FvH4_2g23720

protein trichome birefringence-like

Design Primers CRISPR Design

Basic Information

Type: gene
Biological Identity
fragaria_vesca
Fvb2
OG0015547
Physical Location & Seq
Reverse (-)
19422064 .. 19425359
3296 bp
Loading structure...
UTR
Exon/CDS
Intron
FvH4_2g23720.t1

Sequence Viewer

Length: 1296 bp
ATGACCCAGTTGGAGTTGTTCAAGAAAATCAAGCGCTTCAATCCTCTAGAGCCTTCTCTAAGCATTCTTGGGTTTTTCTTGGTCACTGTTCTGCTCATTGGGTGCTTCTTCTACTTGGATTACCGCACAGTTGTCCATGGGATTCGCTCTCGTCTTTTGCATGGTTCAGGTGGGAAAGCACGGTCAGGGTTTCTTGATCAGGGGAATGAAGGGTGTGATGTGTTTGATGGGAACTGGGTTTGGGATGAGAGCTATCCTTTGTATCAGTCTCAGAATTGCTCCTTTTTGGATGGAGGGTTTAGGTGTTCTGAGAATGGGAGACCTGACAGCTTCTATACCAAGTGGCGTTGGCAACCCAAAGATTGCAACTTGCCCAGATTCGATGCAGGGATGATGTTGGAAAAGCTTCGAAATCGACGTCTTGTATTTGCCGGTGACTCGATTGGAAGGAACCAGTGGGAGTCTCTTCTCTGCATGCTCTCTTCTGTAATTCCTAACAAGACTTCAATCTATGAGGTGAATGGGACGCCAATTACGAAGCACTCAGGCTTTTTGGTCTTCAAGTTTGAGGACTTCAACTGCACAGTGGAATACTACAGGTCACCATTTCTAGTCTTTCAGGGCCATCCTCCAGCTGGTGCCCCAGAGGAGGTGAAGATGACTTTAAAAGTGGATCAAATGGACTGGCTTTCCCTCCACTGGAGGGATGCGGATGTGCTAGTCTTGAATTCCGGGCATTGGTGGAATTATGGGAAGACCATTAGAGAGGGCTGCTACTTTCAAGAAGGGGAGGAGATAAAGATGAATATGACTGTTAATACTGCATACCGGAAATCAATGGAAACTTTGATCGATTGGATTGATAGTGAAGTCAACATGAGCAAGACTTATGTACTTTTTCGAACTTATGCTCCTGTGCATTTCAGTGGTGGTGATTGGAATTCTGGTGGTGGCTGTCACTTGGAGACTCTACCTGGATTAGGCCCATTGCCTGTTTTATCTGACAGTGTTTACACGACTATCTTTGATGTGTTATCTGAGCGATCAAATGAGTCACACGTAGGAAAAGTCGATTTTCTGAATGTAACAAACATGTCTCTGTGGAGAAAAGATGCCCATGCTTCTCTGTACTATTTAGGGCCAGATGGACCAGCCTCAATCAACCATCAAGACTGCAGCCACTGGTGCCTACCTGGCGTCCCTGATTCATGGAATGAACTTCTTTATGCCCTTTTCCTTAAACAGGAATCCATTCGCTCAGGAAATTCAACCAAATCTTCCCAGTTTCCATTGTGA
Functional Annotation
Gene Ontology
Molecular Function
Biological Process
Cellular Component
No ontology terms assigned.
KEGG Pathways
Metabolic & Signaling
No pathways identified.
Pfam Domains
Protein Families
PC-Esterase PMR5N

Protein Analysis

432

Amino Acids

49.24

Weight (kDa)

5.96

Isoelectric Point (pI)

44.19

Instability Index
Protein Domains (Pfam)
Domain Name Pfam ID Position E-value Description
PMR5N PF14416 71 - 123 1.4e-22 PMR5 N terminal Domain
PC-Esterase PF13839 124 - 410 1.1e-95 GDSL/SGNH-like Acyl-Esterase family found in Pmr5 and Cas1p
Hydrophobicity Profile (Kyte-Doolittle)
AI Structure Prediction Report
Calculating structure properties...

