Basic Information

Type: gene

Biological Identity

Species

malus_domestica

Chromosome

Chr11

Orthogroup ID

OG0016794

Physical Location & Seq

Genomic Coordinates Reverse (-)

20022600 .. 20023844

Gene Structure 1245 bp

Loading structure...

UTR

Exon/CDS

Intron

Transcript / Protein ID

MD11G1173600.v1.1.491

Sequence Viewer

CDS Protein Promoter (2k)

Length: 1245 bp

ATGGAGGAATTCGTTATTCCTCCGCATTTTCTCTGCCCAATTTCATTAGAAATCATGAGAGATCCGGTCACGATCTCAACCGGTATAACGTACGATCGAGAAAGCATAGAGCAGTGGTGGTTCACATCGAAGAACAACACTTGCCCCGTAAGCAAAAAGTCATTGTCCGACGTTGACGCAGATCTCACGCCCAACCACACTCTCCGCCGCATAATCCAAGCTTGGTGCACTCTCAACTCTTCGTATGGCGTCGAGCTGATACCAACTCCTAAATCCGCGCTCGACAAATCCCAGGTCGTGATGCTCATCGAACAAGCCAAGAAAATCCCTCACACCAATCTCCATTGCCTCCAAAGGCTCAGATCCATCGCCATTCAAGGTGAGAGGAACAAGAACTGTTTGGAGGCAGCAGGAGCAATCGAATACGTGGCCTCCATCATCAACAAAAGTCAATCCGATTCATCTACAGTCGAAGCGGCTATTGAAGTCCTTTTTCATTTCAAAAGCTCGGAATCTCGTCTCAAAGATCTCATGAGCAATGACAGCAACAATAATAACATCACGGAGTCTCTGGTCAGAGTATTGGATTGCGGGAATTACCAATCTCGAGCTCATGCAATAATGTTATTGAAGTCGTTTTTCGAAGTGGCAGATCCGATCCAGTTGATAAATGTGGACGTTGAGTCATTCCGCCAGATAGTGCATGTGCTAAATGGTCGGATTTCACAAGCAGCGTCAAAGGCGGCATTGAAGCTTCTTGTGGAGCTTTGTCCGTGGGGAAGGAACCGAATCAAAGCTGTCCAAAGCGGTGCAGTTTCGGTGCTCACGGAGGCTCTCCTGGGGACGCAAGAGAAAAGAACGTGTGAGCTAATATTGATCATTTTAGAACAGCTTTGTGGATGTGCGGAAGGGCGAGCTGAGCTTTTGAAGCATGGGGCGGGGCTAGCCATTGTATCGAAGAAAATCCTTAGGGTTTCACATGTGGCAAGTGATCGGGCGGTAAGAATATTGTCCTCCATTTGTAGATTTTCTGCAACTTGTGGTGTTCTTCAGGATATGTTGCAAGTCGGTGCTGTGGCCAAATTGTGTTTGGTGCTTCAAGTGGATTGTAGGTTTAAAACAAAGGAGAAGGCTAGGGTGGTTCTTAAGTTGCACTCTAGGGTTTGGAAGAATTCTTCATGTATTCCTGACCATTTGTTGCCCTTTTATCCGGATCAGGATCCTCTCCTAAGCTTAAGAGGTTGA

Functional Annotation

Gene Ontology

Molecular Function
Biological Process
Cellular Component

GO:0001101 GO:0003674 GO:0003824 GO:0004842 GO:0005575 GO:0005622 GO:0005623 GO:0005737 GO:0005829 GO:0006464 GO:0006807 GO:0006950 GO:0008150 GO:0008152 GO:0009414 GO:0009415 GO:0009628 GO:0009987 GO:0010033 GO:0010035 GO:0010200 GO:0010243 GO:0016567 GO:0016740 GO:0019538 GO:0019787 GO:0032446 GO:0036211 GO:0042221 GO:0042493 GO:0043170 GO:0043412 GO:0044237 GO:0044238 GO:0044260 GO:0044267 GO:0044424 GO:0044444 GO:0044464 GO:0050896 GO:0051865 GO:0070647 GO:0071704 GO:0140096 GO:1901564 GO:1901698 GO:1901700

KEGG Pathways

Metabolic & Signaling

No pathways identified.

