Basic Information

Type: gene

Biological Identity

Species

rosa_samantha

Chromosome

Chr5B

Orthogroup ID

OG0018010

Physical Location & Seq

Genomic Coordinates Forward (+)

73616739 .. 73620725

Gene Structure 3987 bp

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UTR

Exon/CDS

Intron

Transcript / Protein ID

Rh5BG460300.1

Sequence Viewer

CDS Protein Promoter (2k)

Length: 1461 bp

ATGGGTGCCGGATTCTCTCTTTGCTTCCGGCCATCAAATACGATATCCAATCTCAACGACTCGTCGGAATTAGAGAACGTAAGCAAGGAAACTTACTCGCTGCCTGAGTTCAGCTTGAACCAACTTAAAGCTGCCACAGAGGGATTCTTGCCGGAGAACATTGTTTCCGACGACGGGGAGAAAGCTCCAAATGTTGTCTACAGAGGCATGCTCAAAGATGGCCGTTGGATCGCCATTAAGCGCTTCAAGTACTCGGCTTGGCCTGATTCCCTCCAATTTCTTGCTGAGGCTAGAGCTGTGGGGCAGCTGAGAAACAAACGTTTGGCGAATTTGATAGGGAGTTGCTGTGAAGGCAATGAGAGATTGCTTGTGGCTGAGTTCATGCCTAATCAAACTCTCTCTAAACATCTGTTCCATAGGGAGTCCCAGCCCATGAAAGGGACTATGAGGTTGCGAGTGGCTCTGTATCTGGCACAAGCTTTGGTGTATTGCAGTAGCAAAGGGCGGCCATTGTACCATGATCTTAATGCCTCTAGAGTCCTGTTTGATCAGAAGGGAAATCCCAGACTATCCTGCTTTGGCCTTATGAAGAGTAGCAAAGATGGCAAGAGCTACAGCTCCAACCTAGCTTACCCTCCAGAATACACCAGGACAGGAAGAGTTATACCAGAAAGTTTGATTTATAGCTTTGGGACCTTATTGCTTGATCTGCTCAGTAGCAGTCATACCCACCCAAGCCATGCAGTAGATCTCATACACGACAAAGGTTTCCTGATGCAATTGGACTCATGCTTGGTGGATCATTTCTCAAAAGATGATGGAAATGAGCTGTTGTGGTTAGCTTCACAATGCTTGCAGTTTGATCCTTGTAAGAGACCAAATCCTAAGTCTCTTGTGAATACCCTCATTCCTCTTCAGAAAGAAACAGAGGTTCCATCATTTGTTTTGATGGATATTCCACATGGAAATGTGCCTCCAAAGCAATTGTCACCTTTAGGTGAAGCTTGCTCAAGATTGGATCTTACTGCAATACATAAAGTACTGGAGATGGCTGGATACAGGGATGATGTTGAATCTGAGGCTGAAAAGGAGCCTTCTTTCGATATGTGGACTGATCCAATGCGGGAAGCACTAAATTCCAGGAAGCAGGGAGATGCTGCATTTAGAACTAAAGACTTTGGTACTGCAATAGGTTGTTACACAAAATTCATTAATAGTGGGAGCTTGGAATCCCGAACCGTGTATGTTAGACGTTGCTTGTGTTACTTGATGACTGACAAGGCAGAAGAAGCTCTTAAAGATAGTATGAATGCCCTAGTACTACACCATTTATGGCCGACTGCCTTCTATCTTCAAGCTGCTGCCCTAAAGATCCTAGGGATGGATAATGATGCAAAGGATATTCTGAAAGAAGGTGCATCATTGGAAGCCATGAAAAACAAAAAAGCTCAAATTGTTTAA