Genomic Context

Gene Family Tree

Species Filter
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Publication-ready
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Orthologous Genes (Group: OG0015547)

Species Orthologous Gene IDs
fragaria_vesca FvH4_2g23720
malus_domestica MD03G1266900.v1.1 MD11G1287300.v1.1
prunus_persica Prupe.8G242600_v2.0.a1
pyrus_communis pycom03g21320 pycom11g25470
rosa_chinensis RchiOBHm_Chr6g0291221
rosa_laevigata RLG00000012132
rosa_multiflora Rmu_sc0013203.1_g000020
rosa_roxburghii Rroxscaffold_7G00176240
rosa_rugosa Rorug06G0218500
rosa_samantha Rh6AG329300 Rh6BG336500 Rh6CG342800 Rh6DG329800
rosa_wichuraiana Rw6G028630

Restriction Enzyme Sites

1 / 10
Enzyme Recognition Site Cut Count Positions (bp)
AatII GACGTC 1 cut(s) 421
AccB1I GGYRCC 2 cut(s) 638, 1185
AciI CCGC 2 cut(s) 124, 710
AclWI GGATC 1 cut(s) 681
AcsI RAATTY 3 cut(s) 727, 940, 1264
AcyI GRCGYC 3 cut(s) 418, 527, 1197
AfaI GTAC 2 cut(s) 894, 1130
AfeI AGCGCT 1 cut(s) 35
AfiI CCNNNNNNNGG 7 cut(s) 635, 649, 699, 703, 738, 980, 1243
AflIII ACRYGT 2 cut(s) 1057, 1092
AgsI TTSAA 8 cut(s) 22, 40, 507, 562, 577, 727, 782, 1269
AjnI CCWGG 2 cut(s) 973, 1192
AloI GAACNNNNNNTCC 2 cut(s) 895, 927
AluBI AGCT 4 cut(s) 252, 330, 406, 635
AluI AGCT 4 cut(s) 252, 330, 406, 635
Alw26I GTCTC 5 cut(s) 273, 313, 468, 959, 1101
AlwI GGATC 1 cut(s) 681
AlwNI CAGNNNCTG 1 cut(s) 1182
Aor51HI AGCGCT 1 cut(s) 35
AoxI GGCC 3 cut(s) 622, 982, 1139
ApeKI GCWGC 2 cut(s) 771, 1176
ApoI RAATTY 3 cut(s) 727, 940, 1264
ArsI GACNNNNNNTTYG 2 cut(s) 403, 435
Asp700I GAANNNNTTC 2 cut(s) 405, 1251
AspLEI GCGC 1 cut(s) 36
AspS9I GGNCC 4 cut(s) 622, 983, 1139, 1148
AsuC2I CCSGG 1 cut(s) 733
AsuHPI GGTGA 5 cut(s) 446, 529, 594, 664, 944
AsuII TTCGAA 2 cut(s) 409, 901
AvaII GGWCC 1 cut(s) 1148
BaeGI GKGCMC 1 cut(s) 643
BanI GGYRCC 2 cut(s) 638, 1185
BbsI GAAGAC 2 cut(s) 550, 761
BbvI GCAGC 2 cut(s) 758, 1188
BccI CCATC 5 cut(s) 221, 284, 633, 1139, 1173
BciT130I CCWGG 2 cut(s) 975, 1194
BclI TGATCA 1 cut(s) 196
BcnI CCSGG 1 cut(s) 733
BcoDI GTCTC 5 cut(s) 273, 313, 468, 959, 1101
BfaI CTAG 3 cut(s) 47, 611, 719
BfmI CTRYAG 2 cut(s) 595, 1174
BfoI RGCGCY 1 cut(s) 37
BglI GCCNNNNNGGC 1 cut(s) 1194
BisI GCNGC 2 cut(s) 772, 1177
BlsI GCNGC 2 cut(s) 773, 1178
Bme1390I CCNGG 3 cut(s) 733, 975, 1194