Pfam Domains

Protein Families

U-box

Protein Analysis

415

Amino Acids

46.04

Weight (kDa)

8.66

Isoelectric Point (pI)

41.24

Instability Index

Protein Domains (Pfam)

Domain Name	Pfam ID	Position	E-value	Description
U-box	PF04564	6 - 79	1.8e-17	U-box domain
ARM_PUB	PF25598	98 - 398	3.7e-76	Plant U-box protein ARM repeat region

Hydrophobicity Profile (Kyte-Doolittle)

AI Structure Prediction Report

Calculating structure properties...

Genomic Context

Gene Family Tree

Species Filter

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Align Labels (Cladogram)

Image

Tree File

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Publication-ready

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Orthologous Genes (Group: OG0016794)

Species	Orthologous Gene IDs
fragaria_vesca	FvH4_3g26340
malus_domestica	MD03G1156000.v1.1 MD11G1173600.v1.1
prunus_persica	Prupe.6G145300_v2.0.a1
rosa_chinensis	RchiOBHm_Chr5g0048581
rosa_laevigata	RLG00000034537
rosa_multiflora	Rmu_sc0007341.1_g000023
rosa_roxburghii	Rroxscaffold_1G00033250
rosa_rugosa	Rorug05G0239700
rosa_samantha	Rh5AG319300 Rh5BG329700 Rh5DG340900
rosa_wichuraiana	Rw0G021250 Rw5G029860

