Basic Information

Type: gene

Biological Identity

Species

arabidopsis_thaliana

Chromosome

Orthogroup ID

OG0011313

Physical Location & Seq

Genomic Coordinates Forward (+)

26423874 .. 26425356

Gene Structure 1483 bp

Loading structure...

UTR

Exon/CDS

Intron

Transcript / Protein ID

AT1G70170.1

Sequence Viewer

CDS Protein Promoter (2k)

Length: 1137 bp

ATGAGGTTTTGTGTTTTCGGGTTTTTATCGCTTTTCTTGATTGTTTCTCCTGCCTCCGCCTGGTTCTTCCCTAACTCCACCGCGGTACCACCGTCTCTACGTAACACTACACGTGTTTTCTGGGACGCTTTCTCCAATTTCACCGGTTGTCACCACGGCCAAAACGTTGATGGTCTCTACCGTATCAAAAAATACTTCCAACGTTTTGGTTACATCCCTGAAACGTTTTCGGGAAATTTCACCGATGATTTTGATGATATCCTCAAAGCCGCTGTCGAATTGTATCAGACGAATTTCAATCTTAACGTTACCGGTGAGCTTGATGCGCTTACCATTCAACACATCGTGATCCCGCGTTGCGGTAATCCTGACGTGGTCAACGGTACTTCGTTGATGCATGGCGGGAGAAGAAAGACCTTCGAGGTCAACTTCTCCCGCACGCATTTGCACGCGGTTAAACGCTACACGTTATTCCCAGGCGAGCCACGGTGGCCGAGAAACCGCCGTGACTTGACCTACGCGTTCGACCCGAAAAACCCGTTGACGGAAGAGGTCAAGAGCGTGTTCTCACGCGCTTTTGGTCGTTGGTCTGACGTCACCGCGTTAAACTTCACTCTTTCCGAGTCTTTCTCGACGTCTGACATCACAATCGGATTCTACACCGGCGACCATGGTGACGGAGAGCCTTTCGACGGCGTTTTGGGAACATTAGCTCACGCTTTCTCTCCTCCCAGCGGGAAATTCCACCTAGACGCAGATGAGAACTGGGTGGTCTCCGGAGACCTAGACAGTTTCCTCTCCGTAACTGCTGCGGTTGATCTTGAATCAGTGGCGGTTCACGAGATCGGTCATCTTCTAGGTTTAGGACATTCTTCAGTTGAGGAATCCATCATGTATCCAACGATCACGACAGGGAAACGTAAGGTTGACTTAACCAATGATGACGTGGAAGGAATCCAATACTTGTACGGTGCAAACCCTAACTTCAACGGCACAACATCTCCACCGTCGACCACTAAACACCAGCGAGACACTGGTGGCTTCAGTGCCGCCTGGAGAATCGACGGTTCATCGAGATCAACAATTGTTAGTCTCTTGTTGTCAACCGTCGGATTGGTCTTGTGGTTCTTACCGTAG

Functional Annotation

Gene Ontology

Molecular Function
Biological Process
Cellular Component

GO:0001101 GO:0003674 GO:0003824 GO:0004175 GO:0004222 GO:0005575 GO:0005623 GO:0005886 GO:0006508 GO:0006807 GO:0006950 GO:0006970 GO:0008150 GO:0008152 GO:0008233 GO:0008237 GO:0009628 GO:0009651 GO:0009719 GO:0009725 GO:0009753 GO:0010033 GO:0010035 GO:0010038 GO:0016020 GO:0016787 GO:0019538 GO:0031224 GO:0031225 GO:0032502 GO:0040007 GO:0042221 GO:0043170 GO:0044238 GO:0044425 GO:0044464 GO:0046686 GO:0048519 GO:0048580 GO:0048583 GO:0048589 GO:0048831 GO:0050789 GO:0050793 GO:0050896 GO:0051093 GO:0051239 GO:0051241 GO:0065007 GO:0070011 GO:0071704 GO:0071944 GO:0080186 GO:0140096 GO:1900055 GO:1900056 GO:1901564 GO:1901700 GO:1905622 GO:2000024 GO:2000026 GO:2000028 GO:2000241

KEGG Pathways

Metabolic & Signaling

No pathways identified.

