Rorug03G0039600

No description available

Design Primers CRISPR Design

Basic Information

Type: gene
Biological Identity
rosa_rugosa
GWHBQTZ00000003
OG0010297
Physical Location & Seq
Reverse (-)
3086744 .. 3088690
1947 bp
Loading structure...
UTR
Exon/CDS
Intron
Rorug03G0039600.1

Sequence Viewer

Length: 1392 bp
ATGAGTAGTGACCGAAAACCCTTGCACATTGCCATGTATCCCTGGTTTGCTATGGGGCATTTGTTCCCATTTCTCCACATCTCTAACAAACTAGCCGAGAGAGGCCATAGAATCTCCTTCTTCCTACCCCAAAAAACACAAGCCAAGTTGGAGCATTACAATCTCCACCCTGATCTCATCACCTTCTTCCCCCTCACCATCCCTCACGTCGAGGGCCTTCCCGAGGGGGCGGAGACCACTGCCGACGTTCCTTTTCCTCTCCACACCCACGTCGTCACCGCCATGGACCTCACTCGCCCCCAAATTGAGAAATTTCTCCGAGAACTCAAACCGAATTTGGTCTTCTACGATTTCACTCATTGGTTGCCTCAACTGTTGCGTGAGCTGGACGGCAACATGAAGTCAGTCCACTACTGCACTATGAGCCCCGTGACAATGGGGTACCTTCTAAGCCCGGAGAGGAAGCTAACCGAAAAACCGTTGGTTGCAACTGATCTTCTAGAGGCTCCTACGTCATTTCCTTCTTCATCCATCAAGTTGCGGACACATGAAGCTCGAGGACTAACCTATGTGACAAACCTGGAATACGGGCGTGGGATAACATTTCTTCAACGCCAACTGACCTCTTTGGCCGAATGTGATGCCCTTGGTTTCAAGACATGTAAAGAATTCGAAGGGAAGTTTTGTGATTACCTAGAAGTGCAGATGAAGAAGCCAGTGATTCTCGCGGGGCCGGTGGTACCGCCCCCTCCGAGCTCGGCCTTGGAAGAGAAATGGGAAGAGTGGCTTGGAGGGTTTGAGCCAAGAAGTGTGATTTTTTGTGCTTTCGGAAGTGAATGCGTGCTGAAGAAGGAACAGTTGCAAGAACTGGTGTTGGGATTCGAGCTCACGGGTCTTCCGTTCTTTGCAGCGCTGAAAACCCCTGTGGGATTCGAGTCCATCGAAGCTGCATTGCCTGAAGGGTTCGAAGAGAGGGTGAAAGGGAGAGGTGTTGTTTACAGTGGCTGGGTGCAACAGCCGTTGATACTGAAGCACCCCTCTGTGGGCTGCTTTGTGACTCACTGTGGTTCGGGGTCTCTAGCCGAGGCGATGATGAACCATTGCCGGTTGGTGCTTCTGCCGCACGTCGGAGACCAGATCATCAATGCGAGGATGATGGGAGGTGATGATCTGAGAGTGGGAGTTGAAGTCGAGAAGGGAGACGAAGATGGCTTGTTTAGCAAGGAAGGTGTTTGCAAAGCTGTGGAGGCTGTGATGGATGATGAGAGTGAGGTAGGGAAAGAAGTGAAGGCTAACCATGCTCAGTGGAGGGAGTTCTTCTCCAACAAAGGGCTTGAGAAAACGTACATTGATGGTTTTGATCAGAGGCTGCATCAACTGCTGGAATGTTAA
Functional Annotation
Gene Ontology
Molecular Function
Biological Process
Cellular Component
No ontology terms assigned.
KEGG Pathways
Metabolic & Signaling
No pathways identified.
Pfam Domains
Protein Families
No domains found.

Protein Analysis

463

Amino Acids

52.15

Weight (kDa)

5.73

Isoelectric Point (pI)

46.16

Instability Index
Protein Domains (Pfam)
Domain Name Pfam ID Position E-value Description
UDPGT PF00201 335 - 423 8.3e-12 UDP-glucoronosyl and UDP-glucosyl transferase
Hydrophobicity Profile (Kyte-Doolittle)
AI Structure Prediction Report
Calculating structure properties...