Restriction Enzyme Sites

1 / 10

Enzyme	Recognition Site	Cut Count	Positions (bp)
AccII	CGCG	1 cut(s)	278
AccIII	TCCGGA	1 cut(s)	1210
AclWI	GGATC	7 cut(s)	56, 357, 647, 652, 1214, 1221, 1227
AcoI	YGGCCR	1 cut(s)	1077
AcsI	RAATTY	2 cut(s)	8, 1171
AcuI	CTGAAG	1 cut(s)	1034
AcyI	GRCGYC	1 cut(s)	249
AfaI	GTAC	1 cut(s)	92
AflII	CTTAAG	2 cut(s)	1145, 1234
AflIII	ACRYGT	2 cut(s)	860, 979
AgeI	ACCGGT	1 cut(s)	80
AgsI	TTSAA	7 cut(s)	377, 485, 502, 631, 751, 928, 1100
AhdI	GACNNNNNGTC	1 cut(s)	682
AjnI	CCWGG	2 cut(s)	291, 837
Alw21I	GWGCWC	3 cut(s)	230, 613, 825
Alw26I	GTCTC	2 cut(s)	524, 573
Alw44I	GTGCAC	1 cut(s)	226
AlwI	GGATC	7 cut(s)	56, 357, 647, 652, 1214, 1221, 1227
Ama87I	CYCGRG	1 cut(s)	606
Aor13HI	TCCGGA	1 cut(s)	1210
AoxI	GGCC	2 cut(s)	429, 1077
ApaLI	GTGCAC	1 cut(s)	226
ApeKI	GCWGC	2 cut(s)	407, 731
ApoI	RAATTY	2 cut(s)	8, 1171
ArsI	GACNNNNNNTTYG	2 cut(s)	279, 311
AsiGI	ACCGGT	1 cut(s)	80
AspLEI	GCGC	1 cut(s)	280
AsuHPI	GGTGA	1 cut(s)	392
AsuII	TTCGAA	1 cut(s)	642
AsuNHI	GCTAGC	1 cut(s)	943
AvaI	CYCGRG	1 cut(s)	606
AxyI	CCTNAGG	1 cut(s)	968
BaeGI	GKGCMC	1 cut(s)	230
BalI	TGGCCA	1 cut(s)	1079
BamHI	GGATCC	1 cut(s)	1219
BanII	GRGCYC	1 cut(s)	613
Bbv12I	GWGCWC	3 cut(s)	230, 613, 825
BbvI	GCAGC	2 cut(s)	419, 743
BccI	CCATC	2 cut(s)	374, 443
BciT130I	CCWGG	2 cut(s)	293, 839
BclI	TGATCA	1 cut(s)	876
BcoDI	GTCTC	2 cut(s)	524, 573
BfaI	CTAG	3 cut(s)	944, 1134, 1158
BfmI	CTRYAG	1 cut(s)	465
BfrI	CTTAAG	2 cut(s)	1145, 1234
BglII	AGATCT	2 cut(s)	181, 526
BisI	GCNGC	5 cut(s)	208, 408, 477, 732, 744
BlpI	GCTNAGC	1 cut(s)	918
BlsI	GCNGC	5 cut(s)	209, 409, 478, 733, 745
Bme1390I	CCNGG	2 cut(s)	293, 839
BmeRI	GACNNNNNGTC	1 cut(s)	682
BmeT110I	CYCGRG	1 cut(s)	606
BmiI	GGNNCC	2 cut(s)	785, 1221
BmrFI	CCNGG	2 cut(s)	293, 839
BmsI	GCATC	1 cut(s)	291
BmtI	GCTAGC	1 cut(s)	947
Bpu10I	CCTNAGC	1 cut(s)	1229
Bpu1102I	GCTNAGC	1 cut(s)	918
Bpu14I	TTCGAA	1 cut(s)	642
BsaAI	YACGTR	1 cut(s)	427
BsaBI	GATNNNNATC	2 cut(s)	305, 1218
BsaHI	GRCGYC	1 cut(s)	249
BsaJI	CCNNGG	3 cut(s)	291, 773, 838
BsaWI	WCCGGW	3 cut(s)	64, 80, 1210
BsaXI	ACNNNNNCTCC	2 cut(s)	416, 446
Bse118I	RCCGGY	1 cut(s)	80
Bse1I	ACTGG	1 cut(s)	661
Bse21I	CCTNAGG	1 cut(s)	968
Bse3DI	GCAATG	2 cut(s)	343, 544
Bse8I	GATNNNNATC	2 cut(s)	305, 1218
BseAI	TCCGGA	1 cut(s)	1210
BseBI	CCWGG	2 cut(s)	293, 839
BseDI	CCNNGG	3 cut(s)	291, 773, 838
BseGI	GGATG	1 cut(s)	905
BseJI	GATNNNNATC	2 cut(s)	305, 1218
BseMI	GCAATG	2 cut(s)	343, 544
BseMII	CTCAG	2 cut(s)	373, 909
BseNI	ACTGG	1 cut(s)	661
BseSI	GKGCMC	1 cut(s)	230