Functional Annotation

Gene Ontology

Molecular Function
Biological Process
Cellular Component

GO:0001101 GO:0003674 GO:0003824 GO:0004672 GO:0004674 GO:0004675 GO:0004888 GO:0005575 GO:0005623 GO:0005886 GO:0005911 GO:0006464 GO:0006468 GO:0006793 GO:0006796 GO:0006807 GO:0006950 GO:0006970 GO:0007154 GO:0007165 GO:0007166 GO:0007167 GO:0007178 GO:0008150 GO:0008152 GO:0009266 GO:0009409 GO:0009506 GO:0009628 GO:0009651 GO:0009719 GO:0009725 GO:0009737 GO:0009741 GO:0009742 GO:0009755 GO:0009987 GO:0010033 GO:0014070 GO:0016020 GO:0016301 GO:0016310 GO:0016740 GO:0016772 GO:0016773 GO:0019199 GO:0019538 GO:0023052 GO:0030054 GO:0032870 GO:0033993 GO:0036211 GO:0038023 GO:0042221 GO:0043170 GO:0043401 GO:0043412 GO:0044237 GO:0044238 GO:0044260 GO:0044267 GO:0044464 GO:0048545 GO:0050789 GO:0050794 GO:0050896 GO:0051716 GO:0055044 GO:0060089 GO:0065007 GO:0070887 GO:0071310 GO:0071367 GO:0071383 GO:0071396 GO:0071407 GO:0071495 GO:0071704 GO:0071944 GO:0097305 GO:0140096 GO:1901564 GO:1901700 GO:1901701

KEGG Pathways

Metabolic & Signaling

ko04075 map04075

Pfam Domains

Protein Families

Pkinase_Tyr

Protein Analysis

486

Amino Acids

54.42

Weight (kDa)

7.16

Isoelectric Point (pI)

38.19

Instability Index

Protein Domains (Pfam)

Domain Name	Pfam ID	Position	E-value	Description
Pkinase	PF00069	65 - 300	2.9e-17	Protein kinase domain
PK_Tyr_Ser-Thr	PF07714	66 - 302	1.3e-20	Protein tyrosine and serine/threonine kinase
TPR_BSK1_C	PF25575	334 - 453	1e-52	Serine/threonine-protein kinase BSK1-like, TPR repeats

Hydrophobicity Profile (Kyte-Doolittle)

AI Structure Prediction Report

Calculating structure properties...

Genomic Context

Gene Family Tree

Species Filter

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Align Labels (Cladogram)

Image

Tree File

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Publication-ready

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Orthologous Genes (Group: OG0018010)

Species	Orthologous Gene IDs
fragaria_vesca	FvH4_3g37410
rosa_chinensis	RchiOBHm_Chr5g0067291 RchiOBHm_Chr5g0067301
rosa_laevigata	RLG00000035924
rosa_multiflora	Rmu_sc0000015.1_g000009
rosa_roxburghii	Rroxscaffold_1G00013830
rosa_rugosa	Rorug05G0383800
rosa_samantha	Rh5AG443100 Rh5BG460300 Rh5DG475200
rosa_wichuraiana	Rw5G041410

