Basic Information

Type: gene

Biological Identity

Species

rosa_samantha

Chromosome

Chr4B

Orthogroup ID

OG0004299

Physical Location & Seq

Genomic Coordinates Reverse (-)

56622355 .. 56629100

Gene Structure 6746 bp

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UTR

Exon/CDS

Intron

Transcript / Protein ID

Rh4BG420600.1

Sequence Viewer

CDS Protein Promoter (2k)

Length: 1347 bp

ATGGCTGCTCGGAGGCTCTCATCTCTGTTGTCTCGTTCCTTCACTTCTGCTTCTTCCTTTTCTAAAGGGAGGCCTTCTTCTGTAAGTAGAGGAATCGGTAAATATAGCACTGTTGCTGCTGCTGCTGCTGCTGAGGACCCAATCATTCCATCTGTTAAAGTAGATTACACTAAGCTTCTGATTAATGGACAGTTTGTAGATGCAGCGTCAGAAAAAACATTTCCCACATTGGACCCTCGGACTGGGGATGTGATTGCTCATGTTGCTGAAGGTGATGCTAAAGATGTAAATCGGGCAGTTGCTGCTGCTAGAAAAGCATTTGACGAAGGACCATGGCCAAAGATGACAGCATATGAAAGATCTCGGGTCCTTTTCCGCTTTGCTGATTTGATTGAGAAGCATAATGACGAACTAGCGGCACTTGAGACTTGGGATAATGGGAAGCCATTTGAACAATCTGCTAAAATTGAGATACCAATGCTCGCACGTTTCTTTCGGTATTATGCTGGATATGCGGATAAGATTCATGGTCTCACTGTTCCAGCCGATGGACAATATCATGTGCAAACCTTGCATGAACCTATAGGTGTTGCAGGTCAGATTATTCCATGGAACTTCCCTCTTCTCATGTTTGCTTGGAAGGTTGCACCTGCTCTAGCATGTGGCAACACCATTGTTCTGAAGACAGCCGAGCAGACACCATTGTCTGCTCTCTATGCAGCACAGCTGTTGCATGAGGCCGGACTGCCTCCAGGTGCTTTGAATGTAGTTTCTGGTTTTGGTCCAACTGCAGGTGCCGCAGTTGCTAGTCATATGGACATCGATAAGGTTGCTTTTACTGGATCAACTGCTACTGGTAAAATTGTACTTGAACTGGCTGCTAAAAGCAATCTTAAGACAGTAACTTTGGAGCTTGGTGGCAAATCTCCATTCATTGTATGTGAGGATGCTGATGTAGACAAGGCTGTTGAGTTGGCACATTTTGCTCTATTCTTTAATCAGGGACAATGTTGCTGTGCTGGTTCTCGTACTTTTGTACATGAACGTGTGTATGATGAGTTTGTAGAGAAAGCAAAGGCACGTGCTGCGAGACGTGTAGTTGGTGATCCATTCAAGGGGGGCATTGAGCAAGGTCCTCAGATTGATTCAGACCAATTTGAGAAGATCCTGAAGTACATTGATTATGGTATTAAAGGTGGAGCGACACTGGAAACAGGAGGAGAACGATTTGGCTCAAAAGGTTACTATATTCAGCCCACTGTATTCTCAAATGTGAAGGTATTATTCTACATTGCACCTCATAACCATATTCACCAAGAATTCAAAGCTTCAATATGGTTGATCTGA

Functional Annotation

Gene Ontology

Molecular Function
Biological Process
Cellular Component

GO:0000166 GO:0003674 GO:0003824 GO:0004029 GO:0005488 GO:0005524 GO:0005575 GO:0005622 GO:0005623 GO:0005737 GO:0005739 GO:0008144 GO:0008150 GO:0008152 GO:0009507 GO:0009536 GO:0010035 GO:0010038 GO:0016491 GO:0016620 GO:0016903 GO:0017076 GO:0030554 GO:0032553 GO:0032555 GO:0032559 GO:0035639 GO:0036094 GO:0042221 GO:0043167 GO:0043168 GO:0043226 GO:0043227 GO:0043229 GO:0043231 GO:0044424 GO:0044444 GO:0044464 GO:0046686 GO:0050896 GO:0055114 GO:0097159 GO:0097367 GO:1901265 GO:1901363