BseXI	GCAGC	2 cut(s)	419, 743
BsgI	GTGCAG	1 cut(s)	831
Bsh1236I	CGCG	1 cut(s)	278
Bsh1285I	CGRYCG	1 cut(s)	97
BshFI	GGCC	2 cut(s)	431, 1079
BshTI	ACCGGT	1 cut(s)	80
BsiEI	CGRYCG	1 cut(s)	97
BsiHKAI	GWGCWC	3 cut(s)	230, 613, 825
BsiHKCI	CYCGRG	1 cut(s)	606
BsiSI	CCGG	3 cut(s)	65, 81, 1211
BsiWI	CGTACG	1 cut(s)	90
BslFI	GGGAC	1 cut(s)	856
BsmAI	GTCTC	2 cut(s)	524, 573
BsmBI	CGTCTC	1 cut(s)	524
BsmFI	GGGAC	1 cut(s)	856
BsnI	GGCC	2 cut(s)	431, 1079
BsoBI	CYCGRG	1 cut(s)	606
Bsp119I	TTCGAA	1 cut(s)	642
Bsp1286I	GDGCHC	3 cut(s)	230, 613, 825
Bsp13I	TCCGGA	1 cut(s)	1210
Bsp1720I	GCTNAGC	1 cut(s)	918
BspANI	GGCC	2 cut(s)	431, 1079
BspCNI	CTCAG	2 cut(s)	372, 910
BspEI	TCCGGA	1 cut(s)	1210
BspFNI	CGCG	1 cut(s)	278
BspHI	TCATGA	2 cut(s)	54, 531
BspLI	GGNNCC	2 cut(s)	785, 1221
BspOI	GCTAGC	1 cut(s)	947
BspPI	GGATC	7 cut(s)	56, 357, 647, 652, 1214, 1221, 1227
BspT104I	TTCGAA	1 cut(s)	642
BspTI	CTTAAG	2 cut(s)	1145, 1234
BsrDI	GCAATG	2 cut(s)	343, 544
BsrFI	RCCGGY	1 cut(s)	80
BsrI	ACTGG	1 cut(s)	661
BssAI	RCCGGY	1 cut(s)	80
BssECI	CCNNGG	3 cut(s)	291, 773, 838
BssNI	GRCGYC	1 cut(s)	249
Bst2UI	CCWGG	2 cut(s)	293, 839
Bst4CI	ACNGT	2 cut(s)	398, 469
Bst6I	CTCTTC	1 cut(s)	244
BstACI	GRCGYC	1 cut(s)	249
BstAFI	CTTAAG	2 cut(s)	1145, 1234
BstBAI	YACGTR	1 cut(s)	427
BstBI	TTCGAA	1 cut(s)	642
BstC8I	GCNNGC	2 cut(s)	915, 945
BstDEI	CTNAG	4 cut(s)	359, 918, 968, 1229
BstDSI	CCRYGG	1 cut(s)	773
BstF5I	GGATG	1 cut(s)	905
BstFNI	CGCG	1 cut(s)	278
BstHHI	GCGC	1 cut(s)	280
BstMAI	GTCTC	2 cut(s)	524, 573
BstMCI	CGRYCG	1 cut(s)	97
BstMWI	GCNNNNNNNGC	7 cut(s)	150, 413, 543, 740, 919, 928, 944
BstNI	CCWGG	2 cut(s)	293, 839
BstNSI	RCATGY	2 cut(s)	707, 983
BstSCI	CCNGG	2 cut(s)	291, 837
BstSFI	CTRYAG	1 cut(s)	465
BstSLI	GKGCMC	1 cut(s)	230
BstUI	CGCG	1 cut(s)	278
BstV1I	GCAGC	2 cut(s)	419, 743
BstX2I	RGATCY	6 cut(s)	61, 181, 362, 526, 652, 1219
BstYI	RGATCY	6 cut(s)	61, 181, 362, 526, 652, 1219
Bsu36I	CCTNAGG	1 cut(s)	968
BsuRI	GGCC	2 cut(s)	431, 1079
BtgI	CCRYGG	1 cut(s)	773
BtgZI	GCGATG	1 cut(s)	352
BtsCI	GGATG	1 cut(s)	905
BtsI	GCAGTG	1 cut(s)	119
BtsIMutI	CAGTG	1 cut(s)	119
Cac8I	GCNNGC	2 cut(s)	915, 945
CciI	TCATGA	2 cut(s)	54, 531
CfoI	GCGC	1 cut(s)	280
Cfr10I	RCCGGY	1 cut(s)	80
CseI	GACGC	4 cut(s)	185, 238, 723, 853
Csp6I	GTAC	1 cut(s)	91
CspAI	ACCGGT	1 cut(s)	80
CviAII	CATG	7 cut(s)	55, 532, 614, 704, 932, 980, 1179
CviQI	GTAC	1 cut(s)	91
DdeI	CTNAG	4 cut(s)	359, 918, 968, 1229
DraI	TTTAAA	1 cut(s)	1117
DriI	GACNNNNNGTC	1 cut(s)	682