Genomic Context

Gene Family Tree

Species Filter
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Publication-ready
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Restriction Enzyme Sites

1 / 10
Enzyme Recognition Site Cut Count Positions (bp)
Acc65I GGTACC 2 cut(s) 441, 739
AccB1I GGYRCC 2 cut(s) 441, 739
AccII CGCG 1 cut(s) 728
AciI CCGC 6 cut(s) 230, 279, 541, 728, 743, 1121
AcoI YGGCCR 1 cut(s) 630
AcsI RAATTY 3 cut(s) 311, 334, 668
AcuI CTGAAG 3 cut(s) 866, 978, 1049
AfaI GTAC 3 cut(s) 443, 741, 1346
AfeI AGCGCT 1 cut(s) 912
AfiI CCNNNNNNNGG 5 cut(s) 223, 1042, 1043, 1127, 1329
AflIII ACRYGT 1 cut(s) 659
AgsI TTSAA 3 cut(s) 611, 655, 1187
AjiI CACGTC 3 cut(s) 208, 271, 1126
AjnI CCWGG 2 cut(s) 41, 579
AjuI GAANNNNNNNTTGG 2 cut(s) 771, 803
AluBI AGCT 7 cut(s) 385, 466, 554, 756, 886, 947, 1241
AluI AGCT 7 cut(s) 385, 466, 554, 756, 886, 947, 1241
Alw21I GWGCWC 2 cut(s) 758, 888
Alw26I GTCTC 4 cut(s) 227, 1080, 1125, 1194
AlwNI CAGNNNCTG 2 cut(s) 1005, 1369
Ama87I CYCGRG 2 cut(s) 221, 555
Aor51HI AGCGCT 1 cut(s) 912
AoxI GGCC 5 cut(s) 103, 214, 630, 731, 759
ApeKI GCWGC 4 cut(s) 908, 947, 1047, 1369
ApoI RAATTY 3 cut(s) 311, 334, 668
Asp718I GGTACC 2 cut(s) 441, 739
AspLEI GCGC 1 cut(s) 913
AspS9I GGNCC 3 cut(s) 214, 286, 731
AsuC2I CCSGG 1 cut(s) 455
AsuHPI GGTGA 5 cut(s) 172, 187, 268, 988, 1175
AsuII TTCGAA 2 cut(s) 672, 966
AvaI CYCGRG 2 cut(s) 221, 555
AvaII GGWCC 1 cut(s) 286
BanI GGYRCC 2 cut(s) 441, 739
BanII GRGCYC 3 cut(s) 428, 758, 888
BbsI GAAGAC 2 cut(s) 334, 887
Bbv12I GWGCWC 2 cut(s) 758, 888
BbvI GCAGC 4 cut(s) 920, 934, 1034, 1356
BccI CCATC 7 cut(s) 206, 539, 947, 1150, 1202, 1249, 1346
BceAI ACGGC 2 cut(s) 406, 1003
BcgI CGANNNNNNTGC 2 cut(s) 623, 657
BciT130I CCWGG 2 cut(s) 43, 581
BciVI GTATCC 1 cut(s) 48
BclI TGATCA 1 cut(s) 1360
BcnI CCSGG 1 cut(s) 455
BcoDI GTCTC 4 cut(s) 227, 1080, 1125, 1194
BfaI CTAG 4 cut(s) 92, 500, 695, 1079
BfoI RGCGCY 1 cut(s) 914
BfuI GTATCC 1 cut(s) 48
BisI GCNGC 5 cut(s) 909, 948, 1048, 1121, 1370
BlsI GCNGC 5 cut(s) 910, 949, 1049, 1122, 1371
Bme1390I CCNGG 3 cut(s) 43, 455, 581
Bme18I GGWCC 1 cut(s) 286
BmeT110I CYCGRG 2 cut(s) 221, 555
BmgBI CACGTC 3 cut(s) 208, 271, 1126
BmgT120I GGNCC 3 cut(s) 214, 286, 731
BmiI GGNNCC 4 cut(s) 443, 507, 732, 741
BmrFI CCNGG 3 cut(s) 43, 455, 581
