FvH4_5g09900

Pentatricopeptide repeat-containing protein

Design Primers CRISPR Design

Basic Information

Type: gene
Biological Identity
fragaria_vesca
Fvb5
OG0015029
Physical Location & Seq
Forward (+)
5658227 .. 5661208
2982 bp
Loading structure...
UTR
Exon/CDS
Intron
FvH4_5g09900.t1

Sequence Viewer

Length: 1584 bp
ATGAGTAGCACCACCAATCTGATCACCCTAATCAAACACTGCAAAACCCCAAAACAACTCCACCAAATCCATGCTCACACCACCACTGCCACTGCCACTGCCACTGCCTCTCATCCCAAAATCCTACCCAACCTCCTCCACACCTTCACCTCTCTCCTCAAACCCTCCTCCTCTCCAACCTCACTCACCTACGCCACATCTCTCTTCAACCTCATCCCAAACCCCTCCACCTTCTCCTTCAACAACCTCATCCGGGCCCACACCCTCCTCTCCTCCCCTCTCCCCGCCCTCCACCTCTTCGCTTCCCTTCGCCGCCTTTCCCTCCCCCCTGACTTCCACACCTTCCCCTTCGCCCTCAAAGCCTGCGCCCACCTCGCCCCTTCTTCGTTCCCGATCTCCCAGGCCCTACATTCCCAAGCCCTCAAGTTCGGTTTCGCTTCACATTCCTTCGTCGCCAACACCCTCATTCATGTCTATTCAGTATGCCACAATCTTAACCACGCACGCCAGCTGTTCGACGAAAGTCCCCACAAGGACATGGTCTCGTACAATGCGATGCTCGACGCGTTTGTTAAGGGAGGCGAGATGGACAATGCGAGAGAGGTGTTTGATGAAATGCCTGAGAGGGACTCTGTTTCGTGGGGGACTGTTTTGGCAGGGTATGCGCAGACGAGCCGCTGCGAGGAGGCGATTGAGCTGTTTGGTGAGATGATGGAGTCGGAAATTAGGCCTGATGATATCTGCTTGGTTTCGGCACTCTCTGCTTGTGCGCAGCTGGGTGATTTGGAGCGAGGGAGGAGCATCCATGACTATGTTAAACGAAACAGGAGTAGACTAAGTTCGTATTTTTTGACAGCGTTGGTGGACTTGTATGCGAAATGTGGGTGCATTGAGACTGCTATGAAGATTTTCGAGTCGAGCTCTGGGAAGAATGTGTTCACTTGGAATGCCATGCTTGTTGGGCTTGCAATGCATGGTCATGCCCATCTCTCACTGGAGTACTTCTTCAGGATGATAGAGGCTGGAGTTAAACCTGACGGGGTTAGCTTCTTGGGGGTTTTAGTGGGTTGCAGCCATGGAGGTAAAGTAGACGAAGCTAGAAAGCTTTTCGATGAAATGGAATCGGTCTATGGGGTGCCCAGAGAGCTCAAGCATTATGGGTGCATGGCTGATCTGCTTGGAAGAGCAGGGTTGATCAAGGAGGCTATGGAGATGATAGAGAAGATGCCAATGGGGGGTGATGTGTTTGTGTGGGGTGGTTTGCTTGGAGGTTGCAAGATACATGGTGATGTGGAGAGTGCAAAGACAGCCGCTGAGCATGTGATGGAGTTGGACCCTGAAGATGGTGGTGTGTATTCAATCATGGCTAGTGTTTATGCCAATGCAGAGAAATGGGATGATCTGGTTAAAACCAGAAATTCCATAAGTGCTAATAAGAGGGTCAAGAAGAACAATGGCTGTAGTTTGATTAGGTTGAATGGAACCAACCATGAGTTTCTTGCAGGGGATAGCTTACATCCTCAAACAGAAGCAATATATTTTGTTCTGGACGGACTTTGGAAACATCAATTTGAATCATGCTAA
Functional Annotation
Gene Ontology
Molecular Function
Biological Process
Cellular Component
No ontology terms assigned.
KEGG Pathways
Metabolic & Signaling
No pathways identified.
Pfam Domains
Protein Families
PPR

Protein Analysis

528

Amino Acids

58.09

Weight (kDa)

6.37

Isoelectric Point (pI)

43.43

Instability Index
Protein Domains (Pfam)
Domain Name Pfam ID Position E-value Description
PPR_2 PF13041 179 - 209 2.5e-06 PPR repeat family
PPR PF01535 181 - 209 1.6e-07 PPR repeat
PPR_2 PF13041 210 - 257 7e-07 PPR repeat family
PPR PF01535 212 - 241 6.9e-06 PPR repeat
E_motif PF20431 428 - 490 2.6e-10 E motif
Hydrophobicity Profile (Kyte-Doolittle)
AI Structure Prediction Report
Calculating structure properties...