Bme18I GGWCC 1 cut(s) 1148
BmgT120I GGNCC 4 cut(s) 622, 983, 1139, 1148
BmiI GGNNCC 3 cut(s) 452, 640, 1187
BmrFI CCNGG 3 cut(s) 733, 975, 1194
BmrI ACTGGG 2 cut(s) 244, 1276
BmsI GCATC 3 cut(s) 373, 697, 1102
BmuI ACTGGG 2 cut(s) 244, 1276
BpiI GAAGAC 2 cut(s) 550, 761
BpmI CTGGAG 2 cut(s) 615, 721
Bpu10I CCTNAGC 1 cut(s) 1258
Bpu14I TTCGAA 2 cut(s) 409, 901
BpuMI CCSGG 1 cut(s) 733
Bsa29I ATCGAT 1 cut(s) 852
BsaAI YACGTR 1 cut(s) 1060
BsaHI GRCGYC 3 cut(s) 418, 527, 1197
BsaI GGTCTC 1 cut(s) 313
BsaJI CCNNGG 1 cut(s) 136
BsaWI WCCGGW 1 cut(s) 828
BsaXI ACNNNNNCTCC 2 cut(s) 895, 925
Bsc4I CCNNNNNNNGG 7 cut(s) 635, 649, 699, 703, 738, 980, 1243
Bse118I RCCGGY 1 cut(s) 431
Bse1I ACTGG 7 cut(s) 7, 239, 454, 689, 704, 1187, 1282
Bse3DI GCAATG 1 cut(s) 986
BseBI CCWGG 2 cut(s) 975, 1194
BseCI ATCGAT 1 cut(s) 852
BseDI CCNNGG 1 cut(s) 136
BseGI GGATG 6 cut(s) 250, 295, 396, 625, 712, 718
BseLI CCNNNNNNNGG 7 cut(s) 635, 649, 699, 703, 738, 980, 1243
BseMI GCAATG 1 cut(s) 986
BseMII CTCAG 5 cut(s) 284, 300, 558, 1029, 1272
BseNI ACTGG 7 cut(s) 7, 239, 454, 689, 704, 1187, 1282
BseRI GAGGAG 2 cut(s) 662, 806
BseSI GKGCMC 1 cut(s) 643
BseXI GCAGC 2 cut(s) 758, 1188
BsgI GTGCAG 1 cut(s) 565
BshFI GGCC 3 cut(s) 624, 984, 1141
BshNI GGYRCC 2 cut(s) 638, 1185
BshVI ATCGAT 1 cut(s) 852
BsiSI CCGG 3 cut(s) 432, 732, 829
BslFI GGGAC 2 cut(s) 538, 1184
BslI CCNNNNNNNGG 7 cut(s) 635, 649, 699, 703, 738, 980, 1243
BsmAI GTCTC 5 cut(s) 273, 313, 468, 959, 1101
BsmFI GGGAC 2 cut(s) 538, 1184
BsmI GAATGC 1 cut(s) 63
BsnI GGCC 3 cut(s) 624, 984, 1141
Bso31I GGTCTC 1 cut(s) 313
Bsp119I TTCGAA 2 cut(s) 409, 901
Bsp1286I GDGCHC 1 cut(s) 643
Bsp143I GATC 4 cut(s) 196, 673, 849, 1043
Bsp19I CCATGG 1 cut(s) 136
BspACI CCGC 2 cut(s) 124, 710
BspANI GGCC 3 cut(s) 624, 984, 1141
BspCNI CTCAG 5 cut(s) 283, 301, 557, 1030, 1271
BspDI ATCGAT 1 cut(s) 852
BspLI GGNNCC 3 cut(s) 452, 640, 1187
BspMAI CTGCAG 1 cut(s) 1178
BspPI GGATC 1 cut(s) 681
BspT104I TTCGAA 2 cut(s) 409, 901
BspT107I GGYRCC 2 cut(s) 638, 1185
BspTNI GGTCTC 1 cut(s) 313
BsrDI GCAATG 1 cut(s) 986
BsrFI RCCGGY 1 cut(s) 431
BsrI ACTGG 7 cut(s) 7, 239, 454, 689, 704, 1187, 1282
BssAI RCCGGY 1 cut(s) 431
BssECI CCNNGG 1 cut(s) 136
BssMI GATC 4 cut(s) 196, 673, 849, 1043
BssNI GRCGYC 3 cut(s) 418, 527, 1197
BssT1I CCWWGG 1 cut(s) 136
