Rroxscaffold_1G00028270

dihydrokaempferol 4-reductase activity

Design Primers CRISPR Design

Basic Information

Type: gene
Biological Identity
rosa_roxburghii
GWHEROQ00000001
OG0010109
Physical Location & Seq
Forward (+)
36007715 .. 36010711
2997 bp
Loading structure...
UTR
Exon/CDS
Intron
Rroxscaffold_1G00028270.1

Sequence Viewer

Length: 1533 bp
ATGTCTTTCACAACGACAGTAACCGCAACAAGGAGGCCAAAACGGCAGTACCCACCAGCCCCACCGGTCCCAAAGATCCTCCATTTGCCACGTAGGCCTCGCCGGAGAACACCCAAGAGCAGCGTCGCGGCCGTTCATGGAAAACCAGCCGCGCCCACTTCGGTGGAGGACAAGAAGGGCAAGCTGGTGGCTCTGTTTGACCAAGAACGAGTTTTTCAAAAGCTGAGTGAGGGGAGGAGAGAGAGGGTTAGTGAGGGGGAGGAGGATGATGGTGGTGCTGACGTGGAGGTGGTGGAAGAGGAGAAGTGGAGGTTTCAGGCGGAGATGTTGAGGGCAGAGTGCAATTTGTTGAGGATGGAGAAGGAGATTGTGGTTAAGAAGATGGAGAGGATCAAAGTTAAAATGGAAAGGACTCTTAAATCAGCTGTGCACACTCTTGTTTCTGGGAGGAAGAAGATTAGTGAAGGAAGGAATGTAACTATGGTATTGGAGGAAGAGATTCAGCAATTGGTTCAGAAACTAGAGAAGTTGCAAAGGAATCTAGGAGCTAAGGGCGTGGAGGTTCGGAAGTCTAGTAACTTCGACAAACAAGCTTATCTTCTTCAAAGACGGTTAGAGAAGTTTAGAGGAACATCGGATGAGATATGCGTTAAGGAGATCCAGGAGATGGCAGAAGCAAGCTTATCAATCAAAACAAGCTATAGAGGAACTGAGAACTTGGTTTCAAGTGGCAAAAGAAATATGATGCAGGTGGAGATTCTGAGAAGGAGAATGGAGGGGTTGTCAAATGGGATGCTATTAAAGAGGATGAAGGAGGAATATGGTTCAATGCTGTCTACAGCCAACAACTCTGTTGCCAGTTCTGCCTCCTCGTCCAAGAGAATCGAATCCACCGATTTGTCCTCTTCCTTGGAGAAAGAAACTCATGATGAGAACGTATGCTCAGGACGTTGCAAGGCCATTGTTCGAAGGATTGTAGAGCAAGTACGAGCTGAGACAGAGCAATGGTCTCAGATGCAGGAAATGCTTGGGCGGGTGAGGGAGGAGATGGAGGAATTGCAGGCCTCTCGAGACTTTTGGGAAGATCGAGCCCTAGACTCTGATTATCATATCCAATCCTTGAGCTCCTCTGTGCAAGAATGGAGGCAGAAAGCTGTTTCATCTGAAAGCAAAGCAAGGTGCTTGCAAGAACAAGTGTCATCACTCCATGCGGAGCTAAATAGCTTGAGGAAATGTGAAAATACAAAAGTAAAGAAGGAGAATTGCTCACCTAATCCCTGGGATCCGCAGACCGAAATGGAAAAACGGGTTTTGATTTGTCACTTGAAGGAAAACCATTGTACCAAAGAATACAGCAGCAAGCAAAGGGAGGTTTTACTTGAAAGTAGGAAAAAACCAGACTCATGCACCAACAGAACACCTATTGCTCTGAAAAGATCACCATTTCGAGATATTGGAAATTCCTCATTGTCGTTGAGGCAACACAGCAAAGCAGTCTTCCCTTTACATTGTCCTCTACCTTCCAAGACTTAA
Functional Annotation
Gene Ontology
Molecular Function
Biological Process
Cellular Component
No ontology terms assigned.
KEGG Pathways
Metabolic & Signaling
No pathways identified.
Pfam Domains
Protein Families
No domains found.