Genomic Context

Gene Family Tree

Species Filter
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Publication-ready
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Orthologous Genes (Group: OG0015029)

Species Orthologous Gene IDs
arabidopsis_thaliana AT5G61800
fragaria_vesca FvH4_5g09900
malus_domestica MD00G1076000.v1.1
prunus_persica Prupe.5G139600_v2.0.a1
pyrus_communis pycom06g11970
rosa_chinensis RchiOBHm_Chr7g0188381
rosa_laevigata RLG00000004697
rosa_multiflora Rmu_co8347981.1_g000001 Rmu_sc0000952.1_g000005
rosa_roxburghii Rroxscaffold_3G00266440
rosa_rugosa Rorug06G0490300
rosa_samantha Rh7AG095000 Rh7BG096500 Rh7CG097000 Rh7CG097100
rosa_wichuraiana Rw7G008190

Restriction Enzyme Sites

1 / 10
Enzyme Recognition Site Cut Count Positions (bp)
Acc16I TGCGCA 2 cut(s) 666, 771
AccB1I GGYRCC 1 cut(s) 1135
AccI GTMKAC 2 cut(s) 832, 1089
AccII CGCG 1 cut(s) 566
AciI CCGC 4 cut(s) 285, 313, 676, 1311
AcsI RAATTY 1 cut(s) 1417
AcuI CTGAAG 2 cut(s) 991, 1359
AfaI GTAC 2 cut(s) 548, 1001
AfiI CCNNNNNNNGG 4 cut(s) 253, 682, 1235, 1343
AflIII ACRYGT 1 cut(s) 564
AgsI TTSAA 5 cut(s) 208, 241, 1359, 1477, 1574
AjnI CCWGG 1 cut(s) 399
AluBI AGCT 9 cut(s) 511, 697, 775, 921, 1047, 1097, 1105, 1147, 1512
AluI AGCT 9 cut(s) 511, 697, 775, 921, 1047, 1097, 1105, 1147, 1512
Alw21I GWGCWC 2 cut(s) 923, 1149
Alw26I GTCTC 2 cut(s) 547, 887
AlwNI CAGNNNCTG 1 cut(s) 1313
AoxI GGCC 3 cut(s) 255, 402, 728
ApaI GGGCCC 1 cut(s) 259
ApeKI GCWGC 3 cut(s) 678, 772, 1071
ApoI RAATTY 1 cut(s) 1417
Asp700I GAANNNNTTC 2 cut(s) 908, 935
AspLEI GCGC 3 cut(s) 368, 667, 772
AspS9I GGNCC 4 cut(s) 255, 256, 403, 1333
AsuC2I CCSGG 1 cut(s) 254
AsuHPI GGTGA 7 cut(s) 16, 139, 178, 716, 791, 1250, 1298
AvaII GGWCC 1 cut(s) 1333
BaeGI GKGCMC 2 cut(s) 259, 1140
BanI GGYRCC 1 cut(s) 1135
BanII GRGCYC 3 cut(s) 259, 923, 1149
Bbv12I GWGCWC 2 cut(s) 923, 1149
BbvI GCAGC 3 cut(s) 665, 784, 1083
BccI CCATC 5 cut(s) 580, 706, 993, 1318, 1337
BciT130I CCWGG 1 cut(s) 401
BclI TGATCA 2 cut(s) 21, 1194
BcnI CCSGG 1 cut(s) 254
BcoDI GTCTC 2 cut(s) 547, 887
BfaI CTAG 2 cut(s) 1098, 1368
BfmI CTRYAG 1 cut(s) 1459
BisI GCNGC 6 cut(s) 313, 676, 679, 773, 1072, 1311
BlpI GCTNAGC 1 cut(s) 1314
BlsI GCNGC 6 cut(s) 314, 677, 680, 774, 1073, 1312
