Rmu_sc0002513.1_g000007

Mitogen-activated protein kinase kinase kinase

Design Primers CRISPR Design

Basic Information

Type: gene
Biological Identity
rosa_multiflora
Rmu_sc0002513.1
OG0019501
Physical Location & Seq
Reverse (-)
36313 .. 41719
5407 bp
Loading structure...
UTR
Exon/CDS
Intron
Rmu_sc0002513.1_g000007.1.cds

Sequence Viewer

Length: 1515 bp
atgcaaaaccgaaccgagcccaacttcggttcggtttgcggttttggcccaaaaccgaaccgattggaaccgaacccacccctagctggcaaggctaacaagggcgtagcgaggctaacagtaggcacaggggtgcgcggagatggggcaggcgaggctagccggcagggccgattctgggggatccgccagaggttttgtactcctaggatcgaggactttgaggtgtgcggcggaggtagcttgggtttaaaggctacgattttagggactgagcgagatcttgtcgaagctactgtggaatttgtgtacagtaaactacaacacgttgtggaaatcaacactggagattttgtagcaacaaggcaagccactactgaaattgaacctgcaatctctcgacaaactgctgcatctcttccttcatcagctgaaccatcatcaacagctggtcagtggaagtatgacgtgtatttgagtttcttgggtgcagatactcgtaacagttttacagatcatttatactatgggctgcaacatagtggactctcaatgaatcaacatcttgaacgagagcaccccatttctccctctcttgtggaggcaattgaagaatcaaggtttgcaatcgttgttatgtcgaaaagctatgttcattatacttggtgtttggaggaacttacaaagatctatgagtgcatgaaaaaccaaaacagaattcttccaattttttataatgtggcaaacgttgatgtacggtactgcaagacgagtttcgaaaaagctttccaagagcatgaaagctctgggcgacacacatcagacaagatgaagggctggaaagatgcttcaaaccatgtagtcagtctctgcgggactgatagagaacttgtggagacaattgtggattttgtgtgccgtcaagtaaaacctattgaaattgggtttacaatctctactaattggaaaaagggtcatcttctcggctgcggctccttcgggtctgtgtatgaaggaatttctgagatctttgttgatatgatcttgctctctttcagtggcggatgcttcattgctgtcaaggaagtttccttgcttgatcaaggaaaagtttctcaacttgaacaggatgaatcaaagctctacatctttctggaacttgtaaccaaaggctcccttcaaaagctatatcaaacgtaccctcttacagatttgcatgtctctgtatacactagacagatcttgcaaggtctgaagtatcttcacgaccaaaatgtgattcacagggacattaaatgtgcaaatattttggtgcatgctaatggatctgtgaagcttgcagactttggcttggcaaagcttgctaaacctggaaatccaggttatggccttccagctgatatatggagtcttggatgtactgtgttggagatgttaactaggatggttccataccctaatctcgaatgggtttgtgccattttcaatagactctttatggttcagaatgactag
Functional Annotation
Gene Ontology
Molecular Function
Biological Process
Cellular Component
GO:0000165 GO:0000186 GO:0001101 GO:0001932 GO:0001934 GO:0003674 GO:0003676 GO:0003677 GO:0003824 GO:0004672 GO:0004674 GO:0004709 GO:0005488 GO:0005515 GO:0005575 GO:0005622 GO:0005623 GO:0005634 GO:0005737 GO:0005768 GO:0005886 GO:0006464 GO:0006468 GO:0006793 GO:0006796 GO:0006807 GO:0006950 GO:0006970 GO:0007154 GO:0007165 GO:0007275 GO:0007346 GO:0007584 GO:0008150 GO:0008152 GO:0009266 GO:0009409 GO:0009605 GO:0009611 GO:0009628 GO:0009631 GO:0009651 GO:0009888 GO:0009893 GO:0009966 GO:0009967 GO:0009987 GO:0009991 GO:0010033 GO:0010243 GO:0010449 GO:0010562 GO:0010604 GO:0010646 GO:0010647 GO:0012505 GO:0016020 GO:0016301 GO:0016310 GO:0016740 GO:0016772 GO:0016773 GO:0019220 GO:0019222 GO:0019538 GO:0019899 GO:0019900 GO:0022622 GO:0023014 GO:0023051 GO:0023052 GO:0023056 GO:0031098 GO:0031323 GO:0031325 GO:0031347 GO:0031399 GO:0031401 GO:0031410 GO:0031667 GO:0031982 GO:0032147 GO:0032268 GO:0032270 GO:0032501 GO:0032502 GO:0033554 GO:0033674 GO:0035266 GO:0035556 GO:0036211 GO:0040007 GO:0042221 GO:0042325 GO:0042327 GO:0042493 GO:0043085 GO:0043170 GO:0043200 GO:0043226 GO:0043227 GO:0043229 GO:0043231 GO:0043405 GO:0043406 GO:0043408 GO:0043410 GO:0043412 GO:0043549 GO:0044093 GO:0044237 GO:0044238 GO:0044260 GO:0044267 GO:0044424 GO:0044444 GO:0044464 GO:0045859 GO:0045860 GO:0045937 GO:0046777 GO:0048364 GO:0048507 GO:0048518 GO:0048522 GO:0048583 GO:0048584 GO:0048589 GO:0048731 GO:0048856 GO:0050789 GO:0050790 GO:0050794 GO:0050896 GO:0051171 GO:0051173 GO:0051174 GO:0051246 GO:0051247 GO:0051338 GO:0051347 GO:0051716 GO:0051726 GO:0060255 GO:0065007 GO:0065009 GO:0071704 GO:0071900 GO:0071902 GO:0071944 GO:0080090 GO:0080134 GO:0097159 GO:0097708 GO:0099402 GO:0140096 GO:1901363 GO:1901564 GO:1901698 GO:1901700 GO:1902065 GO:1902531 GO:1902533
KEGG Pathways
Metabolic & Signaling
ko04016 ko04626 map04016 map04626
Pfam Domains
Protein Families
Pkinase