Protein Analysis

510

Amino Acids

58.53

Weight (kDa)

9.46

Isoelectric Point (pI)

62.85

Instability Index
Protein Domains (Pfam)
No Pfam domains detected for this protein.
Hydrophobicity Profile (Kyte-Doolittle)
AI Structure Prediction Report
Calculating structure properties...

Genomic Context

Gene Family Tree

Species Filter
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Publication-ready
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Restriction Enzyme Sites

1 / 10
Enzyme Recognition Site Cut Count Positions (bp)
AarI CACCTGC 1 cut(s) 739
Acc36I ACCTGC 1 cut(s) 739
AccB7I CCANNNNNTGG 1 cut(s) 667
AccI GTMKAC 1 cut(s) 836
AccII CGCG 2 cut(s) 128, 152
AciI CCGC 7 cut(s) 24, 128, 150, 320, 1033, 1211, 1286
AclWI GGATC 5 cut(s) 70, 398, 652, 1277, 1290
AcoI YGGCCR 1 cut(s) 129
AcsI RAATTY 1 cut(s) 1459
AfaI GTAC 3 cut(s) 50, 987, 1342
AfiI CCNNNNNNNGG 2 cut(s) 30, 667
AgeI ACCGGT 1 cut(s) 64
AgsI TTSAA 6 cut(s) 218, 605, 726, 828, 1327, 1382
AjiI CACGTC 1 cut(s) 283
AjnI CCWGG 2 cut(s) 660, 1277
AleI CACNNNNGTG 1 cut(s) 161
Alw21I GWGCWC 2 cut(s) 432, 1127
Alw26I GTCTC 3 cut(s) 989, 1014, 1065
Alw44I GTGCAC 1 cut(s) 428
AlwI GGATC 5 cut(s) 70, 398, 652, 1277, 1290
Ama87I CYCGRG 1 cut(s) 1068
AoxI GGCC 5 cut(s) 35, 95, 129, 957, 1062
ApaLI GTGCAC 1 cut(s) 428
ApeKI GCWGC 2 cut(s) 120, 1356
ApoI RAATTY 1 cut(s) 1459
AsiGI ACCGGT 1 cut(s) 64
Asp700I GAANNNNTTC 1 cut(s) 498
AspLEI GCGC 1 cut(s) 154
AspS9I GGNCC 1 cut(s) 67
AsuHPI GGTGA 3 cut(s) 1048, 1260, 1431
AsuII TTCGAA 1 cut(s) 967
AvaI CYCGRG 1 cut(s) 1068
AvaII GGWCC 1 cut(s) 67
BaeGI GKGCMC 1 cut(s) 432
BaeI ACNNNNGTAYC 4 cut(s) 32, 65, 1324, 1357
BamHI GGATCC 1 cut(s) 1282
BanII GRGCYC 2 cut(s) 1093, 1127
BbsI GAAGAC 1 cut(s) 1489
Bbv12I GWGCWC 2 cut(s) 432, 1127
BbvI GCAGC 2 cut(s) 132, 1368
BccI CCATC 5 cut(s) 263, 349, 376, 661, 1042
BceAI ACGGC 2 cut(s) 59, 116
BcgI CGANNNNNNTGC 2 cut(s) 1049, 1083
BciT130I CCWGG 2 cut(s) 662, 1279
BcoDI GTCTC 3 cut(s) 989, 1014, 1065
BfaI CTAG 4 cut(s) 521, 542, 573, 1094
BfmI CTRYAG 2 cut(s) 700, 837
BfuAI ACCTGC 1 cut(s) 739
BglI GCCNNNNNGGC 2 cut(s) 43, 94
BisI GCNGC 4 cut(s) 121, 129, 150, 1357
BlsI GCNGC 4 cut(s) 122, 130, 151, 1358
Bme1390I CCNGG 2 cut(s) 662, 1279
Bme18I GGWCC 1 cut(s) 67
BmeT110I CYCGRG 1 cut(s) 1068