Pfam Domains

Protein Families

PG_binding_1 Peptidase_M10

Protein Analysis

378

Amino Acids

42.01

Weight (kDa)

5.78

Isoelectric Point (pI)

31.18

Instability Index

Protein Domains (Pfam)

Domain Name	Pfam ID	Position	E-value	Description
PG_binding_1	PF01471	60 - 114	1.5e-10	Putative peptidoglycan binding domain
Peptidase_M10	PF00413	164 - 324	1.7e-51	Matrixin

Hydrophobicity Profile (Kyte-Doolittle)

AI Structure Prediction Report

Calculating structure properties...

Genomic Context

Gene Family Tree

Species Filter

Style Settings

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Align Labels (Cladogram)

Image

Tree File

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Publication-ready

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Orthologous Genes (Group: OG0011313)

Species	Orthologous Gene IDs
arabidopsis_thaliana	AT1G70170
fragaria_vesca	FvH4_4g35180 FvH4_4g35180
malus_domestica	MD13G1085700.v1.1 MD16G1085000.v1.1
prunus_persica	Prupe.1G261000_v2.0.a1
pyrus_communis	pycom13g07410 pycom16g07350
rosa_chinensis	RchiOBHm_Chr4g0444791
rosa_laevigata	RLG00000005802
rosa_multiflora	Rmu_sc0000096.1_g000019 Rmu_sc0007130.1_g000003 Rmu_sc0037188.1_g000001
rosa_roxburghii	Rroxscaffold_5G00385280
rosa_rugosa	Rorug04G0355300
rosa_samantha	Rh4AG416500 Rh4BG427900 Rh4CG442700 Rh4DG423600
rosa_wichuraiana	Rw4G035690