Protein Analysis

504

Amino Acids

56.21

Weight (kDa)

6.41

Isoelectric Point (pI)

32.71

Instability Index
Protein Domains (Pfam)
No Pfam domains detected for this protein.
Hydrophobicity Profile (Kyte-Doolittle)
AI Structure Prediction Report
Calculating structure properties...

Genomic Context

Gene Family Tree

Species Filter
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Publication-ready
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Restriction Enzyme Sites

1 / 10
Enzyme Recognition Site Cut Count Positions (bp)
AanI TTATAA 1 cut(s) 735
Acc36I ACCTGC 1 cut(s) 397
AccB7I CCANNNNNTGG 1 cut(s) 1385
AccI GTMKAC 1 cut(s) 1227
AccII CGCG 1 cut(s) 138
AciI CCGC 8 cut(s) 39, 138, 187, 231, 234, 873, 990, 1062
AclI AACGTT 1 cut(s) 747
AclWI GGATC 4 cut(s) 178, 191, 218, 1333
AcsI RAATTY 3 cut(s) 302, 717, 1017
AcuI CTGAAG 1 cut(s) 1274
AdeI CACNNNGTG 1 cut(s) 331
AfaI GTAC 6 cut(s) 202, 311, 756, 761, 1199, 1420
AfiI CCNNNNNNNGG 4 cut(s) 26, 86, 178, 1385
AflIII ACRYGT 2 cut(s) 325, 468
AgsI TTSAA 8 cut(s) 386, 569, 611, 852, 938, 1124, 1181, 1486
AjiI CACGTC 1 cut(s) 469
AjnI CCWGG 2 cut(s) 1369, 1378
AjuI GAANNNNNNNTTGG 4 cut(s) 702, 734, 1263, 1295
AleI CACNNNNGTG 1 cut(s) 131
Alw21I GWGCWC 1 cut(s) 579
Alw26I GTCTC 3 cut(s) 872, 890, 1225
AlwI GGATC 4 cut(s) 178, 191, 218, 1333
AlwNI CAGNNNCTG 1 cut(s) 870
AoxI GGCC 3 cut(s) 46, 169, 1387
ApeKI GCWGC 3 cut(s) 410, 532, 987
ApoI RAATTY 3 cut(s) 302, 717, 1017
Asp700I GAANNNNTTC 1 cut(s) 1111
AspA2I CCTAGG 1 cut(s) 206
AspLEI GCGC 1 cut(s) 138
AspS9I GGNCC 2 cut(s) 47, 169
AsuII TTCGAA 1 cut(s) 777
AsuNHI GCTAGC 1 cut(s) 158
AvrII CCTAGG 1 cut(s) 206
BamHI GGATCC 1 cut(s) 183
BanII GRGCYC 1 cut(s) 21
Bbv12I GWGCWC 1 cut(s) 579
BbvI GCAGC 3 cut(s) 397, 519, 974
BccI CCATC 3 cut(s) 137, 445, 1438
BceAI ACGGC 1 cut(s) 903
BciT130I CCWGG 2 cut(s) 1371, 1380
BclI TGATCA 1 cut(s) 1099
BcoDI GTCTC 3 cut(s) 872, 890, 1225
BfaI CTAG 6 cut(s) 83, 159, 207, 1233, 1440, 1513
BfuAI ACCTGC 1 cut(s) 397
BglI GCCNNNNNGGC 1 cut(s) 168
BglII AGATCT 4 cut(s) 280, 687, 1026, 1239
BisI GCNGC 5 cut(s) 232, 411, 533, 988, 991