KEGG Pathways

Metabolic & Signaling

ko00010 ko00053 ko00071 ko00280 ko00310 ko00330 ko00340 ko00380 ko00410 ko00561 ko00620 ko00625 ko00903 ko00981 ko01100 ko01110 ko01120 ko01130 map00010 map00053 map00071 map00280 map00310 map00330 map00340 map00380 map00410 map00561 map00620 map00625 map00903 map00981 map01100 map01110 map01120 map01130

Pfam Domains

Protein Families

Aldedh

Protein Analysis

448

Amino Acids

48.6

Weight (kDa)

7.19

Isoelectric Point (pI)

29.3

Instability Index

Protein Domains (Pfam)

Domain Name	Pfam ID	Position	E-value	Description
Aldedh	PF00171	65 - 427	1.5e-134	Aldehyde dehydrogenase family

Hydrophobicity Profile (Kyte-Doolittle)

AI Structure Prediction Report

Calculating structure properties...

Genomic Context

Gene Family Tree

Species Filter

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Align Labels (Cladogram)

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Publication-ready

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Orthologous Genes (Group: OG0004299)

Species	Orthologous Gene IDs
arabidopsis_thaliana	AT1G23800 AT1G23800
fragaria_vesca	FvH4_4g34660 FvH4_4g34660 FvH4_4g34660 FvH4_4g34660 FvH4_4g34660 FvH4_4g34660 FvH4_4g34660 FvH4_4g34660 FvH4_4g34660 FvH4_4g34660 FvH4_4g34660 FvH4_4g34660 FvH4_4g34660 FvH4_4g34660
malus_domestica	MD13G1090600.v1.1 MD13G1090800.v1.1
prunus_persica	Prupe.1G255200_v2.0.a1 Prupe.1G255200_v2.0.a1
pyrus_communis	pycom13g07880 pycom13g07910
rosa_chinensis	RchiOBHm_Chr4g0444001
rosa_laevigata	RLG00000005868
rosa_multiflora	Rmu_sc0015522.1_g000013
rosa_roxburghii	Rroxscaffold_5G00384570
rosa_rugosa	Rorug04G0348900
rosa_samantha	Rh4AG409500 Rh4BG420600 Rh4CG434800 Rh4DG415600
rosa_wichuraiana	Rw4G035190