Restriction Enzyme Sites

1 / 10

Enzyme	Recognition Site	Cut Count	Positions (bp)
AccB1I	GGYRCC	1 cut(s)	5
AccI	GTMKAC	1 cut(s)	198
AciI	CCGC	2 cut(s)	505, 1123
AclI	AACGTT	1 cut(s)	319
AclWI	GGATC	6 cut(s)	236, 807, 857, 1026, 1109, 1366
AcoI	YGGCCR	4 cut(s)	29, 220, 506, 1334
AcsI	RAATTY	3 cut(s)	328, 1135, 1205
AcuI	CTGAAG	1 cut(s)	899
AfaI	GTAC	5 cut(s)	251, 515, 1041, 1183, 1320
AfeI	AGCGCT	1 cut(s)	242
AfiI	CCNNNNNNNGG	4 cut(s)	174, 437, 438, 1381
AgsI	TTSAA	4 cut(s)	118, 247, 1073, 1355
AjnI	CCWGG	2 cut(s)	647, 1139
AloI	GAACNNNNNNTCC	2 cut(s)	149, 181
Alw26I	GTCTC	2 cut(s)	868, 894
AlwI	GGATC	6 cut(s)	236, 807, 857, 1026, 1109, 1366
Aor51HI	AGCGCT	1 cut(s)	242
AoxI	GGCC	6 cut(s)	29, 220, 260, 506, 580, 1334
ApeKI	GCWGC	6 cut(s)	100, 131, 304, 1157, 1358, 1361
ApoI	RAATTY	3 cut(s)	328, 1135, 1205
ArsI	GACNNNNNNTTYG	4 cut(s)	872, 904, 971, 1003
AseI	ATTAAT	1 cut(s)	1212
AspA2I	CCTAGG	1 cut(s)	1375
AspLEI	GCGC	1 cut(s)	243
AspS9I	GGNCC	1 cut(s)	693
AsuHPI	GGTGA	2 cut(s)	981, 1010
AvaII	GGWCC	1 cut(s)	693
AvrII	CCTAGG	1 cut(s)	1375
BanI	GGYRCC	1 cut(s)	5
BbvCI	CCTCAGC	1 cut(s)	285
BbvI	GCAGC	6 cut(s)	87, 118, 316, 1144, 1345, 1348
BccI	CCATC	8 cut(s)	40, 212, 596, 812, 943, 943, 1042, 1375
BceAI	ACGGC	1 cut(s)	207
BciT130I	CCWGG	2 cut(s)	649, 1141
BciVI	GTATCC	1 cut(s)	1049
BclI	TGATCA	1 cut(s)	547
BcoDI	GTCTC	2 cut(s)	868, 894
BfaI	CTAG	5 cut(s)	291, 534, 626, 1316, 1376
BfmI	CTRYAG	2 cut(s)	199, 613
BfoI	RGCGCY	1 cut(s)	244
BfuI	GTATCC	1 cut(s)	1049
BglII	AGATCT	1 cut(s)	748
BisI	GCNGC	7 cut(s)	101, 132, 305, 506, 1158, 1359, 1362
BlnI	CCTAGG	1 cut(s)	1375
BlsI	GCNGC	7 cut(s)	102, 133, 306, 507, 1159, 1360, 1363
BmcAI	AGTACT	3 cut(s)	251, 1041, 1320
Bme1390I	CCNGG	2 cut(s)	649, 1141
Bme18I	GGWCC	1 cut(s)	693
BmgT120I	GGNCC	1 cut(s)	693
BmiI	GGNNCC	4 cut(s)	7, 694, 933, 1092
BmrFI	CCNGG	2 cut(s)	649, 1141
BmsI	GCATC	4 cut(s)	765, 1144, 1381, 1427
BplI	GAGNNNNNCTC	2 cut(s)	195, 227
BpmI	CTGGAG	2 cut(s)	621, 1064
Bpu10I	CCTNAGC	1 cut(s)	285
BpuEI	CTTGAG	1 cut(s)	994
BsaI	GGTCTC	1 cut(s)	868
BsaJI	CCNNGG	1 cut(s)	1375
BsaXI	ACNNNNNCTCC	2 cut(s)	331, 361
Bsc4I	CCNNNNNNNGG	4 cut(s)	174, 437, 438, 1381
Bse1I	ACTGG	1 cut(s)	1047
Bse3DI	GCAATG	1 cut(s)	361
BseBI	CCWGG	2 cut(s)	649, 1141
BseDI	CCNNGG	1 cut(s)	1375
BseGI	GGATG	2 cut(s)	1069, 1386
BseLI	CCNNNNNNNGG	4 cut(s)	174, 437, 438, 1381
BseMI	GCAATG	1 cut(s)	361
BseMII	CTCAG	6 cut(s)	96, 276, 299, 366, 727, 1068