BlnI CCTAGG 1 cut(s) 206
BlsI GCNGC 5 cut(s) 233, 412, 534, 989, 992
Bme1390I CCNGG 2 cut(s) 1371, 1380
BmgBI CACGTC 1 cut(s) 469
BmgT120I GGNCC 2 cut(s) 47, 169
BmiI GGNNCC 5 cut(s) 69, 185, 994, 1174, 1449
BmrFI CCNGG 2 cut(s) 1371, 1380
BmsI GCATC 3 cut(s) 422, 835, 1055
BmtI GCTAGC 1 cut(s) 162
BpmI CTGGAG 1 cut(s) 366
Bpu14I TTCGAA 1 cut(s) 777
BsaJI CCNNGG 1 cut(s) 206
BsaXI ACNNNNNCTCC 2 cut(s) 339, 369
Bsc4I CCNNNNNNNGG 4 cut(s) 26, 86, 178, 1385
Bse118I RCCGGY 1 cut(s) 162
Bse1I ACTGG 1 cut(s) 349
Bse3DI GCAATG 1 cut(s) 1071
BseBI CCWGG 2 cut(s) 1371, 1380
BseDI CCNNGG 1 cut(s) 206
BseGI GGATG 4 cut(s) 1070, 1135, 1421, 1449
BseLI CCNNNNNNNGG 4 cut(s) 26, 86, 178, 1385
BseMI GCAATG 1 cut(s) 1071
BseMII CTCAG 2 cut(s) 264, 1014
BseNI ACTGG 1 cut(s) 349
BseXI GCAGC 3 cut(s) 397, 519, 974
BsgI GTGCAG 1 cut(s) 510
Bsh1236I CGCG 1 cut(s) 138
BshFI GGCC 3 cut(s) 48, 171, 1389
BsiHKAI GWGCWC 1 cut(s) 579
BsiSI CCGG 1 cut(s) 163
BslFI GGGAC 3 cut(s) 283, 889, 1301
BslI CCNNNNNNNGG 4 cut(s) 26, 86, 178, 1385
BsmAI GTCTC 3 cut(s) 872, 890, 1225
BsmFI GGGAC 3 cut(s) 283, 889, 1301
BsnI GGCC 3 cut(s) 48, 171, 1389
Bsp119I TTCGAA 1 cut(s) 777
Bsp1286I GDGCHC 2 cut(s) 21, 579
Bsp1407I TGTACA 1 cut(s) 309
BspACI CCGC 8 cut(s) 39, 138, 187, 231, 234, 873, 990, 1062
BspANI GGCC 3 cut(s) 48, 171, 1389
BspCNI CTCAG 2 cut(s) 265, 1015
BspFNI CGCG 1 cut(s) 138
BspLI GGNNCC 5 cut(s) 69, 185, 994, 1174, 1449
BspMI ACCTGC 1 cut(s) 397
BspOI GCTAGC 1 cut(s) 162
BspPI GGATC 4 cut(s) 178, 191, 218, 1333
BspT104I TTCGAA 1 cut(s) 777
BsrDI GCAATG 1 cut(s) 1071
BsrFI RCCGGY 1 cut(s) 162
BsrGI TGTACA 1 cut(s) 309
BsrI ACTGG 1 cut(s) 349
BssAI RCCGGY 1 cut(s) 162
BssECI CCNNGG 1 cut(s) 206
BssNAI GTATAC 1 cut(s) 1228
BssT1I CCWWGG 1 cut(s) 206
Bst1107I GTATAC 1 cut(s) 1228
Bst2UI CCWGG 2 cut(s) 1371, 1380
Bst4CI ACNGT 6 cut(s) 121, 298, 314, 506, 759, 1423
Bst6I CTCTTC 1 cut(s) 423
BstAPI GCANNNNNTGC 1 cut(s) 1361
BstAUI TGTACA 1 cut(s) 309
BstBI TTCGAA 1 cut(s) 777
BstC8I GCNNGC 8 cut(s) 88, 151, 160, 164, 369, 1317, 1338, 1362
BstDEI CTNAG 2 cut(s) 273, 1023
BstF5I GGATG 4 cut(s) 1070, 1135, 1421, 1449