Restriction Enzyme Sites

1 / 10

Enzyme	Recognition Site	Cut Count	Positions (bp)
AatII	GACGTC	2 cut(s)	597, 638
Acc65I	GGTACC	1 cut(s)	85
AccB1I	GGYRCC	1 cut(s)	85
AccI	GTMKAC	1 cut(s)	1010
AccII	CGCG	6 cut(s)	83, 355, 452, 521, 573, 602
AccIII	TCCGGA	1 cut(s)	776
AclI	AACGTT	4 cut(s)	165, 202, 224, 306
AclWI	GGATC	1 cut(s)	343
AcoI	YGGCCR	2 cut(s)	157, 491
AcsI	RAATTY	3 cut(s)	235, 292, 740
AcuI	CTGAAG	2 cut(s)	858, 1027
AcvI	CACGTG	1 cut(s)	113
AcyI	GRCGYC	2 cut(s)	594, 635
AdeI	CACNNNGTG	1 cut(s)	346
AfaI	GTAC	3 cut(s)	87, 385, 968
AfiI	CCNNNNNNNGG	5 cut(s)	60, 359, 544, 692, 734
AflIII	ACRYGT	4 cut(s)	110, 112, 465, 519
AgeI	ACCGGT	2 cut(s)	143, 311
AgsI	TTSAA	4 cut(s)	298, 338, 824, 988
AjiI	CACGTC	2 cut(s)	373, 946
AjnI	CCWGG	3 cut(s)	59, 475, 1052
AluBI	AGCT	2 cut(s)	319, 713
AluI	AGCT	2 cut(s)	319, 713
Alw26I	GTCTC	6 cut(s)	99, 179, 774, 778, 1023, 1097
AlwI	GGATC	1 cut(s)	343
Aor13HI	TCCGGA	1 cut(s)	776
AoxI	GGCC	2 cut(s)	157, 491
ApeKI	GCWGC	1 cut(s)	809
ApoI	RAATTY	3 cut(s)	235, 292, 740
ArsI	GACNNNNNNTTYG	2 cut(s)	258, 290
AsiGI	ACCGGT	2 cut(s)	143, 311
Asp718I	GGTACC	1 cut(s)	85
AspLEI	GCGC	2 cut(s)	328, 575
AsuHPI	GGTGA	6 cut(s)	133, 143, 232, 326, 589, 686
BaeI	ACNNNNGTAYC	4 cut(s)	69, 77, 102, 110
BanI	GGYRCC	1 cut(s)	85
BauI	CACGAG	1 cut(s)	839
BbrPI	CACGTG	1 cut(s)	113
BbvI	GCAGC	1 cut(s)	796
BccI	CCATC	2 cut(s)	164, 896
BceAI	ACGGC	4 cut(s)	172, 489, 709, 1006
BciT130I	CCWGG	3 cut(s)	61, 477, 1054
BciVI	GTATCC	1 cut(s)	906
BcoDI	GTCTC	6 cut(s)	99, 179, 774, 778, 1023, 1097
BfaI	CTAG	3 cut(s)	749, 785, 857
BfuI	GTATCC	1 cut(s)	906
BglI	GCCNNNNNGGC	1 cut(s)	490
BisI	GCNGC	3 cut(s)	270, 810, 1050
BlsI	GCNGC	3 cut(s)	271, 811, 1051
Bme1390I	CCNGG	3 cut(s)	61, 477, 1054
BmgBI	CACGTC	2 cut(s)	373, 946
BmiI	GGNNCC	1 cut(s)	87
BmrFI	CCNGG	3 cut(s)	61, 477, 1054
BmrI	ACTGGG	1 cut(s)	775
BmsI	GCATC	2 cut(s)	313, 384
BmuI	ACTGGG	1 cut(s)	775
BplI	GAGNNNNNCTC	2 cut(s)	614, 646
BpmI	CTGGAG	1 cut(s)	1075
BsaAI	YACGTR	2 cut(s)	101, 113
BsaHI	GRCGYC	2 cut(s)	594, 635
BsaI	GGTCTC	3 cut(s)	179, 774, 778
BsaJI	CCNNGG	5 cut(s)	81, 154, 475, 485, 670
BsaWI	WCCGGW	3 cut(s)	143, 311, 776
BsaXI	ACNNNNNCTCC	4 cut(s)	116, 146, 985, 1015
Bsc4I	CCNNNNNNNGG	5 cut(s)	60, 359, 544, 692, 734
Bse118I	RCCGGY	3 cut(s)	143, 311, 662
Bse1I	ACTGG	2 cut(s)	770, 1039
BseAI	TCCGGA	1 cut(s)	776
BseBI	CCWGG	3 cut(s)	61, 477, 1054
BseDI	CCNNGG	5 cut(s)	81, 154, 475, 485, 670
BseGI	GGATG	1 cut(s)	213
BseLI	CCNNNNNNNGG	5 cut(s)	60, 359, 544, 692, 734
BseNI	ACTGG	2 cut(s)	770, 1039