BmsI GCATC 2 cut(s) 631, 1381
BpiI GAAGAC 2 cut(s) 334, 887
BplI GAGNNNNNCTC 2 cut(s) 1304, 1336
Bpu14I TTCGAA 2 cut(s) 672, 966
BpuEI CTTGAG 1 cut(s) 1355
BpuMI CCSGG 1 cut(s) 455
BsaI GGTCTC 3 cut(s) 227, 1080, 1125
BsaJI CCNNGG 6 cut(s) 41, 222, 282, 646, 762, 1083
BsaXI ACNNNNNCTCC 2 cut(s) 1173, 1203
Bsc4I CCNNNNNNNGG 5 cut(s) 223, 1042, 1043, 1127, 1329
Bse118I RCCGGY 2 cut(s) 733, 1104
Bse1I ACTGG 2 cut(s) 716, 873
Bse3DI GCAATG 3 cut(s) 27, 950, 1099
BseBI CCWGG 2 cut(s) 43, 581
BseDI CCNNGG 6 cut(s) 41, 222, 282, 646, 762, 1083
BseGI GGATG 4 cut(s) 198, 527, 1158, 1264
BseLI CCNNNNNNNGG 5 cut(s) 223, 1042, 1043, 1127, 1329
BseMI GCAATG 3 cut(s) 27, 950, 1099
BseMII CTCAG 2 cut(s) 1163, 1316
BseNI ACTGG 2 cut(s) 716, 873
BseXI GCAGC 4 cut(s) 920, 934, 1034, 1356
BseYI CCCAGC 1 cut(s) 1005
BsgI GTGCAG 2 cut(s) 400, 722
Bsh1236I CGCG 1 cut(s) 728
BshFI GGCC 5 cut(s) 105, 216, 632, 733, 761
BshNI GGYRCC 2 cut(s) 441, 739
BsiHKAI GWGCWC 2 cut(s) 758, 888
BsiHKCI CYCGRG 2 cut(s) 221, 555
BsiSI CCGG 3 cut(s) 455, 734, 1105
BslI CCNNNNNNNGG 5 cut(s) 223, 1042, 1043, 1127, 1329
BsmAI GTCTC 4 cut(s) 227, 1080, 1125, 1194
BsmBI CGTCTC 1 cut(s) 1194
BsmI GAATGC 1 cut(s) 842
BsnI GGCC 5 cut(s) 105, 216, 632, 733, 761
Bso31I GGTCTC 3 cut(s) 227, 1080, 1125
BsoBI CYCGRG 2 cut(s) 221, 555
Bsp119I TTCGAA 2 cut(s) 672, 966
Bsp1286I GDGCHC 3 cut(s) 428, 758, 888
Bsp143I GATC 5 cut(s) 172, 493, 1137, 1168, 1360
Bsp19I CCATGG 1 cut(s) 282
BspACI CCGC 6 cut(s) 230, 279, 541, 728, 743, 1121
BspANI GGCC 5 cut(s) 105, 216, 632, 733, 761
BspCNI CTCAG 2 cut(s) 1164, 1315
BspFNI CGCG 1 cut(s) 728
BspLI GGNNCC 4 cut(s) 443, 507, 732, 741
BspT104I TTCGAA 2 cut(s) 672, 966
BspT107I GGYRCC 2 cut(s) 441, 739
BspTNI GGTCTC 3 cut(s) 227, 1080, 1125
BsrDI GCAATG 3 cut(s) 27, 950, 1099
BsrFI RCCGGY 2 cut(s) 733, 1104
BsrI ACTGG 2 cut(s) 716, 873
BssAI RCCGGY 2 cut(s) 733, 1104
BssECI CCNNGG 6 cut(s) 41, 222, 282, 646, 762, 1083
BssMI GATC 5 cut(s) 172, 493, 1137, 1168, 1360
BssT1I CCWWGG 3 cut(s) 282, 646, 762
Bst2UI CCWGG 2 cut(s) 43, 581
Bst4CI ACNGT 5 cut(s) 375, 480, 858, 1001, 1064
Bst6I CTCTTC 3 cut(s) 762, 774, 963
BstAPI GCANNNNNTGC 1 cut(s) 1378
BstBI TTCGAA 2 cut(s) 672, 966
BstC8I GCNNGC 1 cut(s) 842
BstDEI CTNAG 3 cut(s) 449, 1172, 1302