BmcAI AGTACT 1 cut(s) 1001
Bme1390I CCNGG 2 cut(s) 254, 401
Bme18I GGWCC 1 cut(s) 1333
BmgT120I GGNCC 4 cut(s) 255, 256, 403, 1333
BmiI GGNNCC 4 cut(s) 257, 1137, 1335, 1483
BmrFI CCNGG 2 cut(s) 254, 401
BmsI GCATC 3 cut(s) 546, 810, 1215
BoxI GACNNNNGTC 1 cut(s) 522
BplI GAGNNNNNCTC 4 cut(s) 614, 646, 905, 937
BpmI CTGGAG 2 cut(s) 1016, 1044
Bpu1102I GCTNAGC 1 cut(s) 1314
BpuEI CTTGAG 2 cut(s) 407, 1133
BpuMI CCSGG 1 cut(s) 254
BsaI GGTCTC 1 cut(s) 547
BsaJI CCNNGG 2 cut(s) 399, 1075
BsaXI ACNNNNNCTCC 4 cut(s) 117, 147, 1071, 1101
Bsc4I CCNNNNNNNGG 4 cut(s) 253, 682, 1235, 1343
Bse1I ACTGG 1 cut(s) 999
Bse3DI GCAATG 1 cut(s) 975
BseBI CCWGG 1 cut(s) 401
BseDI CCNNGG 2 cut(s) 399, 1075
BseGI GGATG 7 cut(s) 112, 213, 249, 801, 1017, 1402, 1516
BseLI CCNNNNNNNGG 4 cut(s) 253, 682, 1235, 1343
BseMI GCAATG 1 cut(s) 975
BseMII CTCAG 2 cut(s) 612, 1305
BseNI ACTGG 1 cut(s) 999
BseRI GAGGAG 8 cut(s) 125, 146, 157, 160, 257, 262, 698, 811
BseSI GKGCMC 2 cut(s) 259, 1140
BseXI GCAGC 3 cut(s) 665, 784, 1083
BseYI CCCAGC 1 cut(s) 775
Bsh1236I CGCG 1 cut(s) 566
BshFI GGCC 3 cut(s) 257, 404, 730
BshNI GGYRCC 1 cut(s) 1135
BsiHKAI GWGCWC 2 cut(s) 923, 1149
BsiSI CCGG 1 cut(s) 253
BslFI GGGAC 3 cut(s) 510, 641, 658
BslI CCNNNNNNNGG 4 cut(s) 253, 682, 1235, 1343
BsmAI GTCTC 2 cut(s) 547, 887
BsmFI GGGAC 3 cut(s) 510, 641, 658
BsmI GAATGC 1 cut(s) 952
BsnI GGCC 3 cut(s) 257, 404, 730
Bso31I GGTCTC 1 cut(s) 547
Bsp120I GGGCCC 1 cut(s) 255
Bsp1286I GDGCHC 4 cut(s) 259, 923, 1140, 1149
Bsp143I GATC 5 cut(s) 21, 393, 1171, 1194, 1399
Bsp1720I GCTNAGC 1 cut(s) 1314
Bsp19I CCATGG 1 cut(s) 1075
BspACI CCGC 4 cut(s) 285, 313, 676, 1311
BspANI GGCC 3 cut(s) 257, 404, 730
BspCNI CTCAG 2 cut(s) 613, 1306
BspFNI CGCG 1 cut(s) 566
BspLI GGNNCC 4 cut(s) 257, 1137, 1335, 1483
BspQI GCTCTTC 1 cut(s) 1177
BspT107I GGYRCC 1 cut(s) 1135
BspTNI GGTCTC 1 cut(s) 547
BsrDI GCAATG 1 cut(s) 975
BsrI ACTGG 1 cut(s) 999
BssECI CCNNGG 2 cut(s) 399, 1075
BssMI GATC 5 cut(s) 21, 393, 1171, 1194, 1399
BssT1I CCWWGG 1 cut(s) 1075
Bst2UI CCWGG 1 cut(s) 401
Bst4CI ACNGT 1 cut(s) 649
Bst6I CTCTTC 3 cut(s) 209, 302, 1177
BstAPI GCANNNNNTGC 2 cut(s) 662, 761
BstC8I GCNNGC 4 cut(s) 364, 505, 509, 966