Restriction Enzyme Sites

1 / 10

Enzyme	Recognition Site	Cut Count	Positions (bp)
AarI	CACCTGC	2 cut(s)	658, 782
AasI	GACNNNNNNGTC	1 cut(s)	703
Acc36I	ACCTGC	3 cut(s)	584, 658, 782
AccB1I	GGYRCC	1 cut(s)	794
AccB7I	CCANNNNNTGG	1 cut(s)	548
AccI	GTMKAC	1 cut(s)	957
AciI	CCGC	4 cut(s)	376, 416, 515, 798
AclWI	GGATC	3 cut(s)	850, 1100, 1159
AcoI	YGGCCR	1 cut(s)	335
AcsI	RAATTY	1 cut(s)	1319
AcuI	CTGAAG	3 cut(s)	288, 701, 1190
AcvI	CACGTG	1 cut(s)	1082
AfaI	GTAC	4 cut(s)	867, 1030, 1038, 1175
AfiI	CCNNNNNNNGG	4 cut(s)	242, 548, 791, 1115
AflII	CTTAAG	1 cut(s)	893
AflIII	ACRYGT	2 cut(s)	1045, 1093
AgsI	TTSAA	6 cut(s)	452, 763, 872, 1114, 1324, 1332
AjiI	CACGTC	1 cut(s)	1094
AjnI	CCWGG	1 cut(s)	751
AluBI	AGCT	4 cut(s)	175, 727, 913, 1328
AluI	AGCT	4 cut(s)	175, 727, 913, 1328
Alw26I	GTCTC	4 cut(s)	36, 419, 536, 1084
AlwI	GGATC	3 cut(s)	850, 1100, 1159
AlwNI	CAGNNNCTG	1 cut(s)	302
Ama87I	CYCGRG	1 cut(s)	363
AoxI	GGCC	3 cut(s)	71, 335, 738
ApoI	RAATTY	1 cut(s)	1319
ArsI	GACNNNNNNTTYG	2 cut(s)	889, 921
AseI	ATTAAT	1 cut(s)	183
AspS9I	GGNCC	6 cut(s)	136, 232, 329, 367, 782, 1133
AsuHPI	GGTGA	3 cut(s)	284, 1115, 1304
AvaI	CYCGRG	1 cut(s)	363
AvaII	GGWCC	6 cut(s)	136, 232, 329, 367, 782, 1133
BalI	TGGCCA	1 cut(s)	337
BanI	GGYRCC	1 cut(s)	794
BbrPI	CACGTG	1 cut(s)	1082
BbsI	GAAGAC	1 cut(s)	689
BbvCI	CCTCAGC	1 cut(s)	132
BccI	CCATC	2 cut(s)	157, 542
BcgI	CGANNNNNNTGC	2 cut(s)	812, 846
BciT130I	CCWGG	1 cut(s)	753
BcoDI	GTCTC	4 cut(s)	36, 419, 536, 1084
BfaI	CTAG	4 cut(s)	309, 413, 656, 807
BfmI	CTRYAG	2 cut(s)	582, 789
BfrI	CTTAAG	1 cut(s)	893
BfuAI	ACCTGC	3 cut(s)	584, 658, 782
BglII	AGATCT	1 cut(s)	359
Bme1390I	CCNGG	1 cut(s)	753
Bme18I	GGWCC	6 cut(s)	136, 232, 329, 367, 782, 1133
BmeT110I	CYCGRG	1 cut(s)	363
BmgBI	CACGTC	1 cut(s)	1094
BmgT120I	GGNCC	6 cut(s)	136, 232, 329, 367, 782, 1133
BmiI	GGNNCC	4 cut(s)	138, 234, 368, 796
BmrFI	CCNGG	1 cut(s)	753
BmrI	ACTGGG	1 cut(s)	252
BmsI	GCATC	3 cut(s)	190, 265, 937
BmuI	ACTGGG	1 cut(s)	252
BpiI	GAAGAC	1 cut(s)	689
BpmI	CTGGAG	1 cut(s)	735
Bpu10I	CCTNAGC	1 cut(s)	132
BpuEI	CTTGAG	1 cut(s)	443
Bsa29I	ATCGAT	1 cut(s)	822
BsaAI	YACGTR	1 cut(s)	1082
BsaBI	GATNNNNATC	1 cut(s)	288
BsaI	GGTCTC	1 cut(s)	536
BsaJI	CCNNGG	3 cut(s)	236, 332, 608
Bsc4I	CCNNNNNNNGG	4 cut(s)	242, 548, 791, 1115
Bse1I	ACTGG	5 cut(s)	247, 844, 859, 879, 1212
Bse3DI	GCAATG	1 cut(s)	1290
Bse8I	GATNNNNATC	1 cut(s)	288
BseBI	CCWGG	1 cut(s)	753
BseCI	ATCGAT	1 cut(s)	822