BseRI	GAGGAG	1 cut(s)	717
BseXI	GCAGC	1 cut(s)	796
BseYI	CCCAGC	1 cut(s)	731
Bsh1236I	CGCG	6 cut(s)	83, 355, 452, 521, 573, 602
BshFI	GGCC	2 cut(s)	159, 493
BshNI	GGYRCC	1 cut(s)	85
BshTI	ACCGGT	2 cut(s)	143, 311
BsiSI	CCGG	4 cut(s)	144, 312, 663, 777
BslFI	GGGAC	1 cut(s)	137
BslI	CCNNNNNNNGG	5 cut(s)	60, 359, 544, 692, 734
BsmAI	GTCTC	6 cut(s)	99, 179, 774, 778, 1023, 1097
BsmBI	CGTCTC	1 cut(s)	99
BsmFI	GGGAC	1 cut(s)	137
BsnI	GGCC	2 cut(s)	159, 493
Bso31I	GGTCTC	3 cut(s)	179, 774, 778
Bsp13I	TCCGGA	1 cut(s)	776
Bsp143I	GATC	5 cut(s)	348, 817, 843, 903, 1076
Bsp19I	CCATGG	1 cut(s)	670
BspANI	GGCC	2 cut(s)	159, 493
BspEI	TCCGGA	1 cut(s)	776
BspFNI	CGCG	6 cut(s)	83, 355, 452, 521, 573, 602
BspLI	GGNNCC	1 cut(s)	87
BspPI	GGATC	1 cut(s)	343
BspT107I	GGYRCC	1 cut(s)	85
BspTNI	GGTCTC	3 cut(s)	179, 774, 778
BsrFI	RCCGGY	3 cut(s)	143, 311, 662
BsrI	ACTGG	2 cut(s)	770, 1039
BssAI	RCCGGY	3 cut(s)	143, 311, 662
BssECI	CCNNGG	5 cut(s)	81, 154, 475, 485, 670
BssMI	GATC	5 cut(s)	348, 817, 843, 903, 1076
BssNI	GRCGYC	2 cut(s)	594, 635
BssSI	CACGAG	1 cut(s)	839
BssT1I	CCWWGG	1 cut(s)	670
Bst2BI	CACGAG	1 cut(s)	839
Bst2UI	CCWGG	3 cut(s)	61, 477, 1054
Bst6I	CTCTTC	1 cut(s)	543
BstACI	GRCGYC	2 cut(s)	594, 635
BstBAI	YACGTR	2 cut(s)	101, 113
BstC8I	GCNNGC	3 cut(s)	440, 450, 482
BstDSI	CCRYGG	4 cut(s)	81, 154, 485, 670
BstF5I	GGATG	1 cut(s)	213
BstFNI	CGCG	6 cut(s)	83, 355, 452, 521, 573, 602
BstHHI	GCGC	2 cut(s)	328, 575
BstKTI	GATC	5 cut(s)	351, 820, 846, 906, 1079
BstMAI	GTCTC	6 cut(s)	99, 179, 774, 778, 1023, 1097
BstMBI	GATC	5 cut(s)	348, 817, 843, 903, 1076
BstMWI	GCNNNNNNNGC	2 cut(s)	325, 490
BstNI	CCWGG	3 cut(s)	61, 477, 1054
BstSCI	CCNGG	3 cut(s)	59, 475, 1052
BstSNI	TACGTA	1 cut(s)	101
BstUI	CGCG	6 cut(s)	83, 355, 452, 521, 573, 602
BstV1I	GCAGC	1 cut(s)	796
BstXI	CCANNNNNNTGG	1 cut(s)	206
BsuI	GTATCC	1 cut(s)	906
BsuRI	GGCC	2 cut(s)	159, 493
BtgI	CCRYGG	4 cut(s)	81, 154, 485, 670
BtrI	CACGTC	2 cut(s)	373, 946
BtsCI	GGATG	1 cut(s)	213
BtsIMutI	CAGTG	3 cut(s)	834, 1032, 1051
Cac8I	GCNNGC	3 cut(s)	440, 450, 482
CfoI	GCGC	2 cut(s)	328, 575
Cfr10I	RCCGGY	3 cut(s)	143, 311, 662
Cfr42I	CCGCGG	1 cut(s)	84
CseI	GACGC	2 cut(s)	134, 761
Csp6I	GTAC	3 cut(s)	86, 384, 967
CspAI	ACCGGT	2 cut(s)	143, 311
CviAII	CATG	3 cut(s)	398, 671, 892
CviJI	RGCY	8 cut(s)	159, 269, 319, 484, 493, 685, 713, 1041
CviKI_1	RGCY	8 cut(s)	159, 269, 319, 484, 493, 685, 713, 1041
CviQI	GTAC	3 cut(s)	86, 384, 967
DpnI	GATC	5 cut(s)	350, 819, 845, 905, 1078
DpnII	GATC	5 cut(s)	348, 817, 843, 903, 1076