BstFNI CGCG 1 cut(s) 138
BstHHI GCGC 1 cut(s) 138
BstMAI GTCTC 3 cut(s) 872, 890, 1225
BstMWI GCNNNNNNNGC 7 cut(s) 45, 92, 155, 159, 168, 240, 1361
BstNI CCWGG 2 cut(s) 1371, 1380
BstNSI RCATGY 2 cut(s) 1220, 1319
BstSCI CCNGG 2 cut(s) 1369, 1378
BstUI CGCG 1 cut(s) 138
BstV1I GCAGC 3 cut(s) 397, 519, 974
BstX2I RGATCY 6 cut(s) 183, 280, 687, 1026, 1239, 1325
BstYI RGATCY 6 cut(s) 183, 280, 687, 1026, 1239, 1325
BstZ17I GTATAC 1 cut(s) 1228
BsuRI GGCC 3 cut(s) 48, 171, 1389
BtrI CACGTC 1 cut(s) 469
BtsCI GGATG 4 cut(s) 1070, 1135, 1421, 1449
BtsIMutI CAGTG 3 cut(s) 342, 461, 1063
BveI ACCTGC 1 cut(s) 397
Cac8I GCNNGC 8 cut(s) 88, 151, 160, 164, 369, 1317, 1338, 1362
CaiI CAGNNNCTG 1 cut(s) 870
CfoI GCGC 1 cut(s) 138
Cfr10I RCCGGY 1 cut(s) 162
Cfr13I GGNCC 2 cut(s) 47, 169
Csp6I GTAC 6 cut(s) 201, 310, 755, 760, 1198, 1419
CviAII CATG 5 cut(s) 700, 797, 857, 1217, 1316
CviQI GTAC 6 cut(s) 201, 310, 755, 760, 1198, 1419
DdeI CTNAG 2 cut(s) 273, 1023
DraI TTTAAA 1 cut(s) 252
DraIII CACNNNGTG 1 cut(s) 331
Eam1104I CTCTTC 1 cut(s) 423
EarI CTCTTC 1 cut(s) 423
EciI GGCGGA 3 cut(s) 176, 249, 1077
Eco130I CCWWGG 1 cut(s) 206
Eco24I GRGCYC 1 cut(s) 21
Eco57I CTGAAG 1 cut(s) 1274
EcoRI GAATTC 1 cut(s) 717
EcoRII CCWGG 2 cut(s) 1369, 1378
EcoT14I CCWWGG 1 cut(s) 206
EcoT38I GRGCYC 1 cut(s) 21
ErhI CCWWGG 1 cut(s) 206
FaeI CATG 5 cut(s) 703, 800, 860, 1220, 1319
FalI AAGNNNNNCTT 2 cut(s) 1161, 1193
FaqI GGGAC 3 cut(s) 283, 889, 1301
FatI CATG 5 cut(s) 699, 796, 856, 1216, 1315
FauI CCCGC 1 cut(s) 866
FbaI TGATCA 1 cut(s) 1099
FblI GTMKAC 1 cut(s) 1227
Fnu4HI GCNGC 5 cut(s) 232, 411, 533, 988, 991
FokI GGATG 4 cut(s) 1077, 1142, 1428, 1456
FriOI GRGCYC 1 cut(s) 21
Fsp4HI GCNGC 5 cut(s) 232, 411, 533, 988, 991
FspBI CTAG 6 cut(s) 83, 159, 207, 1233, 1440, 1513
GlaI GCGC 1 cut(s) 137
GluI GCNGC 5 cut(s) 232, 411, 533, 988, 991
GsuI CTGGAG 1 cut(s) 366
HaeIII GGCC 3 cut(s) 48, 171, 1389
HapII CCGG 1 cut(s) 163
HhaI GCGC 1 cut(s) 138
Hin1II CATG 5 cut(s) 703, 800, 860, 1220, 1319
Hin6I GCGC 1 cut(s) 136
HinP1I GCGC 1 cut(s) 136
HincII GTYRAC 1 cut(s) 1437
HindII GTYRAC 1 cut(s) 1437