BseNI	ACTGG	1 cut(s)	1047
BseXI	GCAGC	6 cut(s)	87, 118, 316, 1144, 1345, 1348
BseYI	CCCAGC	1 cut(s)	426
BshFI	GGCC	6 cut(s)	31, 222, 262, 508, 582, 1336
BshNI	GGYRCC	1 cut(s)	5
BsiSI	CCGG	3 cut(s)	9, 28, 152
BslFI	GGGAC	3 cut(s)	409, 454, 706
BslI	CCNNNNNNNGG	4 cut(s)	174, 437, 438, 1381
BsmAI	GTCTC	2 cut(s)	868, 894
BsmFI	GGGAC	3 cut(s)	409, 454, 706
BsmI	GAATGC	1 cut(s)	1315
BsnI	GGCC	6 cut(s)	31, 222, 262, 508, 582, 1336
Bso31I	GGTCTC	1 cut(s)	868
BspACI	CCGC	2 cut(s)	505, 1123
BspANI	GGCC	6 cut(s)	31, 222, 262, 508, 582, 1336
BspCNI	CTCAG	6 cut(s)	97, 277, 300, 367, 726, 1069
BspLI	GGNNCC	4 cut(s)	7, 694, 933, 1092
BspPI	GGATC	6 cut(s)	236, 807, 857, 1026, 1109, 1366
BspT107I	GGYRCC	1 cut(s)	5
BspTNI	GGTCTC	1 cut(s)	868
BsrDI	GCAATG	1 cut(s)	361
BsrI	ACTGG	1 cut(s)	1047
BssECI	CCNNGG	1 cut(s)	1375
BssT1I	CCWWGG	1 cut(s)	1375
Bst2UI	CCWGG	2 cut(s)	649, 1141
Bst4CI	ACNGT	1 cut(s)	1240
Bst6I	CTCTTC	3 cut(s)	584, 652, 918
BstC8I	GCNNGC	3 cut(s)	209, 854, 1006
BstDEI	CTNAG	7 cut(s)	105, 285, 308, 375, 713, 885, 1077
BstF5I	GGATG	2 cut(s)	1069, 1386
BstH2I	RGCGCY	1 cut(s)	244
BstHHI	GCGC	1 cut(s)	243
BstMAI	GTCTC	2 cut(s)	868, 894
BstMWI	GCNNNNNNNGC	5 cut(s)	351, 603, 709, 979, 1289
BstNI	CCWGG	2 cut(s)	649, 1141
BstNSI	RCATGY	1 cut(s)	211
BstSCI	CCNGG	2 cut(s)	647, 1139
BstSFI	CTRYAG	2 cut(s)	199, 613
BstV1I	GCAGC	6 cut(s)	87, 118, 316, 1144, 1345, 1348
BstX2I	RGATCY	3 cut(s)	748, 1018, 1371
BstYI	RGATCY	3 cut(s)	748, 1018, 1371
BsuI	GTATCC	1 cut(s)	1049
BsuRI	GGCC	6 cut(s)	31, 222, 262, 508, 582, 1336
BtsCI	GGATG	2 cut(s)	1069, 1386
Cac8I	GCNNGC	3 cut(s)	209, 854, 1006
CfoI	GCGC	1 cut(s)	243
Cfr13I	GGNCC	1 cut(s)	693
Csp6I	GTAC	5 cut(s)	250, 514, 1040, 1182, 1319
CviAII	CATG	8 cut(s)	208, 382, 433, 518, 740, 789, 962, 1432
CviQI	GTAC	5 cut(s)	250, 514, 1040, 1182, 1319
DdeI	CTNAG	7 cut(s)	105, 285, 308, 375, 713, 885, 1077
EaeI	YGGCCR	4 cut(s)	29, 220, 506, 1334
Eam1104I	CTCTTC	3 cut(s)	584, 652, 918
EarI	CTCTTC	3 cut(s)	584, 652, 918
Eco130I	CCWWGG	1 cut(s)	1375
Eco31I	GGTCTC	1 cut(s)	868
Eco32I	GATATC	1 cut(s)	45
Eco47I	GGWCC	1 cut(s)	693
Eco47III	AGCGCT	1 cut(s)	242
Eco57I	CTGAAG	1 cut(s)	899
EcoO109I	RGGNCCY	1 cut(s)	693
EcoRII	CCWGG	2 cut(s)	647, 1139
EcoRV	GATATC	1 cut(s)	45
EcoT14I	CCWWGG	1 cut(s)	1375
ErhI	CCWWGG	1 cut(s)	1375
FaeI	CATG	8 cut(s)	211, 385, 436, 521, 743, 792, 965, 1435
FalI	AAGNNNNNCTT	2 cut(s)	1278, 1310