DraIII	CACNNNGTG	1 cut(s)	346
EaeI	YGGCCR	2 cut(s)	157, 491
Eam1104I	CTCTTC	1 cut(s)	543
EarI	CTCTTC	1 cut(s)	543
EciI	GGCGGA	1 cut(s)	46
Eco105I	TACGTA	1 cut(s)	101
Eco130I	CCWWGG	1 cut(s)	670
Eco31I	GGTCTC	3 cut(s)	179, 774, 778
Eco32I	GATATC	1 cut(s)	259
Eco57I	CTGAAG	2 cut(s)	858, 1027
Eco72I	CACGTG	1 cut(s)	113
EcoRII	CCWGG	3 cut(s)	59, 475, 1052
EcoRV	GATATC	1 cut(s)	259
EcoT14I	CCWWGG	1 cut(s)	670
EcoT22I	ATGCAT	1 cut(s)	399
ErhI	CCWWGG	1 cut(s)	670
Esp3I	CGTCTC	1 cut(s)	99
FaeI	CATG	3 cut(s)	401, 674, 895
FaiI	YATR	3 cut(s)	399, 672, 893
FalI	AAGNNNNNCTT	2 cut(s)	914, 946
FaqI	GGGAC	1 cut(s)	137
FatI	CATG	3 cut(s)	397, 670, 891
FauI	CCCGC	4 cut(s)	360, 395, 443, 728
FblI	GTMKAC	1 cut(s)	1010
Fnu4HI	GCNGC	3 cut(s)	270, 810, 1050
FokI	GGATG	1 cut(s)	200
Fsp4HI	GCNGC	3 cut(s)	270, 810, 1050
FspBI	CTAG	3 cut(s)	749, 785, 857
GlaI	GCGC	2 cut(s)	327, 574
GluI	GCNGC	3 cut(s)	270, 810, 1050
GsaI	CCCAGC	1 cut(s)	735
GsuI	CTGGAG	1 cut(s)	1075
HaeIII	GGCC	2 cut(s)	159, 493
HapII	CCGG	4 cut(s)	144, 312, 663, 777
HgaI	GACGC	2 cut(s)	134, 761
HhaI	GCGC	2 cut(s)	328, 575
Hin1I	GRCGYC	2 cut(s)	594, 635
Hin1II	CATG	3 cut(s)	401, 674, 895
Hin6I	GCGC	2 cut(s)	326, 573
HinP1I	GCGC	2 cut(s)	326, 573
HincII	GTYRAC	6 cut(s)	379, 427, 543, 928, 1011, 1104
HindII	GTYRAC	6 cut(s)	379, 427, 543, 928, 1011, 1104
HinfI	GANTC	6 cut(s)	623, 654, 824, 884, 954, 1059
HpaII	CCGG	4 cut(s)	144, 312, 663, 777
HphI	GGTGA	6 cut(s)	133, 143, 232, 326, 589, 686
Hpy166II	GTNNAC	7 cut(s)	379, 427, 543, 838, 928, 1011, 1104
Hpy188I	TCNGA	6 cut(s)	288, 592, 622, 640, 653, 1112
Hpy8I	GTNNAC	7 cut(s)	379, 427, 543, 838, 928, 1011, 1104
Hpy99I	CGWCG	5 cut(s)	637, 695, 1012, 1067, 1112
HpyAV	CCTTC	2 cut(s)	427, 944
HpyCH4V	TGCA	3 cut(s)	397, 448, 974
HpyF10VI	GCNNNNNNNGC	2 cut(s)	325, 490
Hsp92I	GRCGYC	2 cut(s)	594, 635
Hsp92II	CATG	3 cut(s)	401, 674, 895
HspAI	GCGC	2 cut(s)	326, 573
Kpn2I	TCCGGA	1 cut(s)	776
KpnI	GGTACC	1 cut(s)	89
KspI	CCGCGG	1 cut(s)	84
Kzo9I	GATC	5 cut(s)	348, 817, 843, 903, 1076
Lsp1109I	GCAGC	1 cut(s)	796
LweI	GCATC	2 cut(s)	313, 384
MaeI	CTAG	3 cut(s)	749, 785, 857
MaeIII	GTNAC	8 cut(s)	101, 149, 209, 307, 506, 595, 674, 802
MalI	GATC	5 cut(s)	350, 819, 845, 905, 1078
MboI	GATC	5 cut(s)	348, 817, 843, 903, 1076
MboII	GAAGA	5 cut(s)	58, 420, 560, 845, 864
MfeI	CAATTG	1 cut(s)	1083
MluCI	AATT	6 cut(s)	136, 235, 278, 292, 740, 1083
MluI	ACGCGT	1 cut(s)	519
MlyI	GAGTC	1 cut(s)	632
MmeI	TCCRAC	3 cut(s)	223, 923, 1090
MnlI	CCTC	7 cut(s)	64, 272, 415, 544, 738, 806, 874