BmgBI CACGTC 1 cut(s) 283
BmgT120I GGNCC 1 cut(s) 67
BmiI GGNNCC 2 cut(s) 69, 1284
BmrFI CCNGG 2 cut(s) 662, 1279
BmsI GCATC 3 cut(s) 735, 783, 1005
BpiI GAAGAC 1 cut(s) 1489
BplI GAGNNNNNCTC 2 cut(s) 1250, 1282
Bpu10I CCTNAGC 2 cut(s) 549, 943
Bpu14I TTCGAA 1 cut(s) 967
BpuEI CTTGAG 2 cut(s) 1141, 1246
BsaAI YACGTR 1 cut(s) 92
BsaI GGTCTC 1 cut(s) 1014
BsaJI CCNNGG 3 cut(s) 909, 1277, 1278
BsaWI WCCGGW 1 cut(s) 64
BsaXI ACNNNNNCTCC 2 cut(s) 767, 797
Bsc4I CCNNNNNNNGG 2 cut(s) 30, 667
Bse118I RCCGGY 1 cut(s) 64
Bse1I ACTGG 1 cut(s) 858
Bse3DI GCAATG 1 cut(s) 1010
BseBI CCWGG 2 cut(s) 662, 1279
BseDI CCNNGG 3 cut(s) 909, 1277, 1278
BseGI GGATG 5 cut(s) 271, 360, 643, 798, 813
BseLI CCNNNNNNNGG 2 cut(s) 30, 667
BseMI GCAATG 1 cut(s) 1010
BseMII CTCAG 6 cut(s) 215, 702, 752, 957, 984, 1025
BseNI ACTGG 1 cut(s) 858
BseRI GAGGAG 6 cut(s) 250, 275, 314, 859, 1058, 1117
BseSI GKGCMC 1 cut(s) 432
BseX3I CGGCCG 1 cut(s) 129
BseXI GCAGC 2 cut(s) 132, 1368
Bsh1236I CGCG 2 cut(s) 128, 152
Bsh1285I CGRYCG 1 cut(s) 132
BshFI GGCC 5 cut(s) 37, 97, 131, 959, 1064
BshTI ACCGGT 1 cut(s) 64
BsiEI CGRYCG 1 cut(s) 132
BsiHKAI GWGCWC 2 cut(s) 432, 1127
BsiHKCI CYCGRG 1 cut(s) 1068
BsiSI CCGG 2 cut(s) 65, 103
BslFI GGGAC 1 cut(s) 53
BslI CCNNNNNNNGG 2 cut(s) 30, 667
BsmAI GTCTC 3 cut(s) 989, 1014, 1065
BsmFI GGGAC 1 cut(s) 53
BsnI GGCC 5 cut(s) 37, 97, 131, 959, 1064
Bso31I GGTCTC 1 cut(s) 1014
BsoBI CYCGRG 1 cut(s) 1068
Bsp119I TTCGAA 1 cut(s) 967
Bsp1286I GDGCHC 3 cut(s) 432, 1093, 1127
Bsp143I GATC 6 cut(s) 75, 390, 657, 1084, 1282, 1436
BspACI CCGC 7 cut(s) 24, 128, 150, 320, 1033, 1211, 1286
BspANI GGCC 5 cut(s) 37, 97, 131, 959, 1064
BspCNI CTCAG 6 cut(s) 216, 703, 753, 956, 985, 1024
BspFNI CGCG 2 cut(s) 128, 152
BspHI TCATGA 1 cut(s) 925
BspLI GGNNCC 2 cut(s) 69, 1284
BspMI ACCTGC 1 cut(s) 739
BspPI GGATC 5 cut(s) 70, 398, 652, 1277, 1290
BspT104I TTCGAA 1 cut(s) 967
BspTNI GGTCTC 1 cut(s) 1014
BsrDI GCAATG 1 cut(s) 1010
BsrFI RCCGGY 1 cut(s) 64
BsrI ACTGG 1 cut(s) 858
BssAI RCCGGY 1 cut(s) 64
BssECI CCNNGG 3 cut(s) 909, 1277, 1278
BssMI GATC 6 cut(s) 75, 390, 657, 1084, 1282, 1436
BssT1I CCWWGG 1 cut(s) 909
Bst2UI CCWGG 2 cut(s) 662, 1279
Bst4CI ACNGT 2 cut(s) 19, 612
Bst6I CTCTTC 3 cut(s) 291, 489, 910
BstAPI GCANNNNNTGC 1 cut(s) 1024