HindIII AAGCTT 3 cut(s) 783, 1334, 1358
HinfI GANTC 8 cut(s) 174, 546, 556, 614, 1133, 1279, 1408, 1491
HpaI GTTAAC 1 cut(s) 1437
HpaII CCGG 1 cut(s) 163
Hpy166II GTNNAC 6 cut(s) 310, 317, 545, 948, 1228, 1437
Hpy188I TCNGA 4 cut(s) 823, 1024, 1254, 1506
Hpy188III TCNNGA 5 cut(s) 399, 566, 1154, 1265, 1463
Hpy8I GTNNAC 6 cut(s) 310, 317, 545, 948, 1228, 1437
HpyAV CCTTC 6 cut(s) 432, 826, 1006, 1007, 1187, 1400
HpyCH4III ACNGT 6 cut(s) 121, 298, 314, 506, 759, 1423
HpyCH4IV ACGT 4 cut(s) 327, 468, 747, 1196
HpyF10VI GCNNNNNNNGC 7 cut(s) 45, 92, 155, 159, 168, 240, 1361
HpyF3I CTNAG 2 cut(s) 273, 1023
HpySE526I ACGT 4 cut(s) 327, 468, 747, 1196
Hsp92II CATG 5 cut(s) 703, 800, 860, 1220, 1319
HspAI GCGC 1 cut(s) 136
KroI GCCGGC 1 cut(s) 162
KroNI GCCGGC 1 cut(s) 164
Ksp22I TGATCA 1 cut(s) 1099
KspAI GTTAAC 1 cut(s) 1437
LmnI GCTCC 2 cut(s) 998, 1178
Lsp1109I GCAGC 3 cut(s) 397, 519, 974
LweI GCATC 3 cut(s) 422, 835, 1055
MaeI CTAG 6 cut(s) 83, 159, 207, 1233, 1440, 1513
MaeII ACGT 4 cut(s) 327, 468, 747, 1196
MaeIII GTNAC 2 cut(s) 500, 1162
MboII GAAGA 5 cut(s) 410, 623, 713, 971, 1253
MfeI CAATTG 2 cut(s) 606, 900
MflI RGATCY 6 cut(s) 183, 280, 687, 1026, 1239, 1325
MhlI GDGCHC 2 cut(s) 21, 579
MluCI AATT 9 cut(s) 302, 381, 606, 717, 726, 900, 939, 961, 1017
MlyI GAGTC 3 cut(s) 540, 1417, 1485
MmeI TCCRAC 1 cut(s) 1407
MroNI GCCGGC 1 cut(s) 162
MroXI GAANNNNTTC 1 cut(s) 1111
MseI TTAA 3 cut(s) 251, 1293, 1436
MslI CAYNNNNRTG 2 cut(s) 131, 1320
MspA1I CMGCKG 3 cut(s) 431, 449, 1397
MspI CCGG 1 cut(s) 163
MspR9I CCNGG 2 cut(s) 1371, 1380
MunI CAATTG 2 cut(s) 606, 900
MvaI CCWGG 2 cut(s) 1371, 1380
MvnI CGCG 1 cut(s) 138
MwoI GCNNNNNNNGC 7 cut(s) 45, 92, 155, 159, 168, 240, 1361
NaeI GCCGGC 1 cut(s) 164
NgoMIV GCCGGC 1 cut(s) 162
NheI GCTAGC 1 cut(s) 158
NlaIII CATG 5 cut(s) 703, 800, 860, 1220, 1319
NlaIV GGNNCC 5 cut(s) 69, 185, 994, 1174, 1449
NmeAIII GCCGAG 1 cut(s) 963
NspI RCATGY 2 cut(s) 1220, 1319
NspV TTCGAA 1 cut(s) 777
OliI CACNNNNGTG 1 cut(s) 131
PaeI GCATGC 1 cut(s) 1319
PdiI GCCGGC 1 cut(s) 164
PdmI GAANNNNTTC 1 cut(s) 1111
PfeI GAWTC 5 cut(s) 174, 556, 614, 1133, 1279