EaeI	YGGCCR	1 cut(s)	1077
Eam1104I	CTCTTC	1 cut(s)	244
Eam1105I	GACNNNNNGTC	1 cut(s)	682
EarI	CTCTTC	1 cut(s)	244
EciI	GGCGGA	2 cut(s)	194, 680
Ecl136II	GAGCTC	1 cut(s)	611
Eco24I	GRGCYC	1 cut(s)	613
Eco53kI	GAGCTC	1 cut(s)	611
Eco57I	CTGAAG	1 cut(s)	1034
Eco81I	CCTNAGG	1 cut(s)	968
Eco88I	CYCGRG	1 cut(s)	606
EcoICRI	GAGCTC	1 cut(s)	611
EcoRI	GAATTC	2 cut(s)	8, 1171
EcoRII	CCWGG	2 cut(s)	291, 837
EcoT38I	GRGCYC	1 cut(s)	613
Esp3I	CGTCTC	1 cut(s)	524
FaeI	CATG	7 cut(s)	58, 535, 617, 707, 935, 983, 1182
FaqI	GGGAC	1 cut(s)	856
FatI	CATG	7 cut(s)	54, 531, 613, 703, 931, 979, 1178
FauI	CCCGC	2 cut(s)	584, 931
FbaI	TGATCA	1 cut(s)	876
Fnu4HI	GCNGC	5 cut(s)	208, 408, 477, 732, 744
FokI	GGATG	1 cut(s)	912
FriOI	GRGCYC	1 cut(s)	613
Fsp4HI	GCNGC	5 cut(s)	208, 408, 477, 732, 744
FspBI	CTAG	3 cut(s)	944, 1134, 1158
GlaI	GCGC	1 cut(s)	279
GluI	GCNGC	5 cut(s)	208, 408, 477, 732, 744
HaeIII	GGCC	2 cut(s)	431, 1079
HapII	CCGG	3 cut(s)	65, 81, 1211
HgaI	GACGC	4 cut(s)	185, 238, 723, 853
HhaI	GCGC	1 cut(s)	280
Hin1I	GRCGYC	1 cut(s)	249
Hin1II	CATG	7 cut(s)	58, 535, 617, 707, 935, 983, 1182
Hin6I	GCGC	1 cut(s)	278
HinP1I	GCGC	1 cut(s)	278
HincII	GTYRAC	1 cut(s)	175
HindII	GTYRAC	1 cut(s)	175
HindIII	AAGCTT	3 cut(s)	219, 752, 1231
HinfI	GANTC	5 cut(s)	458, 512, 566, 683, 789
HpaII	CCGG	3 cut(s)	65, 81, 1211
HphI	GGTGA	1 cut(s)	392
Hpy166II	GTNNAC	4 cut(s)	123, 175, 228, 676
Hpy188I	TCNGA	7 cut(s)	169, 362, 457, 511, 578, 657, 720
Hpy8I	GTNNAC	4 cut(s)	123, 175, 228, 676
Hpy99I	CGWCG	2 cut(s)	173, 254
HpyAV	CCTTC	3 cut(s)	774, 902, 1123
HpyCH4III	ACNGT	2 cut(s)	398, 469
HpyCH4IV	ACGT	5 cut(s)	89, 171, 426, 678, 860
HpyCH4V	TGCA	7 cut(s)	228, 617, 703, 812, 1034, 1063, 1153
HpyF10VI	GCNNNNNNNGC	7 cut(s)	150, 413, 543, 740, 919, 928, 944
HpyF3I	CTNAG	4 cut(s)	359, 918, 968, 1229
HpySE526I	ACGT	5 cut(s)	89, 171, 426, 678, 860
Hsp92I	GRCGYC	1 cut(s)	249
Hsp92II	CATG	7 cut(s)	58, 535, 617, 707, 935, 983, 1182
HspAI	GCGC	1 cut(s)	278
Kpn2I	TCCGGA	1 cut(s)	1210
Ksp22I	TGATCA	1 cut(s)	876
LmnI	GCTCC	2 cut(s)	413, 763
Lsp1109I	GCAGC	2 cut(s)	419, 743
LweI	GCATC	1 cut(s)	291
MaeI	CTAG	3 cut(s)	944, 1134, 1158
MaeII	ACGT	5 cut(s)	89, 171, 426, 678, 860
MaeIII	GTNAC	1 cut(s)	67
MboII	GAAGA	6 cut(s)	142, 231, 970, 1040, 1167, 1180
MflI	RGATCY	6 cut(s)	61, 181, 362, 526, 652, 1219
MhlI	GDGCHC	3 cut(s)	230, 613, 825
MlsI	TGGCCA	1 cut(s)	1079
MluCI	AATT	5 cut(s)	8, 39, 595, 1082, 1171
MluNI	TGGCCA	1 cut(s)	1079
MlyI	GAGTC	2 cut(s)	575, 692
MmeI	TCCRAC	2 cut(s)	192, 698