Bst2UI CCWGG 2 cut(s) 975, 1194
Bst4CI ACNGT 6 cut(s) 88, 130, 183, 586, 814, 1007
Bst6I CTCTTC 2 cut(s) 471, 487
BstACI GRCGYC 3 cut(s) 418, 527, 1197
BstBAI YACGTR 1 cut(s) 1060
BstBI TTCGAA 2 cut(s) 409, 901
BstC8I GCNNGC 1 cut(s) 476
BstDEI CTNAG 6 cut(s) 59, 270, 309, 544, 1038, 1258
BstDSI CCRYGG 1 cut(s) 136
BstEII GGTNACC 1 cut(s) 600
BstENI CCTNNNNNAGG 2 cut(s) 978, 1241
BstF5I GGATG 6 cut(s) 250, 295, 396, 625, 712, 718
BstH2I RGCGCY 1 cut(s) 37
BstHHI GCGC 1 cut(s) 36
BstKTI GATC 4 cut(s) 199, 676, 852, 1046
BstMAI GTCTC 5 cut(s) 273, 313, 468, 959, 1101
BstMBI GATC 4 cut(s) 196, 673, 849, 1043
BstMWI GCNNNNNNNGC 2 cut(s) 1185, 1194
BstNI CCWGG 2 cut(s) 975, 1194
BstNSI RCATGY 2 cut(s) 478, 1096
BstPI GGTNACC 1 cut(s) 600
BstSCI CCNGG 3 cut(s) 731, 973, 1192
BstSFI CTRYAG 2 cut(s) 595, 1174
BstSLI GKGCMC 1 cut(s) 643
BstV1I GCAGC 2 cut(s) 758, 1188
BstV2I GAAGAC 2 cut(s) 550, 761
Bsu15I ATCGAT 1 cut(s) 852
BsuRI GGCC 3 cut(s) 624, 984, 1141
BsuTUI ATCGAT 1 cut(s) 852
BtgI CCRYGG 1 cut(s) 136
BtsCI GGATG 6 cut(s) 250, 295, 396, 625, 712, 718
BtsIMutI CAGTG 7 cut(s) 84, 461, 591, 697, 931, 1012, 1180
Cac8I GCNNGC 1 cut(s) 476
CaiI CAGNNNCTG 1 cut(s) 1182
CfoI GCGC 1 cut(s) 36
Cfr10I RCCGGY 1 cut(s) 431
Cfr13I GGNCC 4 cut(s) 622, 983, 1139, 1148
ClaI ATCGAT 1 cut(s) 852
CseI GACGC 2 cut(s) 535, 1186
Csp6I GTAC 2 cut(s) 893, 1129
CviAII CATG 7 cut(s) 137, 161, 475, 877, 1093, 1118, 1209
CviQI GTAC 2 cut(s) 893, 1129
DdeI CTNAG 6 cut(s) 59, 270, 309, 544, 1038, 1258
DpnI GATC 4 cut(s) 198, 675, 851, 1045
DpnII GATC 4 cut(s) 196, 673, 849, 1043
DraI TTTAAA 1 cut(s) 666
Eam1104I CTCTTC 2 cut(s) 471, 487
EarI CTCTTC 2 cut(s) 471, 487
Eco130I CCWWGG 1 cut(s) 136
Eco31I GGTCTC 1 cut(s) 313
Eco47I GGWCC 1 cut(s) 1148
Eco47III AGCGCT 1 cut(s) 35
Eco91I GGTNACC 1 cut(s) 600
EcoNI CCTNNNNNAGG 2 cut(s) 978, 1241
EcoO65I GGTNACC 1 cut(s) 600
EcoRI GAATTC 2 cut(s) 727, 940
EcoRII CCWGG 2 cut(s) 973, 1192
EcoT14I CCWWGG 1 cut(s) 136
ErhI CCWWGG 1 cut(s) 136
FaeI CATG 7 cut(s) 140, 164, 478, 880, 1096, 1121, 1212
FaqI GGGAC 2 cut(s) 538, 1184
FatI CATG 7 cut(s) 136, 160, 474, 876, 1092, 1117, 1208
FbaI TGATCA 1 cut(s) 196
Fnu4HI GCNGC 2 cut(s) 772, 1177
FokI GGATG 6 cut(s) 257, 302, 403, 612, 719, 725
Fsp4HI GCNGC 2 cut(s) 772, 1177
FspBI CTAG 3 cut(s) 47, 611, 719
GlaI GCGC 1 cut(s) 35
