1272
BsrBI CCGCTC 1 cut(s) 1367
BsrFI RCCGGY 2 cut(s) 200, 1101
BsrGI TGTACA 1 cut(s) 368
BssAI RCCGGY 2 cut(s) 200, 1101
BssECI CCNNGG 1 cut(s) 474
BssNI GRCGYC 2 cut(s) 129, 901
BssT1I CCWWGG 1 cut(s) 474
Bst2UI CCWGG 2 cut(s) 927, 1409
Bst4CI ACNGT 3 cut(s) 64, 123, 779
Bst6I CTCTTC 2 cut(s) 627, 1272
BstACI GRCGYC 2 cut(s) 129, 901
BstAUI TGTACA 1 cut(s) 368
BstC8I GCNNGC 5 cut(s) 231, 442, 1071, 1294, 1365
BstDEI CTNAG 3 cut(s) 415, 480, 803
BstEII GGTNACC 1 cut(s) 1369
BstF5I GGATG 1 cut(s) 694
BstFNI CGCG 4 cut(s) 111, 150, 380, 948
BstH2I RGCGCY 2 cut(s) 447, 1334
BstHHI GCGC 4 cut(s) 113, 216, 446, 1333
BstMAI GTCTC 3 cut(s) 146, 503, 1225
BstMCI CGRYCG 1 cut(s) 534
BstMWI GCNNNNNNNGC 4 cut(s) 476, 752, 815, 1298
BstNI CCWGG 2 cut(s) 927, 1409
BstNSI RCATGY 2 cut(s) 157, 1363
BstPAI GACNNNNGTC 1 cut(s) 65
BstPI GGTNACC 1 cut(s) 1369
BstSCI CCNGG 3 cut(s) 925, 1373, 1407
BstSFI CTRYAG 4 cut(s) 119, 260, 1086, 1113
BstSLI GKGCMC 1 cut(s) 40
BstUI CGCG 4 cut(s) 111, 150, 380, 948
BstV1I GCAGC 7 cut(s) 184, 246, 607, 827, 1320, 1386, 1408
BstV2I GAAGAC 1 cut(s) 981
BstX2I RGATCY 1 cut(s) 265
BstYI RGATCY 1 cut(s) 265
BtgZI GCGATG 1 cut(s) 1140
BtsCI GGATG 1 cut(s) 694
BtsIMutI CAGTG 1 cut(s) 811
BtuMI TCGCGA 2 cut(s) 150, 948
BveI ACCTGC 1 cut(s) 418
Cac8I GCNNGC 5 cut(s) 231, 442, 1071, 1294, 1365
CciI TCATGA 2 cut(s) 1167, 1258
CfoI GCGC 4 cut(s) 113, 216, 446, 1333
Cfr10I RCCGGY 2 cut(s) 200, 1101
Cfr13I GGNCC 3 cut(s) 79, 514, 1017
CseI GACGC 2 cut(s) 629, 890
CsiI ACCWGGT 1 cut(s) 1407
Csp6I GTAC 3 cut(s) 369, 798, 1095
CviAII CATG 7 cut(s) 154, 913, 1021, 1168, 1259, 1360, 1420
CviQI GTAC 3 cut(s) 369, 798, 1095
DdeI CTNAG 3 cut(s) 415, 480, 803
DraIII CACNNNGTG 2 cut(s) 366, 806
DrdI GACNNNNNNGTC 1 cut(s) 77
DriI GACNNNNNGTC 1 cut(s) 825
DseDI GACNNNNNNGTC 1 cut(s) 77
Eam1104I CTCTTC 2 cut(s) 627, 1272
Eam1105I GACNNNNNGTC 1 cut(s) 825
EarI CTCTTC 2 cut(s) 627, 1272
EciI GGCGGA 2 cut(s) 26, 1017
Eco130I CCWWGG 1 cut(s) 474
Eco47I GGWCC 3 cut(s) 79, 514, 1017
Eco47III AGCGCT 1 cut(s) 1332
Eco91I GGTNACC 1 cut(s) 1369
EcoO109I RGGNCCY 1 cut(s) 79
EcoO65I GGTNACC 1 cut(s) 1369
EcoRII CCWGG 2 cut(s) 925, 1407
EcoT14I CCWWGG 1 cut(s) 474
ErhI CCWWGG 1 cut(s) 474
Esp3I CGTCTC 1 cut(s) 503
FaeI CATG 7 cut(s) 157, 916, 1024, 1171, 1262, 1363, 1423
FalI AAGNNNNNCTT 4 cut(s) 609, 641, 1145, 1177
FaqI GGGAC 2 cut(s) 648, 932