FaqI	GGGAC	3 cut(s)	409, 454, 706
FatI	CATG	8 cut(s)	207, 381, 432, 517, 739, 788, 961, 1431
FauI	CCCGC	1 cut(s)	1116
FbaI	TGATCA	1 cut(s)	547
FblI	GTMKAC	1 cut(s)	198
Fnu4HI	GCNGC	7 cut(s)	101, 132, 305, 506, 1158, 1359, 1362
FokI	GGATG	2 cut(s)	1076, 1393
Fsp4HI	GCNGC	7 cut(s)	101, 132, 305, 506, 1158, 1359, 1362
FspBI	CTAG	5 cut(s)	291, 534, 626, 1316, 1376
GlaI	GCGC	1 cut(s)	242
GluI	GCNGC	7 cut(s)	101, 132, 305, 506, 1158, 1359, 1362
GsaI	CCCAGC	1 cut(s)	430
GsuI	CTGGAG	2 cut(s)	621, 1064
HaeII	RGCGCY	1 cut(s)	244
HaeIII	GGCC	6 cut(s)	31, 222, 262, 508, 582, 1336
HapII	CCGG	3 cut(s)	9, 28, 152
HhaI	GCGC	1 cut(s)	243
Hin1II	CATG	8 cut(s)	211, 385, 436, 521, 743, 792, 965, 1435
Hin6I	GCGC	1 cut(s)	241
HinP1I	GCGC	1 cut(s)	241
HindIII	AAGCTT	2 cut(s)	477, 1002
HinfI	GANTC	9 cut(s)	12, 59, 144, 266, 422, 537, 785, 1073, 1229
HpaII	CCGG	3 cut(s)	9, 28, 152
HphI	GGTGA	2 cut(s)	981, 1010
Hpy166II	GTNNAC	2 cut(s)	199, 1110
Hpy188I	TCNGA	6 cut(s)	67, 169, 552, 918, 1078, 1407
Hpy188III	TCNNGA	5 cut(s)	534, 638, 772, 1011, 1233
Hpy8I	GTNNAC	2 cut(s)	199, 1110
Hpy99I	CGWCG	3 cut(s)	67, 173, 176
HpyAV	CCTTC	5 cut(s)	344, 547, 1104, 1354, 1406
HpyCH4III	ACNGT	1 cut(s)	1240
HpyCH4IV	ACGT	3 cut(s)	78, 319, 1252
HpyCH4V	TGCA	9 cut(s)	492, 743, 778, 856, 1028, 1160, 1187, 1394, 1418
HpyF10VI	GCNNNNNNNGC	5 cut(s)	351, 603, 709, 979, 1289
HpyF3I	CTNAG	7 cut(s)	105, 285, 308, 375, 713, 885, 1077
HpySE526I	ACGT	3 cut(s)	78, 319, 1252
Hsp92II	CATG	8 cut(s)	211, 385, 436, 521, 743, 792, 965, 1435
HspAI	GCGC	1 cut(s)	241
Ksp22I	TGATCA	1 cut(s)	547
LmnI	GCTCC	4 cut(s)	190, 623, 1090, 1221
Lsp1109I	GCAGC	6 cut(s)	87, 118, 316, 1144, 1345, 1348
LweI	GCATC	4 cut(s)	765, 1144, 1381, 1427
MaeI	CTAG	5 cut(s)	291, 534, 626, 1316, 1376
MaeII	ACGT	3 cut(s)	78, 319, 1252
MaeIII	GTNAC	3 cut(s)	987, 1196, 1262
MboII	GAAGA	5 cut(s)	601, 669, 905, 1298, 1343
MfeI	CAATTG	2 cut(s)	779, 983
MflI	RGATCY	3 cut(s)	748, 1018, 1371
MluCI	AATT	8 cut(s)	68, 275, 328, 779, 983, 1135, 1205, 1452
MlyI	GAGTC	4 cut(s)	53, 431, 546, 779
MmeI	TCCRAC	4 cut(s)	45, 192, 206, 645
MseI	TTAA	6 cut(s)	126, 237, 525, 1212, 1296, 1459
MslI	CAYNNNNRTG	1 cut(s)	966
MspA1I	CMGCKG	1 cut(s)	307
MspI	CCGG	3 cut(s)	9, 28, 152
MspR9I	CCNGG	2 cut(s)	649, 1141
MunI	CAATTG	2 cut(s)	779, 983
Mva1269I	GAATGC	1 cut(s)	1315
MvaI	CCWGG	2 cut(s)	649, 1141
MwoI	GCNNNNNNNGC	5 cut(s)	351, 603, 709, 979, 1289