GluI GCNGC 2 cut(s) 772, 1177
GsuI CTGGAG 2 cut(s) 615, 721
HaeII RGCGCY 1 cut(s) 37
HaeIII GGCC 3 cut(s) 624, 984, 1141
HapII CCGG 3 cut(s) 432, 732, 829
HgaI GACGC 2 cut(s) 535, 1186
HhaI GCGC 1 cut(s) 36
Hin1I GRCGYC 3 cut(s) 418, 527, 1197
Hin1II CATG 7 cut(s) 140, 164, 478, 880, 1096, 1121, 1212
Hin6I GCGC 1 cut(s) 34
HinP1I GCGC 1 cut(s) 34
HincII GTYRAC 1 cut(s) 874
HindII GTYRAC 1 cut(s) 874
HindIII AAGCTT 1 cut(s) 404
HinfI GANTC 8 cut(s) 142, 378, 437, 461, 967, 1052, 1205, 1247
HpaII CCGG 3 cut(s) 432, 732, 829
HphI GGTGA 5 cut(s) 446, 529, 594, 664, 944
Hpy166II GTNNAC 2 cut(s) 874, 1012
Hpy188I TCNGA 5 cut(s) 273, 310, 1003, 1039, 1080
Hpy188III TCNNGA 7 cut(s) 22, 47, 194, 724, 782, 1169, 1260
Hpy8I GTNNAC 2 cut(s) 874, 1012
Hpy99I CGWCG 1 cut(s) 420
HpyAV CCTTC 4 cut(s) 63, 203, 441, 779
HpyCH4III ACNGT 6 cut(s) 88, 130, 183, 586, 814, 1007
HpyCH4IV ACGT 2 cut(s) 418, 1059
HpyCH4V TGCA 8 cut(s) 160, 366, 386, 474, 582, 824, 919, 1176
HpyF10VI GCNNNNNNNGC 2 cut(s) 1185, 1194
HpyF3I CTNAG 6 cut(s) 59, 270, 309, 544, 1038, 1258
HpySE526I ACGT 2 cut(s) 418, 1059
Hsp92I GRCGYC 3 cut(s) 418, 527, 1197
Hsp92II CATG 7 cut(s) 140, 164, 478, 880, 1096, 1121, 1212
HspAI GCGC 1 cut(s) 34
Ksp22I TGATCA 1 cut(s) 196
Kzo9I GATC 4 cut(s) 196, 673, 849, 1043
LmnI GCTCC 2 cut(s) 284, 916
Lsp1109I GCAGC 2 cut(s) 758, 1188
LweI GCATC 3 cut(s) 373, 697, 1102
MaeI CTAG 3 cut(s) 47, 611, 719
MaeII ACGT 2 cut(s) 418, 1059
MaeIII GTNAC 6 cut(s) 82, 434, 600, 956, 1053, 1084
MalI GATC 4 cut(s) 198, 675, 851, 1045
MboI GATC 4 cut(s) 196, 673, 849, 1043
MboII GAAGA 7 cut(s) 100, 458, 474, 550, 667, 766, 1269
MhlI GDGCHC 1 cut(s) 643
MluCI AATT 7 cut(s) 274, 489, 531, 727, 745, 940, 1264
MlyI GAGTC 4 cut(s) 431, 470, 961, 1061
MmeI TCCRAC 1 cut(s) 378
MroXI GAANNNNTTC 2 cut(s) 405, 1251
MseI TTAA 3 cut(s) 665, 816, 1239
MslI CAYNNNNRTG 1 cut(s) 924
MspA1I CMGCKG 1 cut(s) 635
MspI CCGG 3 cut(s) 432, 732, 829
MspR9I CCNGG 3 cut(s) 733, 975, 1194
Mva1269I GAATGC 1 cut(s) 63
MvaI CCWGG 2 cut(s) 975, 1194
MwoI GCNNNNNNNGC 2 cut(s) 1185, 1194
NciI CCSGG 1 cut(s) 733
NcoI CCATGG 1 cut(s) 136
NdeII GATC 4 cut(s) 196, 673, 849, 1043
NlaIII CATG 7 cut(s) 140, 164, 478, 880, 1096, 1121, 1212
NlaIV GGNNCC 3 cut(s) 452, 640, 1187
NmuCI GTSAC 5 cut(s) 82, 434, 600, 956, 1053
NspI RCATGY 2 cut(s) 478, 1096