BstDSI CCRYGG 1 cut(s) 282
BstENI CCTNNNNNAGG 1 cut(s) 221
BstF5I GGATG 4 cut(s) 198, 527, 1158, 1264
BstFNI CGCG 1 cut(s) 728
BstH2I RGCGCY 1 cut(s) 914
BstHHI GCGC 1 cut(s) 913
BstKTI GATC 5 cut(s) 175, 496, 1140, 1171, 1363
BstMAI GTCTC 4 cut(s) 227, 1080, 1125, 1194
BstMBI GATC 5 cut(s) 172, 493, 1137, 1168, 1360
BstMWI GCNNNNNNNGC 6 cut(s) 423, 1120, 1218, 1247, 1298, 1378
BstNI CCWGG 2 cut(s) 43, 581
BstNSI RCATGY 1 cut(s) 663
BstSCI CCNGG 3 cut(s) 41, 453, 579
BstUI CGCG 1 cut(s) 728
BstV1I GCAGC 4 cut(s) 920, 934, 1034, 1356
BstV2I GAAGAC 2 cut(s) 334, 887
BsuI GTATCC 1 cut(s) 48
BsuRI GGCC 5 cut(s) 105, 216, 632, 733, 761
BtgI CCRYGG 1 cut(s) 282
BtgZI GCGATG 1 cut(s) 1103
BtrI CACGTC 3 cut(s) 208, 271, 1126
BtsCI GGATG 4 cut(s) 198, 527, 1158, 1264
BtsI GCAGTG 1 cut(s) 237
BtsIMutI CAGTG 5 cut(s) 237, 723, 1006, 1060, 1310
Cac8I GCNNGC 1 cut(s) 842
CaiI CAGNNNCTG 2 cut(s) 1005, 1369
CfoI GCGC 1 cut(s) 913
Cfr10I RCCGGY 2 cut(s) 733, 1104
Cfr13I GGNCC 3 cut(s) 214, 286, 731
Csp6I GTAC 3 cut(s) 442, 740, 1345
CviAII CATG 6 cut(s) 34, 283, 397, 548, 660, 1298
CviQI GTAC 3 cut(s) 442, 740, 1345
DdeI CTNAG 3 cut(s) 449, 1172, 1302
DpnI GATC 5 cut(s) 174, 495, 1139, 1170, 1362
DpnII GATC 5 cut(s) 172, 493, 1137, 1168, 1360
EaeI YGGCCR 1 cut(s) 630
Eam1104I CTCTTC 3 cut(s) 762, 774, 963
EarI CTCTTC 3 cut(s) 762, 774, 963
EciI GGCGGA 1 cut(s) 245
Ecl136II GAGCTC 2 cut(s) 756, 886
Eco130I CCWWGG 3 cut(s) 282, 646, 762
Eco24I GRGCYC 3 cut(s) 428, 758, 888
Eco31I GGTCTC 3 cut(s) 227, 1080, 1125
Eco47I GGWCC 1 cut(s) 286
Eco47III AGCGCT 1 cut(s) 912
Eco53kI GAGCTC 2 cut(s) 756, 886
Eco57I CTGAAG 3 cut(s) 866, 978, 1049
Eco88I CYCGRG 2 cut(s) 221, 555
EcoICRI GAGCTC 2 cut(s) 756, 886
EcoNI CCTNNNNNAGG 1 cut(s) 221
EcoO109I RGGNCCY 1 cut(s) 214
EcoRI GAATTC 1 cut(s) 668
EcoRII CCWGG 2 cut(s) 41, 579
EcoT14I CCWWGG 3 cut(s) 282, 646, 762
EcoT38I GRGCYC 3 cut(s) 428, 758, 888
ErhI CCWWGG 3 cut(s) 282, 646, 762
Esp3I CGTCTC 1 cut(s) 1194
FaeI CATG 6 cut(s) 37, 286, 400, 551, 663, 1301
FalI AAGNNNNNCTT 2 cut(s) 771, 803
FatI CATG 6 cut(s) 33, 282, 396, 547, 659, 1297
FauI CCCGC 1 cut(s) 721
FbaI TGATCA 1 cut(s) 1360
Fnu4HI GCNGC 5 cut(s) 909, 948, 1048, 1121, 1370
FokI GGATG 4 cut(s) 185, 514, 1165, 1271
FriOI GRGCYC 3 cut(s) 428, 758, 888