BstBAI YACGTR 1 cut(s) 92
BstBI TTCGAA 1 cut(s) 967
BstC8I GCNNGC 5 cut(s) 182, 679, 1062, 1184, 1361
BstDEI CTNAG 7 cut(s) 224, 549, 711, 761, 943, 993, 1011
BstF5I GGATG 5 cut(s) 271, 360, 643, 798, 813
BstFNI CGCG 2 cut(s) 128, 152
BstHHI GCGC 1 cut(s) 154
BstKTI GATC 6 cut(s) 78, 393, 660, 1087, 1285, 1439
BstMAI GTCTC 3 cut(s) 989, 1014, 1065
BstMBI GATC 6 cut(s) 75, 390, 657, 1084, 1282, 1436
BstMCI CGRYCG 1 cut(s) 132
BstMWI GCNNNNNNNGC 4 cut(s) 43, 94, 863, 1024
BstNI CCWGG 2 cut(s) 662, 1279
BstSCI CCNGG 2 cut(s) 660, 1277
BstSFI CTRYAG 2 cut(s) 700, 837
BstSLI GKGCMC 1 cut(s) 432
BstUI CGCG 2 cut(s) 128, 152
BstV1I GCAGC 2 cut(s) 132, 1368
BstV2I GAAGAC 1 cut(s) 1489
BstX2I RGATCY 3 cut(s) 75, 657, 1282
BstXI CCANNNNNNTGG 1 cut(s) 163
BstYI RGATCY 3 cut(s) 75, 657, 1282
BstZI CGGCCG 1 cut(s) 129
BsuRI GGCC 5 cut(s) 37, 97, 131, 959, 1064
BtrI CACGTC 1 cut(s) 283
BtsCI GGATG 5 cut(s) 271, 360, 643, 798, 813
BveI ACCTGC 1 cut(s) 739
Cac8I GCNNGC 5 cut(s) 182, 679, 1062, 1184, 1361
CciI TCATGA 1 cut(s) 925
CfoI GCGC 1 cut(s) 154
Cfr10I RCCGGY 1 cut(s) 64
Cfr13I GGNCC 1 cut(s) 67
CseI GACGC 1 cut(s) 112
Csp6I GTAC 3 cut(s) 49, 986, 1341
CspAI ACCGGT 1 cut(s) 64
CviAII CATG 4 cut(s) 137, 926, 1208, 1404
CviQI GTAC 3 cut(s) 49, 986, 1341
DdeI CTNAG 7 cut(s) 224, 549, 711, 761, 943, 993, 1011
DpnI GATC 6 cut(s) 77, 392, 659, 1086, 1284, 1438
DpnII GATC 6 cut(s) 75, 390, 657, 1084, 1282, 1436
EaeI YGGCCR 1 cut(s) 129
EagI CGGCCG 1 cut(s) 129
Eam1104I CTCTTC 3 cut(s) 291, 489, 910
EarI CTCTTC 3 cut(s) 291, 489, 910
EciI GGCGGA 1 cut(s) 335
Ecl136II GAGCTC 1 cut(s) 1125
EclXI CGGCCG 1 cut(s) 129
Eco130I CCWWGG 1 cut(s) 909
Eco147I AGGCCT 2 cut(s) 97, 1064
Eco24I GRGCYC 2 cut(s) 1093, 1127
Eco31I GGTCTC 1 cut(s) 1014
Eco47I GGWCC 1 cut(s) 67
Eco52I CGGCCG 1 cut(s) 129
Eco53kI GAGCTC 1 cut(s) 1125
Eco88I CYCGRG 1 cut(s) 1068
EcoICRI GAGCTC 1 cut(s) 1125
EcoRII CCWGG 2 cut(s) 660, 1277
EcoT14I CCWWGG 1 cut(s) 909
EcoT38I GRGCYC 2 cut(s) 1093, 1127
ErhI CCWWGG 1 cut(s) 909
FaeI CATG 4 cut(s) 140, 929, 1211, 1407
FalI AAGNNNNNCTT 2 cut(s) 582, 614
FaqI GGGAC 1 cut(s) 53
FatI CATG 4 cut(s) 136, 925, 1207, 1403
FauI CCCGC 1 cut(s) 1026
FblI GTMKAC 1 cut(s) 836
Fnu4HI GCNGC 4 cut(s) 121, 129, 150, 1357
FokI GGATG 5 cut(s) 278, 367, 650, 805, 820