BstDEI CTNAG 3 cut(s) 621, 836, 1314
BstDSI CCRYGG 1 cut(s) 1075
BstF5I GGATG 7 cut(s) 112, 213, 249, 801, 1017, 1402, 1516
BstFNI CGCG 1 cut(s) 566
BstHHI GCGC 3 cut(s) 368, 667, 772
BstKTI GATC 5 cut(s) 24, 396, 1174, 1197, 1402
BstMAI GTCTC 2 cut(s) 547, 887
BstMBI GATC 5 cut(s) 21, 393, 1171, 1194, 1399
BstMWI GCNNNNNNNGC 8 cut(s) 359, 374, 662, 761, 961, 970, 1144, 1307
BstNI CCWGG 1 cut(s) 401
BstNSI RCATGY 1 cut(s) 1322
BstPAI GACNNNNGTC 1 cut(s) 522
BstSCI CCNGG 2 cut(s) 252, 399
BstSFI CTRYAG 1 cut(s) 1459
BstSLI GKGCMC 2 cut(s) 259, 1140
BstUI CGCG 1 cut(s) 566
BstV1I GCAGC 3 cut(s) 665, 784, 1083
BsuRI GGCC 3 cut(s) 257, 404, 730
BtgI CCRYGG 1 cut(s) 1075
BtgZI GCGATG 1 cut(s) 569
BtsCI GGATG 7 cut(s) 112, 213, 249, 801, 1017, 1402, 1516
BtsI GCAGTG 5 cut(s) 37, 84, 90, 96, 102
BtsIMutI CAGTG 6 cut(s) 37, 84, 90, 96, 102, 992
Cac8I GCNNGC 4 cut(s) 364, 505, 509, 966
CaiI CAGNNNCTG 1 cut(s) 1313
CfoI GCGC 3 cut(s) 368, 667, 772
Cfr13I GGNCC 4 cut(s) 255, 256, 403, 1333
CseI GACGC 1 cut(s) 572
Csp6I GTAC 2 cut(s) 547, 1000
CviQI GTAC 2 cut(s) 547, 1000
DdeI CTNAG 3 cut(s) 621, 836, 1314
DpnI GATC 5 cut(s) 23, 395, 1173, 1196, 1401
DpnII GATC 5 cut(s) 21, 393, 1171, 1194, 1399
Eam1104I CTCTTC 3 cut(s) 209, 302, 1177
EarI CTCTTC 3 cut(s) 209, 302, 1177
Ecl136II GAGCTC 2 cut(s) 921, 1147
Eco130I CCWWGG 1 cut(s) 1075
Eco147I AGGCCT 1 cut(s) 730
Eco24I GRGCYC 3 cut(s) 259, 923, 1149
Eco31I GGTCTC 1 cut(s) 547
Eco32I GATATC 1 cut(s) 739
Eco47I GGWCC 1 cut(s) 1333
Eco53kI GAGCTC 2 cut(s) 921, 1147
Eco57I CTGAAG 2 cut(s) 991, 1359
EcoICRI GAGCTC 2 cut(s) 921, 1147
EcoO109I RGGNCCY 1 cut(s) 403
EcoRII CCWGG 1 cut(s) 399
EcoRV GATATC 1 cut(s) 739
EcoT14I CCWWGG 1 cut(s) 1075
EcoT22I ATGCAT 1 cut(s) 975
EcoT38I GRGCYC 3 cut(s) 259, 923, 1149
ErhI CCWWGG 1 cut(s) 1075
FaqI GGGAC 3 cut(s) 510, 641, 658
FauI CCCGC 1 cut(s) 292
FbaI TGATCA 2 cut(s) 21, 1194
FblI GTMKAC 2 cut(s) 832, 1089
Fnu4HI GCNGC 6 cut(s) 313, 676, 679, 773, 1072, 1311
FokI GGATG 7 cut(s) 99, 200, 236, 788, 1024, 1409, 1503
FriOI GRGCYC 3 cut(s) 259, 923, 1149
Fsp4HI GCNGC 6 cut(s) 313, 676, 679, 773, 1072, 1311
FspBI CTAG 2 cut(s) 1098, 1368
FspI TGCGCA 2 cut(s) 666, 771
GlaI GCGC 3 cut(s) 367, 666, 771