BseDI	CCNNGG	3 cut(s)	236, 332, 608
BseGI	GGATG	2 cut(s)	253, 952
BseJI	GATNNNNATC	1 cut(s)	288
BseLI	CCNNNNNNNGG	4 cut(s)	242, 548, 791, 1115
BseMI	GCAATG	1 cut(s)	1290
BseMII	CTCAG	2 cut(s)	123, 1151
BseNI	ACTGG	5 cut(s)	247, 844, 859, 879, 1212
BseRI	GAGGAG	1 cut(s)	1233
BshFI	GGCC	3 cut(s)	73, 337, 740
BshNI	GGYRCC	1 cut(s)	794
BshVI	ATCGAT	1 cut(s)	822
BsiHKCI	CYCGRG	1 cut(s)	363
BsiSI	CCGG	1 cut(s)	741
BslFI	GGGAC	1 cut(s)	1017
BslI	CCNNNNNNNGG	4 cut(s)	242, 548, 791, 1115
BsmAI	GTCTC	4 cut(s)	36, 419, 536, 1084
BsmBI	CGTCTC	1 cut(s)	1084
BsmFI	GGGAC	1 cut(s)	1017
BsnI	GGCC	3 cut(s)	73, 337, 740
Bso31I	GGTCTC	1 cut(s)	536
BsoBI	CYCGRG	1 cut(s)	363
Bsp1407I	TGTACA	1 cut(s)	1036
Bsp143I	GATC	5 cut(s)	359, 842, 1105, 1164, 1341
Bsp19I	CCATGG	2 cut(s)	332, 608
BspACI	CCGC	4 cut(s)	376, 416, 515, 798
BspANI	GGCC	3 cut(s)	73, 337, 740
BspCNI	CTCAG	2 cut(s)	124, 1150
BspDI	ATCGAT	1 cut(s)	822
BspLI	GGNNCC	4 cut(s)	138, 234, 368, 796
BspMAI	CTGCAG	1 cut(s)	793
BspMI	ACCTGC	3 cut(s)	584, 658, 782
BspPI	GGATC	3 cut(s)	850, 1100, 1159
BspT107I	GGYRCC	1 cut(s)	794
BspTI	CTTAAG	1 cut(s)	893
BspTNI	GGTCTC	1 cut(s)	536
BsrDI	GCAATG	1 cut(s)	1290
BsrGI	TGTACA	1 cut(s)	1036
BsrI	ACTGG	5 cut(s)	247, 844, 859, 879, 1212
BssECI	CCNNGG	3 cut(s)	236, 332, 608
BssMI	GATC	5 cut(s)	359, 842, 1105, 1164, 1341
BssT1I	CCWWGG	2 cut(s)	332, 608
Bst2UI	CCWGG	1 cut(s)	753
Bst4CI	ACNGT	5 cut(s)	112, 192, 538, 901, 1261
Bst6I	CTCTTC	1 cut(s)	627
BstAFI	CTTAAG	1 cut(s)	893
BstAPI	GCANNNNNTGC	4 cut(s)	302, 571, 983, 1085
BstAUI	TGTACA	1 cut(s)	1036
BstBAI	YACGTR	1 cut(s)	1082
BstC8I	GCNNGC	1 cut(s)	483
BstDEI	CTNAG	3 cut(s)	132, 171, 1137
BstDSI	CCRYGG	2 cut(s)	332, 608
BstF5I	GGATG	2 cut(s)	253, 952
BstKTI	GATC	5 cut(s)	362, 845, 1108, 1167, 1344
BstMAI	GTCTC	4 cut(s)	36, 419, 536, 1084
BstMBI	GATC	5 cut(s)	359, 842, 1105, 1164, 1341
BstNI	CCWGG	1 cut(s)	753
BstNSI	RCATGY	1 cut(s)	663
BstSCI	CCNGG	1 cut(s)	751
BstSFI	CTRYAG	2 cut(s)	582, 789
BstV2I	GAAGAC	1 cut(s)	689
BstX2I	RGATCY	2 cut(s)	359, 1164
BstYI	RGATCY	2 cut(s)	359, 1164
Bsu15I	ATCGAT	1 cut(s)	822
BsuRI	GGCC	3 cut(s)	73, 337, 740
BsuTUI	ATCGAT	1 cut(s)	822
BtgI	CCRYGG	2 cut(s)	332, 608
BtrI	CACGTC	1 cut(s)	1094
BtsCI	GGATG	2 cut(s)	253, 952
BtsIMutI	CAGTG	4 cut(s)	108, 534, 1205, 1257
BveI	ACCTGC	3 cut(s)	584, 658, 782
Cac8I	GCNNGC	1 cut(s)	483
CaiI	CAGNNNCTG	1 cut(s)	302
Cfr13I	GGNCC	6 cut(s)	136, 232, 329, 367, 782, 1133