FriOI GRGCYC 2 cut(s) 1093, 1127
Fsp4HI GCNGC 4 cut(s) 121, 129, 150, 1357
FspBI CTAG 4 cut(s) 521, 542, 573, 1094
GlaI GCGC 1 cut(s) 153
GluI GCNGC 4 cut(s) 121, 129, 150, 1357
HaeIII GGCC 5 cut(s) 37, 97, 131, 959, 1064
HapII CCGG 2 cut(s) 65, 103
HgaI GACGC 1 cut(s) 112
HhaI GCGC 1 cut(s) 154
Hin1II CATG 4 cut(s) 140, 929, 1211, 1407
Hin6I GCGC 1 cut(s) 152
HinP1I GCGC 1 cut(s) 152
HindIII AAGCTT 2 cut(s) 591, 679
HinfI GANTC 8 cut(s) 412, 499, 538, 757, 882, 887, 1097, 1400
HpaII CCGG 2 cut(s) 65, 103
HphI GGTGA 3 cut(s) 1048, 1260, 1431
Hpy166II GTNNAC 2 cut(s) 430, 837
Hpy188I TCNGA 8 cut(s) 516, 567, 637, 762, 1014, 1102, 1165, 1431
Hpy188III TCNNGA 5 cut(s) 926, 945, 1068, 1070, 1448
Hpy8I GTNNAC 2 cut(s) 430, 837
Hpy99I CGWCG 1 cut(s) 128
HpyCH4III ACNGT 2 cut(s) 19, 612
HpyCH4IV ACGT 4 cut(s) 91, 282, 936, 949
HpyF10VI GCNNNNNNNGC 4 cut(s) 43, 94, 863, 1024
HpyF3I CTNAG 7 cut(s) 224, 549, 711, 761, 943, 993, 1011
HpySE526I ACGT 4 cut(s) 91, 282, 936, 949
Hsp92II CATG 4 cut(s) 140, 929, 1211, 1407
HspAI GCGC 1 cut(s) 152
Kzo9I GATC 6 cut(s) 75, 390, 657, 1084, 1282, 1436
LmnI GCTCC 3 cut(s) 545, 1130, 1213
Lsp1109I GCAGC 2 cut(s) 132, 1368
LweI GCATC 3 cut(s) 735, 783, 1005
MaeI CTAG 4 cut(s) 521, 542, 573, 1094
MaeII ACGT 4 cut(s) 91, 282, 936, 949
MaeIII GTNAC 4 cut(s) 19, 475, 575, 1319
MalI GATC 6 cut(s) 77, 392, 659, 1086, 1284, 1438
MboI GATC 6 cut(s) 75, 390, 657, 1084, 1282, 1436
MfeI CAATTG 1 cut(s) 506
MflI RGATCY 3 cut(s) 75, 657, 1282
MhlI GDGCHC 3 cut(s) 432, 1093, 1127
MluCI AATT 5 cut(s) 343, 506, 1055, 1261, 1459
MlyI GAGTC 3 cut(s) 406, 1091, 1394
MroXI GAANNNNTTC 1 cut(s) 498
MseI TTAA 6 cut(s) 375, 399, 417, 651, 800, 1531
MslI CAYNNNNRTG 1 cut(s) 161
MspA1I CMGCKG 1 cut(s) 425
MspI CCGG 2 cut(s) 65, 103
MspR9I CCNGG 2 cut(s) 662, 1279
MunI CAATTG 1 cut(s) 506
MvaI CCWGG 2 cut(s) 662, 1279
MvnI CGCG 2 cut(s) 128, 152
MwoI GCNNNNNNNGC 4 cut(s) 43, 94, 863, 1024
NdeII GATC 6 cut(s) 75, 390, 657, 1084, 1282, 1436
NlaIII CATG 4 cut(s) 140, 929, 1211, 1407
NlaIV GGNNCC 2 cut(s) 69, 1284
NmuCI GTSAC 1 cut(s) 1319
NspV TTCGAA 1 cut(s) 967
OliI CACNNNNGTG 1 cut(s) 161
PaeR7I CTCGAG 1 cut(s) 1068
PagI TCATGA 1 cut(s) 925
PaqCI CACCTGC 1 cut(s) 739
PasI CCCWGGG 1 cut(s) 1278
PceI AGGCCT 2 cut(s) 97, 1064