NlaIII	CATG	8 cut(s)	211, 385, 436, 521, 743, 792, 965, 1435
NlaIV	GGNNCC	4 cut(s)	7, 694, 933, 1092
NmeAIII	GCCGAG	1 cut(s)	233
NmuCI	GTSAC	1 cut(s)	987
NspI	RCATGY	1 cut(s)	211
PaeI	GCATGC	1 cut(s)	211
PctI	GAATGC	1 cut(s)	1315
PfeI	GAWTC	5 cut(s)	12, 144, 266, 1073, 1229
PfoI	TCCNGGA	1 cut(s)	1139
PkrI	GCNGC	7 cut(s)	102, 133, 306, 507, 1159, 1360, 1363
PleI	GAGTC	4 cut(s)	53, 430, 545, 779
PpsI	GAGTC	4 cut(s)	53, 430, 545, 779
PpuMI	RGGWCCY	1 cut(s)	693
PshBI	ATTAAT	1 cut(s)	1212
Psp1406I	AACGTT	1 cut(s)	319
Psp5II	RGGWCCY	1 cut(s)	693
Psp6I	CCWGG	2 cut(s)	647, 1139
PspFI	CCCAGC	1 cut(s)	426
PspGI	CCWGG	2 cut(s)	647, 1139
PspN4I	GGNNCC	4 cut(s)	7, 694, 933, 1092
PspPI	GGNCC	1 cut(s)	693
PspPPI	RGGWCCY	1 cut(s)	693
PsuI	RGATCY	3 cut(s)	748, 1018, 1371
PvuII	CAGCTG	1 cut(s)	307
RsaI	GTAC	5 cut(s)	251, 515, 1041, 1183, 1320
RsaNI	GTAC	5 cut(s)	250, 514, 1040, 1182, 1319
RseI	CAYNNNNRTG	1 cut(s)	966
SaqAI	TTAA	6 cut(s)	126, 237, 525, 1212, 1296, 1459
SatI	GCNGC	7 cut(s)	101, 132, 305, 506, 1158, 1359, 1362
Sau96I	GGNCC	1 cut(s)	693
ScaI	AGTACT	3 cut(s)	251, 1041, 1320
SchI	GAGTC	4 cut(s)	53, 431, 546, 779
ScrFI	CCNGG	2 cut(s)	649, 1141
SfaNI	GCATC	4 cut(s)	765, 1144, 1381, 1427
SfcI	CTRYAG	2 cut(s)	199, 613
SinI	GGWCC	1 cut(s)	693
SmiMI	CAYNNNNRTG	1 cut(s)	966
SmlI	CTYRAG	1 cut(s)	1009
SmoI	CTYRAG	1 cut(s)	1009
SphI	GCATGC	1 cut(s)	211
Sse9I	AATT	8 cut(s)	68, 275, 328, 779, 983, 1135, 1205, 1452
SsiI	CCGC	2 cut(s)	505, 1123
SspMI	CTAG	5 cut(s)	291, 534, 626, 1316, 1376
StyD4I	CCNGG	2 cut(s)	647, 1139
StyI	CCWWGG	1 cut(s)	1375
TaaI	ACNGT	1 cut(s)	1240
TaiI	ACGT	3 cut(s)	81, 322, 1255
TaqI	TCGA	1 cut(s)	1101
TasI	AATT	8 cut(s)	68, 275, 328, 779, 983, 1135, 1205, 1452
TatI	WGTACW	3 cut(s)	249, 1039, 1318
TauI	GCSGC	1 cut(s)	508
TfiI	GAWTC	5 cut(s)	12, 144, 266, 1073, 1229
Tru1I	TTAA	6 cut(s)	126, 237, 525, 1212, 1296, 1459
Tru9I	TTAA	6 cut(s)	126, 237, 525, 1212, 1296, 1459
TseFI	GTSAC	1 cut(s)	987
TseI	GCWGC	6 cut(s)	100, 131, 304, 1157, 1358, 1361
Tsp45I	GTSAC	1 cut(s)	987
TspDTI	ATGAA	6 cut(s)	370, 449, 602, 1198, 1322, 1448
VpaK11BI	GGWCC	1 cut(s)	693
VspI	ATTAAT	1 cut(s)	1212
XapI	RAATTY	3 cut(s)	328, 1135, 1205
XbaI	TCTAGA	1 cut(s)	533
XceI	RCATGY	1 cut(s)	211
XmaJI	CCTAGG	1 cut(s)	1375
XmiI	GTMKAC	1 cut(s)	198
XspI	CTAG	5 cut(s)	291, 534, 626, 1316, 1376
ZrmI	AGTACT	3 cut(s)	251, 1041, 1320

Using CommOnly database (standard laboratory enzymes). Scanned on CDS sequence.