FatI CATG 7 cut(s) 153, 912, 1020, 1167, 1258, 1359, 1419
FbaI TGATCA 2 cut(s) 51, 1129
FblI GTMKAC 1 cut(s) 69
FokI GGATG 1 cut(s) 701
FspBI CTAG 1 cut(s) 597
GlaI GCGC 4 cut(s) 112, 215, 445, 1332
GsuI CTGGAG 1 cut(s) 26
HaeII RGCGCY 2 cut(s) 447, 1334
HapII CCGG 5 cut(s) 201, 898, 1102, 1339, 1375
HgaI GACGC 2 cut(s) 629, 890
HhaI GCGC 4 cut(s) 113, 216, 446, 1333
Hin1I GRCGYC 2 cut(s) 129, 901
Hin1II CATG 7 cut(s) 157, 916, 1024, 1171, 1262, 1363, 1423
Hin6I GCGC 4 cut(s) 111, 214, 444, 1331
HinP1I GCGC 4 cut(s) 111, 214, 444, 1331
HincII GTYRAC 2 cut(s) 70, 644
HindII GTYRAC 2 cut(s) 70, 644
HindIII AAGCTT 1 cut(s) 1435
HinfI GANTC 5 cut(s) 826, 943, 1119, 1175, 1444
HpaII CCGG 5 cut(s) 201, 898, 1102, 1339, 1375
Hpy166II GTNNAC 5 cut(s) 70, 341, 369, 644, 800
Hpy8I GTNNAC 5 cut(s) 70, 341, 369, 644, 800
Hpy99I CGWCG 3 cut(s) 64, 74, 131
HpyAV CCTTC 3 cut(s) 70, 530, 886
HpyCH4III ACNGT 3 cut(s) 64, 123, 779
HpyCH4IV ACGT 3 cut(s) 129, 165, 365
HpyF10VI GCNNNNNNNGC 4 cut(s) 476, 752, 815, 1298
HpyF3I CTNAG 3 cut(s) 415, 480, 803
HpySE526I ACGT 3 cut(s) 129, 165, 365
Hsp92I GRCGYC 2 cut(s) 129, 901
Hsp92II CATG 7 cut(s) 157, 916, 1024, 1171, 1262, 1363, 1423
HspAI GCGC 4 cut(s) 111, 214, 444, 1331
Kpn2I TCCGGA 1 cut(s) 1338
Ksp22I TGATCA 2 cut(s) 51, 1129
LguI GCTCTTC 1 cut(s) 1272
LmnI GCTCC 4 cut(s) 24, 752, 1086, 1341
Lsp1109I GCAGC 7 cut(s) 184, 246, 607, 827, 1320, 1386, 1408
LweI GCATC 3 cut(s) 627, 679, 1068
MabI ACCWGGT 1 cut(s) 1407
MaeI CTAG 1 cut(s) 597
MaeII ACGT 3 cut(s) 129, 165, 365
MaeIII GTNAC 5 cut(s) 203, 500, 763, 779, 1369
MbiI CCGCTC 1 cut(s) 1367
MboII GAAGA 4 cut(s) 319, 644, 986, 1289
MflI RGATCY 1 cut(s) 265
MhlI GDGCHC 2 cut(s) 40, 858
MluCI AATT 5 cut(s) 14, 134, 525, 585, 1312
MluI ACGCGT 1 cut(s) 378
MlyI GAGTC 2 cut(s) 835, 1113
MmeI TCCRAC 4 cut(s) 150, 730, 849, 1176
MnlI CCTC 6 cut(s) 14, 50, 104, 540, 597, 1446
MroI TCCGGA 1 cut(s) 1338
MroXI GAANNNNTTC 2 cut(s) 954, 1157
MseI TTAA 3 cut(s) 168, 551, 732
MslI CAYNNNNRTG 1 cut(s) 1025
MspA1I CMGCKG 2 cut(s) 1353, 1399
MspI CCGG 5 cut(s) 201, 898, 1102, 1339, 1375
MspR9I CCNGG 3 cut(s) 927, 1375, 1409
MvaI CCWGG 2 cut(s) 927, 1409
MvnI CGCG 4 cut(s) 111, 150, 380, 948
MwoI GCNNNNNNNGC 4 cut(s) 476, 752, 815, 1298
NciI CCSGG 1 cut(s) 1375
NlaIII CATG 7 cut(s) 157, 916, 1024, 1171, 1262, 1363, 1423
NlaIV GGNNCC 3 cut(s) 267, 486, 516
NmeAIII GCCGAG 2 cut(s) 1032, 1101
NmuCI GTSAC 2 cut(s) 203, 779