NspV TTCGAA 2 cut(s) 409, 901
PaeI GCATGC 1 cut(s) 478
PciI ACATGT 1 cut(s) 1092
PcsI WCGNNNNNNNCGW 2 cut(s) 415, 533
PctI GAATGC 1 cut(s) 63
PdmI GAANNNNTTC 2 cut(s) 405, 1251
PfeI GAWTC 4 cut(s) 142, 378, 1205, 1247
PkrI GCNGC 2 cut(s) 773, 1178
PleI GAGTC 4 cut(s) 431, 469, 961, 1060
PpsI GAGTC 4 cut(s) 431, 469, 961, 1060
Ppu21I YACGTR 1 cut(s) 1060
PscI ACATGT 1 cut(s) 1092
Psp6I CCWGG 2 cut(s) 973, 1192
PspEI GGTNACC 1 cut(s) 600
PspGI CCWGG 2 cut(s) 973, 1192
PspN4I GGNNCC 3 cut(s) 452, 640, 1187
PspPI GGNCC 4 cut(s) 622, 983, 1139, 1148
PstI CTGCAG 1 cut(s) 1178
PstNI CAGNNNCTG 1 cut(s) 1182
PvuII CAGCTG 1 cut(s) 635
RsaI GTAC 2 cut(s) 894, 1130
RsaNI GTAC 2 cut(s) 893, 1129
RseI CAYNNNNRTG 1 cut(s) 924
SaqAI TTAA 3 cut(s) 665, 816, 1239
SatI GCNGC 2 cut(s) 772, 1177
Sau3AI GATC 4 cut(s) 196, 673, 849, 1043
Sau96I GGNCC 4 cut(s) 622, 983, 1139, 1148
SchI GAGTC 4 cut(s) 431, 470, 961, 1061
ScrFI CCNGG 3 cut(s) 733, 975, 1194
SduI GDGCHC 1 cut(s) 643
SfaNI GCATC 3 cut(s) 373, 697, 1102
SfcI CTRYAG 2 cut(s) 595, 1174
SfuI TTCGAA 2 cut(s) 409, 901
SinI GGWCC 1 cut(s) 1148
SmiMI CAYNNNNRTG 1 cut(s) 924
SphI GCATGC 1 cut(s) 478
Sse9I AATT 7 cut(s) 274, 489, 531, 727, 745, 940, 1264
SsiI CCGC 2 cut(s) 124, 710
SspMI CTAG 3 cut(s) 47, 611, 719
StyD4I CCNGG 3 cut(s) 731, 973, 1192
StyI CCWWGG 1 cut(s) 136
TaaI ACNGT 6 cut(s) 88, 130, 183, 586, 814, 1007
TaiI ACGT 2 cut(s) 421, 1062
TaqI TCGA 7 cut(s) 381, 409, 415, 440, 852, 901, 1071
TasI AATT 7 cut(s) 274, 489, 531, 727, 745, 940, 1264
TatI WGTACW 2 cut(s) 892, 1128
TfiI GAWTC 4 cut(s) 142, 378, 1205, 1247
Tru1I TTAA 3 cut(s) 665, 816, 1239
Tru9I TTAA 3 cut(s) 665, 816, 1239
TscAI CASTG 7 cut(s) 91, 461, 591, 704, 931, 1012, 1187
TseFI GTSAC 5 cut(s) 82, 434, 600, 956, 1053
TseI GCWGC 2 cut(s) 771, 1176
Tsp45I GTSAC 5 cut(s) 82, 434, 600, 956, 1053
TspDTI ATGAA 4 cut(s) 222, 818, 1197, 1230
TspRI CASTG 7 cut(s) 91, 461, 591, 704, 931, 1012, 1187
VpaK11BI GGWCC 1 cut(s) 1148
XagI CCTNNNNNAGG 2 cut(s) 978, 1241
XapI RAATTY 3 cut(s) 727, 940, 1264
XbaI TCTAGA 1 cut(s) 46
XceI RCATGY 2 cut(s) 478, 1096
XcmI CCANNNNNNNNNTGG 1 cut(s) 632
XmnI GAANNNNTTC 2 cut(s) 405, 1251
XspI CTAG 3 cut(s) 47, 611, 719
ZraI GACGTC 1 cut(s) 419
Using CommOnly database (standard laboratory enzymes). Scanned on CDS sequence.