Fsp4HI GCNGC 5 cut(s) 909, 948, 1048, 1121, 1370
FspBI CTAG 4 cut(s) 92, 500, 695, 1079
GlaI GCGC 1 cut(s) 912
GluI GCNGC 5 cut(s) 909, 948, 1048, 1121, 1370
GsaI CCCAGC 1 cut(s) 1009
HaeII RGCGCY 1 cut(s) 914
HaeIII GGCC 5 cut(s) 105, 216, 632, 733, 761
HapII CCGG 3 cut(s) 455, 734, 1105
HhaI GCGC 1 cut(s) 913
Hin1II CATG 6 cut(s) 37, 286, 400, 551, 663, 1301
Hin6I GCGC 1 cut(s) 911
HinP1I GCGC 1 cut(s) 911
HinfI GANTC 6 cut(s) 111, 721, 879, 930, 935, 1057
HpaII CCGG 3 cut(s) 455, 734, 1105
HphI GGTGA 5 cut(s) 172, 187, 268, 988, 1175
Hpy166II GTNNAC 2 cut(s) 409, 997
Hpy188I TCNGA 6 cut(s) 320, 753, 830, 1130, 1173, 1365
Hpy188III TCNNGA 4 cut(s) 221, 500, 655, 1192
Hpy8I GTNNAC 2 cut(s) 409, 997
Hpy99I CGWCG 4 cut(s) 212, 248, 275, 1130
HpyCH4III ACNGT 5 cut(s) 375, 480, 858, 1001, 1064
HpyCH4IV ACGT 6 cut(s) 207, 246, 270, 512, 1125, 1343
HpyF10VI GCNNNNNNNGC 6 cut(s) 423, 1120, 1218, 1247, 1298, 1378
HpyF3I CTNAG 3 cut(s) 449, 1172, 1302
HpySE526I ACGT 6 cut(s) 207, 246, 270, 512, 1125, 1343
Hsp92II CATG 6 cut(s) 37, 286, 400, 551, 663, 1301
HspAI GCGC 1 cut(s) 911
KpnI GGTACC 2 cut(s) 445, 743
Ksp22I TGATCA 1 cut(s) 1360
Kzo9I GATC 5 cut(s) 172, 493, 1137, 1168, 1360
LmnI GCTCC 2 cut(s) 151, 511
Lsp1109I GCAGC 4 cut(s) 920, 934, 1034, 1356
LweI GCATC 2 cut(s) 631, 1381
MaeI CTAG 4 cut(s) 92, 500, 695, 1079
MaeII ACGT 6 cut(s) 207, 246, 270, 512, 1125, 1343
MaeIII GTNAC 5 cut(s) 8, 274, 430, 571, 1054
MalI GATC 5 cut(s) 174, 495, 1139, 1170, 1362
MboI GATC 5 cut(s) 172, 493, 1137, 1168, 1360
MhlI GDGCHC 3 cut(s) 428, 758, 888
MluCI AATT 4 cut(s) 303, 311, 334, 668
MlyI GAGTC 2 cut(s) 944, 1051
MmeI TCCRAC 3 cut(s) 129, 1108, 1347
MseI TTAA 1 cut(s) 1390
MslI CAYNNNNRTG 2 cut(s) 32, 281
MspI CCGG 3 cut(s) 455, 734, 1105
MspR9I CCNGG 3 cut(s) 43, 455, 581
Mva1269I GAATGC 1 cut(s) 842
MvaI CCWGG 2 cut(s) 43, 581
MvnI CGCG 1 cut(s) 728
MwoI GCNNNNNNNGC 6 cut(s) 423, 1120, 1218, 1247, 1298, 1378
NciI CCSGG 1 cut(s) 455
NcoI CCATGG 1 cut(s) 282
NdeII GATC 5 cut(s) 172, 493, 1137, 1168, 1360
NlaIII CATG 6 cut(s) 37, 286, 400, 551, 663, 1301
NlaIV GGNNCC 4 cut(s) 443, 507, 732, 741
NmeAIII GCCGAG 3 cut(s) 121, 737, 1108
NmuCI GTSAC 5 cut(s) 8, 274, 430, 571, 1054
NspI RCATGY 1 cut(s) 663
NspV TTCGAA 2 cut(s) 672, 966
PaeR7I CTCGAG 1 cut(s) 555