NruI TCGCGA 2 cut(s) 150, 948
NspI RCATGY 2 cut(s) 157, 1363
PagI TCATGA 2 cut(s) 1167, 1258
PciSI GCTCTTC 1 cut(s) 1272
PdmI GAANNNNTTC 2 cut(s) 954, 1157
PfeI GAWTC 3 cut(s) 943, 1175, 1444
PleI GAGTC 2 cut(s) 834, 1113
PpsI GAGTC 2 cut(s) 834, 1113
PpuMI RGGWCCY 1 cut(s) 79
PshAI GACNNNNGTC 1 cut(s) 65
PsiI TTATAA 1 cut(s) 1459
Psp1406I AACGTT 1 cut(s) 165
Psp5II RGGWCCY 1 cut(s) 79
Psp6I CCWGG 2 cut(s) 925, 1407
PspEI GGTNACC 1 cut(s) 1369
PspGI CCWGG 2 cut(s) 925, 1407
PspN4I GGNNCC 3 cut(s) 267, 486, 516
PspPI GGNCC 3 cut(s) 79, 514, 1017
PspPPI RGGWCCY 1 cut(s) 79
PstI CTGCAG 3 cut(s) 264, 1090, 1117
PsuI RGATCY 1 cut(s) 265
PvuII CAGCTG 1 cut(s) 1399
RruI TCGCGA 2 cut(s) 150, 948
RsaI GTAC 3 cut(s) 370, 799, 1096
RsaNI GTAC 3 cut(s) 369, 798, 1095
RseI CAYNNNNRTG 1 cut(s) 1025
SalI GTCGAC 1 cut(s) 68
SapI GCTCTTC 1 cut(s) 1272
SaqAI TTAA 3 cut(s) 168, 551, 732
Sau96I GGNCC 3 cut(s) 79, 514, 1017
SbfI CCTGCAGG 1 cut(s) 1117
SchI GAGTC 2 cut(s) 835, 1113
ScrFI CCNGG 3 cut(s) 927, 1375, 1409
SdaI CCTGCAGG 1 cut(s) 1117
SduI GDGCHC 2 cut(s) 40, 858
SexAI ACCWGGT 1 cut(s) 1407
SfaNI GCATC 3 cut(s) 627, 679, 1068
SfcI CTRYAG 4 cut(s) 119, 260, 1086, 1113
SinI GGWCC 3 cut(s) 79, 514, 1017
SmiMI CAYNNNNRTG 1 cut(s) 1025
SmlI CTYRAG 3 cut(s) 458, 723, 1188
SmoI CTYRAG 3 cut(s) 458, 723, 1188
SpeI ACTAGT 1 cut(s) 596
Sse8387I CCTGCAGG 1 cut(s) 1117
Sse9I AATT 5 cut(s) 14, 134, 525, 585, 1312
SspI AATATT 1 cut(s) 991
SspMI CTAG 1 cut(s) 597
StyD4I CCNGG 3 cut(s) 925, 1373, 1407
StyI CCWWGG 1 cut(s) 474
TaaI ACNGT 3 cut(s) 64, 123, 779
TaiI ACGT 3 cut(s) 132, 168, 368
TaqI TCGA 6 cut(s) 69, 240, 450, 534, 879, 1076
TasI AATT 5 cut(s) 14, 134, 525, 585, 1312
TatI WGTACW 1 cut(s) 368
TauI GCSGC 3 cut(s) 473, 795, 1356
TfiI GAWTC 3 cut(s) 943, 1175, 1444
Tru1I TTAA 3 cut(s) 168, 551, 732
Tru9I TTAA 3 cut(s) 168, 551, 732
TscAI CASTG 1 cut(s) 811
TseFI GTSAC 2 cut(s) 203, 779
TseI GCWGC 7 cut(s) 197, 259, 620, 815, 1333, 1396, 1399
Tsp45I GTSAC 2 cut(s) 203, 779
TspDTI ATGAA 4 cut(s) 704, 1166, 1275, 1457
TspGWI ACGGA 4 cut(s) 477, 527, 875, 1252
TspRI CASTG 1 cut(s) 811
VpaK11BI GGWCC 3 cut(s) 79, 514, 1017
XapI RAATTY 1 cut(s) 1312
XceI RCATGY 2 cut(s) 157, 1363
XmiI GTMKAC 1 cut(s) 69
XmnI GAANNNNTTC 2 cut(s) 954, 1157
XspI CTAG 1 cut(s) 597
ZraI GACGTC 1 cut(s) 130
Using CommOnly database (standard laboratory enzymes). Scanned on CDS sequence.