GluI GCNGC 6 cut(s) 313, 676, 679, 773, 1072, 1311
GsaI CCCAGC 1 cut(s) 779
GsuI CTGGAG 2 cut(s) 1016, 1044
HaeIII GGCC 3 cut(s) 257, 404, 730
HapII CCGG 1 cut(s) 253
HgaI GACGC 1 cut(s) 572
HhaI GCGC 3 cut(s) 368, 667, 772
Hin6I GCGC 3 cut(s) 366, 665, 770
HinP1I GCGC 3 cut(s) 366, 665, 770
HindIII AAGCTT 1 cut(s) 1103
HinfI GANTC 5 cut(s) 629, 716, 914, 1121, 1574
HpaII CCGG 1 cut(s) 253
HphI GGTGA 7 cut(s) 16, 139, 178, 716, 791, 1250, 1298
Hpy166II GTNNAC 4 cut(s) 833, 865, 939, 1090
Hpy188I TCNGA 2 cut(s) 21, 721
Hpy188III TCNNGA 4 cut(s) 391, 1009, 1444, 1547
Hpy8I GTNNAC 4 cut(s) 833, 865, 939, 1090
Hpy99I CGWCG 3 cut(s) 455, 521, 566
HpyAV CCTTC 8 cut(s) 154, 241, 247, 317, 352, 358, 390, 457
HpyCH4III ACNGT 1 cut(s) 649
HpyF10VI GCNNNNNNNGC 8 cut(s) 359, 374, 662, 761, 961, 970, 1144, 1307
HpyF3I CTNAG 3 cut(s) 621, 836, 1314
HspAI GCGC 3 cut(s) 366, 665, 770
Ksp22I TGATCA 2 cut(s) 21, 1194
Kzo9I GATC 5 cut(s) 21, 393, 1171, 1194, 1399
LguI GCTCTTC 1 cut(s) 1177
LmnI GCTCC 2 cut(s) 787, 798
Lsp1109I GCAGC 3 cut(s) 665, 784, 1083
LweI GCATC 3 cut(s) 546, 810, 1215
MaeI CTAG 2 cut(s) 1098, 1368
MalI GATC 5 cut(s) 23, 395, 1173, 1196, 1401
MboI GATC 5 cut(s) 21, 393, 1171, 1194, 1399
MhlI GDGCHC 4 cut(s) 259, 923, 1140, 1149
MluCI AATT 3 cut(s) 723, 1417, 1568
MluI ACGCGT 1 cut(s) 564
MlyI GAGTC 3 cut(s) 623, 725, 923
MmeI TCCRAC 3 cut(s) 200, 699, 1311
Mph1103I ATGCAT 1 cut(s) 975
MroXI GAANNNNTTC 2 cut(s) 908, 935
MseI TTAA 5 cut(s) 495, 573, 816, 1029, 1407
MslI CAYNNNNRTG 3 cut(s) 810, 978, 1287
MspA1I CMGCKG 4 cut(s) 511, 678, 775, 1313
MspI CCGG 1 cut(s) 253
MspR9I CCNGG 2 cut(s) 254, 401
Mva1269I GAATGC 1 cut(s) 952
MvaI CCWGG 1 cut(s) 401
MvnI CGCG 1 cut(s) 566
MwoI GCNNNNNNNGC 8 cut(s) 359, 374, 662, 761, 961, 970, 1144, 1307
NciI CCSGG 1 cut(s) 254
NcoI CCATGG 1 cut(s) 1075
NdeII GATC 5 cut(s) 21, 393, 1171, 1194, 1399
NlaIV GGNNCC 4 cut(s) 257, 1137, 1335, 1483
NsbI TGCGCA 2 cut(s) 666, 771
NsiI ATGCAT 1 cut(s) 975
NspI RCATGY 1 cut(s) 1322
PceI AGGCCT 1 cut(s) 730
PciSI GCTCTTC 1 cut(s) 1177
PcsI WCGNNNNNNNCGW 1 cut(s) 551
PctI GAATGC 1 cut(s) 952
PdmI GAANNNNTTC 2 cut(s) 908, 935
PfeI GAWTC 2 cut(s) 1121, 1574
PflFI GACNNNGTC 1 cut(s) 539
PkrI GCNGC 6 cut(s) 314, 677, 680, 774, 1073, 1312