ClaI	ATCGAT	1 cut(s)	822
CseI	GACGC	1 cut(s)	195
Csp6I	GTAC	4 cut(s)	866, 1029, 1037, 1174
CviQI	GTAC	4 cut(s)	866, 1029, 1037, 1174
DdeI	CTNAG	3 cut(s)	132, 171, 1137
DpnI	GATC	5 cut(s)	361, 844, 1107, 1166, 1343
DpnII	GATC	5 cut(s)	359, 842, 1105, 1164, 1341
DrdI	GACNNNNNNGTC	1 cut(s)	703
DseDI	GACNNNNNNGTC	1 cut(s)	703
EaeI	YGGCCR	1 cut(s)	335
Eam1104I	CTCTTC	1 cut(s)	627
EarI	CTCTTC	1 cut(s)	627
Eco130I	CCWWGG	2 cut(s)	332, 608
Eco147I	AGGCCT	1 cut(s)	73
Eco31I	GGTCTC	1 cut(s)	536
Eco47I	GGWCC	6 cut(s)	136, 232, 329, 367, 782, 1133
Eco57I	CTGAAG	3 cut(s)	288, 701, 1190
Eco72I	CACGTG	1 cut(s)	1082
Eco88I	CYCGRG	1 cut(s)	363
EcoO109I	RGGNCCY	3 cut(s)	136, 367, 1133
EcoRI	GAATTC	1 cut(s)	1319
EcoRII	CCWGG	1 cut(s)	751
EcoT14I	CCWWGG	2 cut(s)	332, 608
ErhI	CCWWGG	2 cut(s)	332, 608
Esp3I	CGTCTC	1 cut(s)	1084
FaqI	GGGAC	1 cut(s)	1017
FauNDI	CATATG	2 cut(s)	352, 813
FblI	GTMKAC	1 cut(s)	957
FokI	GGATG	2 cut(s)	260, 959
FspBI	CTAG	4 cut(s)	309, 413, 656, 807
GsuI	CTGGAG	1 cut(s)	735
HaeIII	GGCC	3 cut(s)	73, 337, 740
HapII	CCGG	1 cut(s)	741
HgaI	GACGC	1 cut(s)	195
HindIII	AAGCTT	2 cut(s)	173, 1326
HinfI	GANTC	3 cut(s)	93, 523, 1145
HpaII	CCGG	1 cut(s)	741
HphI	GGTGA	3 cut(s)	284, 1115, 1304
Hpy166II	GTNNAC	1 cut(s)	958
Hpy188I	TCNGA	9 cut(s)	12, 180, 211, 240, 600, 681, 1140, 1150, 1346
Hpy188III	TCNNGA	1 cut(s)	1168
Hpy8I	GTNNAC	1 cut(s)	958
HpyAV	CCTTC	6 cut(s)	49, 84, 263, 320, 634, 1270
HpyCH4III	ACNGT	5 cut(s)	112, 192, 538, 901, 1261
HpyCH4IV	ACGT	4 cut(s)	487, 1045, 1081, 1093
HpyCH4V	TGCA	9 cut(s)	203, 565, 574, 593, 647, 719, 733, 791, 1295
HpyF3I	CTNAG	3 cut(s)	132, 171, 1137
HpySE526I	ACGT	4 cut(s)	487, 1045, 1081, 1093
Kzo9I	GATC	5 cut(s)	359, 842, 1105, 1164, 1341
LmnI	GCTCC	2 cut(s)	910, 1199
LweI	GCATC	3 cut(s)	190, 265, 937
MaeI	CTAG	4 cut(s)	309, 413, 656, 807
MaeII	ACGT	4 cut(s)	487, 1045, 1081, 1093
MaeIII	GTNAC	2 cut(s)	901, 1241
MalI	GATC	5 cut(s)	361, 844, 1107, 1166, 1343
MboI	GATC	5 cut(s)	359, 842, 1105, 1164, 1341
MboII	GAAGA	5 cut(s)	45, 69, 614, 694, 1174
MflI	RGATCY	2 cut(s)	359, 1164
MlsI	TGGCCA	1 cut(s)	337
MluCI	AATT	4 cut(s)	465, 861, 1154, 1319
MluNI	TGGCCA	1 cut(s)	337
MmeI	TCCRAC	1 cut(s)	809
Mox20I	TGGCCA	1 cut(s)	337
MscI	TGGCCA	1 cut(s)	337
MseI	TTAA	5 cut(s)	156, 183, 894, 996, 1191
MslI	CAYNNNNRTG	1 cut(s)	1044
Msp20I	TGGCCA	1 cut(s)	337
MspA1I	CMGCKG	1 cut(s)	727
MspCI	CTTAAG	1 cut(s)	893
MspI	CCGG	1 cut(s)	741