Mph1103I	ATGCAT	1 cut(s)	399
MroI	TCCGGA	1 cut(s)	776
MseI	TTAA	4 cut(s)	303, 456, 605, 932
MspA1I	CMGCKG	3 cut(s)	83, 272, 735
MspI	CCGG	4 cut(s)	144, 312, 663, 777
MspR9I	CCNGG	3 cut(s)	61, 477, 1054
MunI	CAATTG	1 cut(s)	1083
MvaI	CCWGG	3 cut(s)	61, 477, 1054
MvnI	CGCG	6 cut(s)	83, 355, 452, 521, 573, 602
MwoI	GCNNNNNNNGC	2 cut(s)	325, 490
NcoI	CCATGG	1 cut(s)	670
NdeII	GATC	5 cut(s)	348, 817, 843, 903, 1076
NlaIII	CATG	3 cut(s)	401, 674, 895
NlaIV	GGNNCC	1 cut(s)	87
NmeAIII	GCCGAG	1 cut(s)	519
NmuCI	GTSAC	4 cut(s)	149, 506, 595, 674
NsiI	ATGCAT	1 cut(s)	399
PcsI	WCGNNNNNNNCGW	1 cut(s)	162
PfeI	GAWTC	5 cut(s)	654, 824, 884, 954, 1059
PflFI	GACNNNGTC	1 cut(s)	374
PinAI	ACCGGT	2 cut(s)	143, 311
PkrI	GCNGC	3 cut(s)	271, 811, 1051
PleI	GAGTC	1 cut(s)	631
PmaCI	CACGTG	1 cut(s)	113
PmlI	CACGTG	1 cut(s)	113
PpsI	GAGTC	1 cut(s)	631
Ppu21I	YACGTR	2 cut(s)	101, 113
Psp1406I	AACGTT	4 cut(s)	165, 202, 224, 306
Psp6I	CCWGG	3 cut(s)	59, 475, 1052
PspCI	CACGTG	1 cut(s)	113
PspFI	CCCAGC	1 cut(s)	731
PspGI	CCWGG	3 cut(s)	59, 475, 1052
PspN4I	GGNNCC	1 cut(s)	87
PsyI	GACNNNGTC	1 cut(s)	374
RsaI	GTAC	3 cut(s)	87, 385, 968
RsaNI	GTAC	3 cut(s)	86, 384, 967
SacII	CCGCGG	1 cut(s)	84
SalI	GTCGAC	1 cut(s)	1009
SaqAI	TTAA	4 cut(s)	303, 456, 605, 932
SatI	GCNGC	3 cut(s)	270, 810, 1050
Sau3AI	GATC	5 cut(s)	348, 817, 843, 903, 1076
SchI	GAGTC	1 cut(s)	632
ScrFI	CCNGG	3 cut(s)	61, 477, 1054
SfaNI	GCATC	2 cut(s)	313, 384
Sfr303I	CCGCGG	1 cut(s)	84
SgrAI	CRCCGGYG	1 cut(s)	662
SgrBI	CCGCGG	1 cut(s)	84
SnaBI	TACGTA	1 cut(s)	101
Sse9I	AATT	6 cut(s)	136, 235, 278, 292, 740, 1083
SspMI	CTAG	3 cut(s)	749, 785, 857
StyD4I	CCNGG	3 cut(s)	59, 475, 1052
StyI	CCWWGG	1 cut(s)	670
TaqI	TCGA	8 cut(s)	276, 420, 525, 632, 690, 1010, 1062, 1073
TaqII	GACCGA	1 cut(s)	836
TasI	AATT	6 cut(s)	136, 235, 278, 292, 740, 1083
TauI	GCSGC	2 cut(s)	272, 1052
TfiI	GAWTC	5 cut(s)	654, 824, 884, 954, 1059
Tru1I	TTAA	4 cut(s)	303, 456, 605, 932
Tru9I	TTAA	4 cut(s)	303, 456, 605, 932
TscAI	CASTG	3 cut(s)	834, 1039, 1051
TseFI	GTSAC	4 cut(s)	149, 506, 595, 674
TseI	GCWGC	1 cut(s)	809
Tsp45I	GTSAC	4 cut(s)	149, 506, 595, 674
TspDTI	ATGAA	1 cut(s)	1059
TspGWI	ACGGA	3 cut(s)	560, 693, 790
TspRI	CASTG	3 cut(s)	834, 1039, 1051
Tth111I	GACNNNGTC	1 cut(s)	374
XapI	RAATTY	3 cut(s)	235, 292, 740
XcmI	CCANNNNNNNNNTGG	3 cut(s)	167, 943, 1031
XmiI	GTMKAC	1 cut(s)	1010
XspI	CTAG	3 cut(s)	749, 785, 857
ZraI	GACGTC	2 cut(s)	595, 636
Zsp2I	ATGCAT	1 cut(s)	399

Using CommOnly database (standard laboratory enzymes). Scanned on CDS sequence.