PleI GAGTC 3 cut(s) 623, 724, 922
PpsI GAGTC 3 cut(s) 623, 724, 922
PshAI GACNNNNGTC 1 cut(s) 522
Psp124BI GAGCTC 2 cut(s) 923, 1149
Psp6I CCWGG 1 cut(s) 399
PspFI CCCAGC 1 cut(s) 775
PspGI CCWGG 1 cut(s) 399
PspN4I GGNNCC 4 cut(s) 257, 1137, 1335, 1483
PspOMI GGGCCC 1 cut(s) 255
PspPI GGNCC 4 cut(s) 255, 256, 403, 1333
PsrI GAACNNNNNNTAC 2 cut(s) 823, 855
PstNI CAGNNNCTG 1 cut(s) 1313
PsyI GACNNNGTC 1 cut(s) 539
PvuII CAGCTG 2 cut(s) 511, 775
RsaI GTAC 2 cut(s) 548, 1001
RsaNI GTAC 2 cut(s) 547, 1000
RseI CAYNNNNRTG 3 cut(s) 810, 978, 1287
SacI GAGCTC 2 cut(s) 923, 1149
SapI GCTCTTC 1 cut(s) 1177
SaqAI TTAA 5 cut(s) 495, 573, 816, 1029, 1407
SatI GCNGC 6 cut(s) 313, 676, 679, 773, 1072, 1311
Sau3AI GATC 5 cut(s) 21, 393, 1171, 1194, 1399
Sau96I GGNCC 4 cut(s) 255, 256, 403, 1333
ScaI AGTACT 1 cut(s) 1001
SchI GAGTC 3 cut(s) 623, 725, 923
ScrFI CCNGG 2 cut(s) 254, 401
SduI GDGCHC 4 cut(s) 259, 923, 1140, 1149
SfaNI GCATC 3 cut(s) 546, 810, 1215
SfcI CTRYAG 1 cut(s) 1459
SinI GGWCC 1 cut(s) 1333
SmiMI CAYNNNNRTG 3 cut(s) 810, 978, 1287
SmlI CTYRAG 2 cut(s) 422, 1148
SmoI CTYRAG 2 cut(s) 422, 1148
Sse9I AATT 3 cut(s) 723, 1417, 1568
SseBI AGGCCT 1 cut(s) 730
SsiI CCGC 4 cut(s) 285, 313, 676, 1311
SspMI CTAG 2 cut(s) 1098, 1368
SstI GAGCTC 2 cut(s) 923, 1149
StuI AGGCCT 1 cut(s) 730
StyD4I CCNGG 2 cut(s) 252, 399
StyI CCWWGG 1 cut(s) 1075
TaaI ACNGT 1 cut(s) 649
TaqI TCGA 5 cut(s) 516, 561, 912, 917, 1110
TaqII GACCGA 1 cut(s) 1114
TasI AATT 3 cut(s) 723, 1417, 1568
TatI WGTACW 1 cut(s) 999
TauI GCSGC 3 cut(s) 315, 678, 1313
TfiI GAWTC 2 cut(s) 1121, 1574
Tru1I TTAA 5 cut(s) 495, 573, 816, 1029, 1407
Tru9I TTAA 5 cut(s) 495, 573, 816, 1029, 1407
TscAI CASTG 6 cut(s) 44, 91, 97, 103, 109, 999
TseI GCWGC 3 cut(s) 678, 772, 1071
TspDTI ATGAA 4 cut(s) 458, 627, 917, 1128
TspGWI ACGGA 1 cut(s) 1566
TspRI CASTG 6 cut(s) 44, 91, 97, 103, 109, 999
Tth111I GACNNNGTC 1 cut(s) 539
VpaK11BI GGWCC 1 cut(s) 1333
XapI RAATTY 1 cut(s) 1417
XceI RCATGY 1 cut(s) 1322
XmiI GTMKAC 2 cut(s) 832, 1089
XmnI GAANNNNTTC 2 cut(s) 908, 935
XspI CTAG 2 cut(s) 1098, 1368
ZrmI AGTACT 1 cut(s) 1001
Zsp2I ATGCAT 1 cut(s) 975
Using CommOnly database (standard laboratory enzymes). Scanned on CDS sequence.