PflMI CCANNNNNTGG 1 cut(s) 1385
PkrI GCNGC 5 cut(s) 233, 412, 534, 989, 992
PleI GAGTC 3 cut(s) 540, 1416, 1485
PpsI GAGTC 3 cut(s) 540, 1416, 1485
PsiI TTATAA 1 cut(s) 735
Psp1406I AACGTT 1 cut(s) 747
Psp6I CCWGG 2 cut(s) 1369, 1378
PspGI CCWGG 2 cut(s) 1369, 1378
PspN4I GGNNCC 5 cut(s) 69, 185, 994, 1174, 1449
PspPI GGNCC 2 cut(s) 47, 169
PstNI CAGNNNCTG 1 cut(s) 870
PsuI RGATCY 6 cut(s) 183, 280, 687, 1026, 1239, 1325
PvuII CAGCTG 3 cut(s) 431, 449, 1397
RsaI GTAC 6 cut(s) 202, 311, 756, 761, 1199, 1420
RsaNI GTAC 6 cut(s) 201, 310, 755, 760, 1198, 1419
RseI CAYNNNNRTG 2 cut(s) 131, 1320
SaqAI TTAA 3 cut(s) 251, 1293, 1436
SatI GCNGC 5 cut(s) 232, 411, 533, 988, 991
Sau96I GGNCC 2 cut(s) 47, 169
SchI GAGTC 3 cut(s) 540, 1417, 1485
ScrFI CCNGG 2 cut(s) 1371, 1380
SduI GDGCHC 2 cut(s) 21, 579
SfaNI GCATC 3 cut(s) 422, 835, 1055
SfuI TTCGAA 1 cut(s) 777
SmiMI CAYNNNNRTG 2 cut(s) 131, 1320
SphI GCATGC 1 cut(s) 1319
Sse9I AATT 9 cut(s) 302, 381, 606, 717, 726, 900, 939, 961, 1017
SsiI CCGC 8 cut(s) 39, 138, 187, 231, 234, 873, 990, 1062
SspI AATATT 1 cut(s) 1306
SspMI CTAG 6 cut(s) 83, 159, 207, 1233, 1440, 1513
StyD4I CCNGG 2 cut(s) 1369, 1378
StyI CCWWGG 1 cut(s) 206
TaaI ACNGT 6 cut(s) 121, 298, 314, 506, 759, 1423
TaiI ACGT 4 cut(s) 330, 471, 750, 1199
TaqI TCGA 6 cut(s) 213, 288, 400, 641, 777, 1464
TasI AATT 9 cut(s) 302, 381, 606, 717, 726, 900, 939, 961, 1017
TatI WGTACW 3 cut(s) 200, 309, 1418
TauI GCSGC 2 cut(s) 234, 993
TfiI GAWTC 5 cut(s) 174, 556, 614, 1133, 1279
Tru1I TTAA 3 cut(s) 251, 1293, 1436
Tru9I TTAA 3 cut(s) 251, 1293, 1436
TscAI CASTG 3 cut(s) 349, 461, 1063
TseI GCWGC 3 cut(s) 410, 532, 987
TspDTI ATGAA 9 cut(s) 414, 569, 644, 716, 813, 845, 1026, 1060, 1146
TspRI CASTG 3 cut(s) 349, 461, 1063
Van91I CCANNNNNTGG 1 cut(s) 1385
XapI RAATTY 3 cut(s) 302, 717, 1017
XceI RCATGY 2 cut(s) 1220, 1319
XcmI CCANNNNNNNNNTGG 1 cut(s) 1401
XmaJI CCTAGG 1 cut(s) 206
XmiI GTMKAC 1 cut(s) 1227
XmnI GAANNNNTTC 1 cut(s) 1111
XspI CTAG 6 cut(s) 83, 159, 207, 1233, 1440, 1513
Using CommOnly database (standard laboratory enzymes). Scanned on CDS sequence.