PcsI WCGNNNNNNNCGW 1 cut(s) 891
PdmI GAANNNNTTC 1 cut(s) 498
PfeI GAWTC 5 cut(s) 499, 538, 757, 882, 887
PflMI CCANNNNNTGG 1 cut(s) 667
PfoI TCCNGGA 1 cut(s) 660
PinAI ACCGGT 1 cut(s) 64
PkrI GCNGC 4 cut(s) 122, 130, 151, 1358
PleI GAGTC 3 cut(s) 406, 1091, 1394
PpsI GAGTC 3 cut(s) 406, 1091, 1394
Ppu21I YACGTR 1 cut(s) 92
Psp124BI GAGCTC 1 cut(s) 1127
Psp6I CCWGG 2 cut(s) 660, 1277
PspGI CCWGG 2 cut(s) 660, 1277
PspN4I GGNNCC 2 cut(s) 69, 1284
PspPI GGNCC 1 cut(s) 67
PsuI RGATCY 3 cut(s) 75, 657, 1282
PvuII CAGCTG 1 cut(s) 425
RsaI GTAC 3 cut(s) 50, 987, 1342
RsaNI GTAC 3 cut(s) 49, 986, 1341
RseI CAYNNNNRTG 1 cut(s) 161
SacI GAGCTC 1 cut(s) 1127
SaqAI TTAA 6 cut(s) 375, 399, 417, 651, 800, 1531
SatI GCNGC 4 cut(s) 121, 129, 150, 1357
Sau3AI GATC 6 cut(s) 75, 390, 657, 1084, 1282, 1436
Sau96I GGNCC 1 cut(s) 67
SchI GAGTC 3 cut(s) 406, 1091, 1394
ScrFI CCNGG 2 cut(s) 662, 1279
SduI GDGCHC 3 cut(s) 432, 1093, 1127
SfaNI GCATC 3 cut(s) 735, 783, 1005
SfcI CTRYAG 2 cut(s) 700, 837
Sfr274I CTCGAG 1 cut(s) 1068
SfuI TTCGAA 1 cut(s) 967
SinI GGWCC 1 cut(s) 67
SlaI CTCGAG 1 cut(s) 1068
SmiMI CAYNNNNRTG 1 cut(s) 161
SmlI CTYRAG 3 cut(s) 1068, 1120, 1225
SmoI CTYRAG 3 cut(s) 1068, 1120, 1225
Sse9I AATT 5 cut(s) 343, 506, 1055, 1261, 1459
SseBI AGGCCT 2 cut(s) 97, 1064
SsiI CCGC 7 cut(s) 24, 128, 150, 320, 1033, 1211, 1286
SspMI CTAG 4 cut(s) 521, 542, 573, 1094
SstI GAGCTC 1 cut(s) 1127
StuI AGGCCT 2 cut(s) 97, 1064
StyD4I CCNGG 2 cut(s) 660, 1277
StyI CCWWGG 1 cut(s) 909
TaaI ACNGT 2 cut(s) 19, 612
TaiI ACGT 4 cut(s) 94, 285, 939, 952
TaqI TCGA 6 cut(s) 582, 885, 967, 1069, 1087, 1447
TaqII GACCGA 1 cut(s) 1307
TasI AATT 5 cut(s) 343, 506, 1055, 1261, 1459
TauI GCSGC 2 cut(s) 131, 152
TfiI GAWTC 5 cut(s) 499, 538, 757, 882, 887
Tru1I TTAA 6 cut(s) 375, 399, 417, 651, 800, 1531
Tru9I TTAA 6 cut(s) 375, 399, 417, 651, 800, 1531
TseFI GTSAC 1 cut(s) 1319
TseI GCWGC 2 cut(s) 120, 1356
Tsp45I GTSAC 1 cut(s) 1319
TspDTI ATGAA 3 cut(s) 125, 824, 1149
Van91I CCANNNNNTGG 1 cut(s) 667
VneI GTGCAC 1 cut(s) 428
VpaK11BI GGWCC 1 cut(s) 67
XapI RAATTY 1 cut(s) 1459
XhoI CTCGAG 1 cut(s) 1068
XmiI GTMKAC 1 cut(s) 836
XmnI GAANNNNTTC 1 cut(s) 498
XspI CTAG 4 cut(s) 521, 542, 573, 1094
Using CommOnly database (standard laboratory enzymes). Scanned on CDS sequence.