MspR9I	CCNGG	1 cut(s)	753
MvaI	CCWGG	1 cut(s)	753
NcoI	CCATGG	2 cut(s)	332, 608
NdeI	CATATG	2 cut(s)	352, 813
NdeII	GATC	5 cut(s)	359, 842, 1105, 1164, 1341
NlaIV	GGNNCC	4 cut(s)	138, 234, 368, 796
NmeAIII	GCCGAG	1 cut(s)	715
NspI	RCATGY	1 cut(s)	663
PaqCI	CACCTGC	2 cut(s)	658, 782
PceI	AGGCCT	1 cut(s)	73
PfeI	GAWTC	3 cut(s)	93, 523, 1145
PflMI	CCANNNNNTGG	1 cut(s)	548
PmaCI	CACGTG	1 cut(s)	1082
PmlI	CACGTG	1 cut(s)	1082
Ppu21I	YACGTR	1 cut(s)	1082
PpuMI	RGGWCCY	3 cut(s)	136, 367, 1133
PshBI	ATTAAT	1 cut(s)	183
Psp5II	RGGWCCY	3 cut(s)	136, 367, 1133
Psp6I	CCWGG	1 cut(s)	751
PspCI	CACGTG	1 cut(s)	1082
PspGI	CCWGG	1 cut(s)	751
PspN4I	GGNNCC	4 cut(s)	138, 234, 368, 796
PspPI	GGNCC	6 cut(s)	136, 232, 329, 367, 782, 1133
PspPPI	RGGWCCY	3 cut(s)	136, 367, 1133
PstI	CTGCAG	1 cut(s)	793
PstNI	CAGNNNCTG	1 cut(s)	302
PsuI	RGATCY	2 cut(s)	359, 1164
PvuII	CAGCTG	1 cut(s)	727
RsaI	GTAC	4 cut(s)	867, 1030, 1038, 1175
RsaNI	GTAC	4 cut(s)	866, 1029, 1037, 1174
RseI	CAYNNNNRTG	1 cut(s)	1044
SaqAI	TTAA	5 cut(s)	156, 183, 894, 996, 1191
Sau3AI	GATC	5 cut(s)	359, 842, 1105, 1164, 1341
Sau96I	GGNCC	6 cut(s)	136, 232, 329, 367, 782, 1133
ScrFI	CCNGG	1 cut(s)	753
SfaNI	GCATC	3 cut(s)	190, 265, 937
SfcI	CTRYAG	2 cut(s)	582, 789
SinI	GGWCC	6 cut(s)	136, 232, 329, 367, 782, 1133
SmiMI	CAYNNNNRTG	1 cut(s)	1044
SmlI	CTYRAG	2 cut(s)	422, 893
SmoI	CTYRAG	2 cut(s)	422, 893
Sse9I	AATT	4 cut(s)	465, 861, 1154, 1319
SseBI	AGGCCT	1 cut(s)	73
SsiI	CCGC	4 cut(s)	376, 416, 515, 798
SspMI	CTAG	4 cut(s)	309, 413, 656, 807
StuI	AGGCCT	1 cut(s)	73
StyD4I	CCNGG	1 cut(s)	751
StyI	CCWWGG	2 cut(s)	332, 608
TaaI	ACNGT	5 cut(s)	112, 192, 538, 901, 1261
TaiI	ACGT	4 cut(s)	490, 1048, 1084, 1096
TaqI	TCGA	1 cut(s)	822
TasI	AATT	4 cut(s)	465, 861, 1154, 1319
TatI	WGTACW	3 cut(s)	865, 1036, 1173
TauI	GCSGC	2 cut(s)	419, 800
TfiI	GAWTC	3 cut(s)	93, 523, 1145
Tru1I	TTAA	5 cut(s)	156, 183, 894, 996, 1191
Tru9I	TTAA	5 cut(s)	156, 183, 894, 996, 1191
TscAI	CASTG	4 cut(s)	115, 541, 1212, 1264
TspDTI	ATGAA	5 cut(s)	369, 515, 591, 922, 1056
TspRI	CASTG	4 cut(s)	115, 541, 1212, 1264
Van91I	CCANNNNNTGG	1 cut(s)	548
Vha464I	CTTAAG	1 cut(s)	893
VpaK11BI	GGWCC	6 cut(s)	136, 232, 329, 367, 782, 1133
VspI	ATTAAT	1 cut(s)	183
XapI	RAATTY	1 cut(s)	1319
XceI	RCATGY	1 cut(s)	663
XmiI	GTMKAC	1 cut(s)	957
XspI	CTAG	4 cut(s)	309, 413, 656, 807

Using CommOnly database (standard laboratory enzymes). Scanned on CDS sequence.