Mox20I	TGGCCA	1 cut(s)	1079
MroI	TCCGGA	1 cut(s)	1210
MscI	TGGCCA	1 cut(s)	1079
MseI	TTAA	3 cut(s)	1116, 1146, 1235
Msp20I	TGGCCA	1 cut(s)	1079
MspCI	CTTAAG	2 cut(s)	1145, 1234
MspI	CCGG	3 cut(s)	65, 81, 1211
MspR9I	CCNGG	2 cut(s)	293, 839
MvaI	CCWGG	2 cut(s)	293, 839
MvnI	CGCG	1 cut(s)	278
MwoI	GCNNNNNNNGC	7 cut(s)	150, 413, 543, 740, 919, 928, 944
NheI	GCTAGC	1 cut(s)	943
NlaIII	CATG	7 cut(s)	58, 535, 617, 707, 935, 983, 1182
NlaIV	GGNNCC	2 cut(s)	785, 1221
NmuCI	GTSAC	1 cut(s)	67
NspI	RCATGY	2 cut(s)	707, 983
NspV	TTCGAA	1 cut(s)	642
PaeR7I	CTCGAG	1 cut(s)	606
PagI	TCATGA	2 cut(s)	54, 531
PciI	ACATGT	1 cut(s)	979
PfeI	GAWTC	3 cut(s)	458, 512, 789
Pfl23II	CGTACG	1 cut(s)	90
PinAI	ACCGGT	1 cut(s)	80
PkrI	GCNGC	5 cut(s)	209, 409, 478, 733, 745
Ple19I	CGATCG	1 cut(s)	97
PleI	GAGTC	2 cut(s)	574, 691
PpsI	GAGTC	2 cut(s)	574, 691
Ppu21I	YACGTR	1 cut(s)	427
PscI	ACATGT	1 cut(s)	979
Psp124BI	GAGCTC	1 cut(s)	613
Psp6I	CCWGG	2 cut(s)	291, 837
PspGI	CCWGG	2 cut(s)	291, 837
PspLI	CGTACG	1 cut(s)	90
PspN4I	GGNNCC	2 cut(s)	785, 1221
PsuI	RGATCY	6 cut(s)	61, 181, 362, 526, 652, 1219
PvuI	CGATCG	1 cut(s)	97
RsaI	GTAC	1 cut(s)	92
RsaNI	GTAC	1 cut(s)	91
SacI	GAGCTC	1 cut(s)	613
SaqAI	TTAA	3 cut(s)	1116, 1146, 1235
SatI	GCNGC	5 cut(s)	208, 408, 477, 732, 744
SchI	GAGTC	2 cut(s)	575, 692
ScrFI	CCNGG	2 cut(s)	293, 839
SduI	GDGCHC	3 cut(s)	230, 613, 825
SfaNI	GCATC	1 cut(s)	291
SfcI	CTRYAG	1 cut(s)	465
Sfr274I	CTCGAG	1 cut(s)	606
SfuI	TTCGAA	1 cut(s)	642
SlaI	CTCGAG	1 cut(s)	606
SmlI	CTYRAG	3 cut(s)	606, 1145, 1234
SmoI	CTYRAG	3 cut(s)	606, 1145, 1234
Sse9I	AATT	5 cut(s)	8, 39, 595, 1082, 1171
SspI	AATATT	2 cut(s)	873, 1008
SspMI	CTAG	3 cut(s)	944, 1134, 1158
SstI	GAGCTC	1 cut(s)	613
StyD4I	CCNGG	2 cut(s)	291, 837
TaaI	ACNGT	2 cut(s)	398, 469
TaiI	ACGT	5 cut(s)	92, 174, 429, 681, 863
TasI	AATT	5 cut(s)	8, 39, 595, 1082, 1171
TauI	GCSGC	3 cut(s)	210, 479, 746
TfiI	GAWTC	3 cut(s)	458, 512, 789
Tru1I	TTAA	3 cut(s)	1116, 1146, 1235
Tru9I	TTAA	3 cut(s)	1116, 1146, 1235
TscAI	CASTG	1 cut(s)	119
TseFI	GTSAC	1 cut(s)	67
TseI	GCWGC	2 cut(s)	407, 731
Tsp45I	GTSAC	1 cut(s)	67
TspDTI	ATGAA	4 cut(s)	33, 450, 485, 1167
TspGWI	ACGGA	3 cut(s)	578, 762, 842
TspRI	CASTG	1 cut(s)	119
Vha464I	CTTAAG	2 cut(s)	1145, 1234
VneI	GTGCAC	1 cut(s)	226
XapI	RAATTY	2 cut(s)	8, 1171
XceI	RCATGY	2 cut(s)	707, 983
XcmI	CCANNNNNNNNNTGG	1 cut(s)	1087
XhoI	CTCGAG	1 cut(s)	606
XspI	CTAG	3 cut(s)	944, 1134, 1158

Using CommOnly database (standard laboratory enzymes). Scanned on CDS sequence.