PciI ACATGT 1 cut(s) 659
PcsI WCGNNNNNNNCGW 2 cut(s) 896, 939
PctI GAATGC 1 cut(s) 842
PfeI GAWTC 4 cut(s) 111, 721, 879, 930
PkrI GCNGC 5 cut(s) 910, 949, 1049, 1122, 1371
PleI GAGTC 2 cut(s) 943, 1051
PpsI GAGTC 2 cut(s) 943, 1051
PscI ACATGT 1 cut(s) 659
Psp124BI GAGCTC 2 cut(s) 758, 888
Psp6I CCWGG 2 cut(s) 41, 579
PspFI CCCAGC 1 cut(s) 1005
PspGI CCWGG 2 cut(s) 41, 579
PspN4I GGNNCC 4 cut(s) 443, 507, 732, 741
PspPI GGNCC 3 cut(s) 214, 286, 731
PspXI VCTCGAGB 1 cut(s) 555
PstNI CAGNNNCTG 2 cut(s) 1005, 1369
RsaI GTAC 3 cut(s) 443, 741, 1346
RsaNI GTAC 3 cut(s) 442, 740, 1345
RseI CAYNNNNRTG 2 cut(s) 32, 281
SacI GAGCTC 2 cut(s) 758, 888
SaqAI TTAA 1 cut(s) 1390
SatI GCNGC 5 cut(s) 909, 948, 1048, 1121, 1370
Sau3AI GATC 5 cut(s) 172, 493, 1137, 1168, 1360
Sau96I GGNCC 3 cut(s) 214, 286, 731
SchI GAGTC 2 cut(s) 944, 1051
ScrFI CCNGG 3 cut(s) 43, 455, 581
SduI GDGCHC 3 cut(s) 428, 758, 888
SfaNI GCATC 2 cut(s) 631, 1381
Sfr274I CTCGAG 1 cut(s) 555
SfuI TTCGAA 2 cut(s) 672, 966
SinI GGWCC 1 cut(s) 286
SlaI CTCGAG 1 cut(s) 555
SmiMI CAYNNNNRTG 2 cut(s) 32, 281
SmlI CTYRAG 2 cut(s) 555, 1334
SmoI CTYRAG 2 cut(s) 555, 1334
Sse9I AATT 4 cut(s) 303, 311, 334, 668
SsiI CCGC 6 cut(s) 230, 279, 541, 728, 743, 1121
SspMI CTAG 4 cut(s) 92, 500, 695, 1079
SstI GAGCTC 2 cut(s) 758, 888
StyD4I CCNGG 3 cut(s) 41, 453, 579
StyI CCWWGG 3 cut(s) 282, 646, 762
TaaI ACNGT 5 cut(s) 375, 480, 858, 1001, 1064
TaiI ACGT 6 cut(s) 210, 249, 273, 515, 1128, 1346
TaqI TCGA 8 cut(s) 210, 556, 672, 882, 933, 942, 966, 1191
TaqII GACCGA 1 cut(s) 27
TasI AATT 4 cut(s) 303, 311, 334, 668
TauI GCSGC 1 cut(s) 1123
TfiI GAWTC 4 cut(s) 111, 721, 879, 930
Tru1I TTAA 1 cut(s) 1390
Tru9I TTAA 1 cut(s) 1390
TscAI CASTG 5 cut(s) 244, 723, 1006, 1067, 1310
TseFI GTSAC 5 cut(s) 8, 274, 430, 571, 1054
TseI GCWGC 4 cut(s) 908, 947, 1047, 1369
Tsp45I GTSAC 5 cut(s) 8, 274, 430, 571, 1054
TspDTI ATGAA 5 cut(s) 413, 516, 564, 722, 1109
TspGWI ACGGA 1 cut(s) 888
TspRI CASTG 5 cut(s) 244, 723, 1006, 1067, 1310
VpaK11BI GGWCC 1 cut(s) 286
XagI CCTNNNNNAGG 1 cut(s) 221
XapI RAATTY 3 cut(s) 311, 334, 668
XbaI TCTAGA 1 cut(s) 499
XceI RCATGY 1 cut(s) 663
XhoI CTCGAG 1 cut(s) 555
XspI CTAG 4 cut(s) 92, 500, 695, 1079
Using CommOnly database (standard laboratory enzymes). Scanned on CDS sequence.