Basic Information

Type: gene

Biological Identity

Species

rosa_rugosa

Chromosome

GWHBQTZ00000006

Orthogroup ID

OG0010975

Physical Location & Seq

Genomic Coordinates Reverse (-)

22117778 .. 22119091

Gene Structure 1314 bp

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UTR

Exon/CDS

Intron

Transcript / Protein ID

Rorug06G0154100.1

Sequence Viewer

CDS Protein Promoter (2k)

Length: 1314 bp

ATGGCTAGTATTGATTCTGTCACCAAAAGCTTTGCTACCTCGCTAGCTATCGTTACCAATGAGGTTGTTGATATCTCCTCCATGGAGAATGGGGGGGCGCAGCGGAATGCACAGTCCTACCTCTTGGCAAGGCCTCTGACTTCCAAGCCAACAAATGCTTCTGATCTGCGGAGGCATTTTCGTCGAATATGGGTGCTAAACAAGGGTTTCAGTGTTCAGCCGAAAGCTCCGGATAGATTCCTCTTCTCCTTTGATCTCCGGGGAGATAGAAATAAAGTTTTGATGGGTGGTCCGTGGAGTTTCAATCGTGCTTTGGTTGTCATGCAGCCATATGATGGAGTAGCACCTCTGCAATCTGTGGAGATGGAGGACCTTTTCTTTTGGGTTCGCATCACTGACATGCCGCCCGCATATGAGCACAGGAACACTATTGCCAATGTTGCCTCCATTGCGGGTACATTTCTTCATCTGGATCAAAAGTTATTTGATACAACTCGGGAAATTAGGGTTAGGGTTTCTCATGCAATTTCGAAACCCTTCTTCCTGCACAAAACCCTAAAATTTGGCAAAGGGGTAGAACAAGAGGTTCACTTCTTTTTTGAGAATCTTGTGGGCAAATGCACTATATGCCTTTGTGTGTATCATAAGGGGGCAGCGTGTCAGGCGGCAACTACTGCGCCGGTGGTCCAGGAAGCTCCCACACCGGCGGCACGAAAATTAAATCTCACTCGCCCCTTCTTAGGTTTTCACGAAAATCAGTTCAAGGATGGATTGTTTCGCTTCCATGGAATTTCCACCCCCATTCTCCCACCTGTGGGAAGGTCTCTTTTTGGAGGTGCTGGGACTAGTAAGGTACGCAAACCCATTGTTCTTCGTCATGCAGGAAAGACACATACTAATGACAACAATGAATTGGCTTTGGTACCTGTGGATACCGATCAGCCTACTAAGAGAAATCGGCCCTTTCCATCTCCAAAAAAATTGCAGTTTATGATGTCTGATTTGCTAGAGGGTAAAACAAGCCCTATTGCTCCCTCAAAGAAGGTAAAACTACCTGAATTCCCTACAAAATTCAATGCAAAATCTCTGGGTTTGATGGAAACACCAGGAGGTATGCTTGTGCCTGCAGAGGGTATGATGCCCAAACCTTTACAACTTGAGCTTCCTAGTGGCAAAAAGAAGAGAGGTCGGCCTTTGGGAGCAAAGAACAGAAATCCACCAAAAGCAAAAAAGATTCCTGCTGAAGATGTCCTGAGTTTTCTGTATTCAGGCAAGCCTCCAAGCCCTAGCTTCAAGGGCAAGGAGAAGATATAG

Functional Annotation

Gene Ontology

Molecular Function
Biological Process
Cellular Component

GO:0003674 GO:0003824 GO:0004672 GO:0004674 GO:0005488 GO:0005515 GO:0005575 GO:0005622 GO:0005623 GO:0005634 GO:0005737 GO:0005886 GO:0006464 GO:0006468 GO:0006793 GO:0006796 GO:0006807 GO:0006950 GO:0006979 GO:0007154 GO:0007165 GO:0008150 GO:0008152 GO:0009611 GO:0009987 GO:0016020 GO:0016301 GO:0016310 GO:0016740 GO:0016772 GO:0016773 GO:0019538 GO:0019899 GO:0019900 GO:0019901 GO:0023052 GO:0035556 GO:0036211 GO:0043170 GO:0043226 GO:0043227 GO:0043229 GO:0043231 GO:0043412 GO:0044237 GO:0044238 GO:0044260 GO:0044267 GO:0044424 GO:0044464 GO:0050789 GO:0050794 GO:0050896 GO:0051716 GO:0065007 GO:0071704 GO:0071944 GO:0140096 GO:1901564

KEGG Pathways

Metabolic & Signaling

ko04016 map04016

Pfam Domains

Protein Families

Pkinase

Protein Analysis

437

Amino Acids

48.48

Weight (kDa)

10.06

Isoelectric Point (pI)

42.32

Instability Index

Protein Domains (Pfam)

Domain Name	Pfam ID	Position	E-value	Description
DUF4283	PF14111	37 - 112	1.1e-08	Domain of unknown function (DUF4283)

Hydrophobicity Profile (Kyte-Doolittle)

AI Structure Prediction Report

Calculating structure properties...

Genomic Context

Gene Family Tree

Species Filter

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Align Labels (Cladogram)

Image

Tree File

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Publication-ready

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Orthologous Genes (Group: OG0010975)

Species	Orthologous Gene IDs
arabidopsis_thaliana	AT3G25250 AT4G13000
fragaria_vesca	FvH4_2g18040
malus_domestica	MD05G1164900.v1.1 MD10G1154200.v1.1
prunus_persica	Prupe.8G193100_v2.0.a1
pyrus_communis	pycom05g15350 pycom10g13330
rosa_chinensis	RchiOBHm_Chr6g0283431
rosa_laevigata	RLG00000012827
rosa_multiflora	Rmu_sc0003166.1_g000017 Rmu_sc0004959.1_g000020 Rmu_sc0039373.1_g000001
rosa_roxburghii	Rroxscaffold_7G00185540
rosa_rugosa	Rorug06G0154100
rosa_samantha	Rh6AG266400 Rh6BG268900 Rh6CG268800 Rh6DG262200
rosa_wichuraiana	Rw6G023060

Restriction Enzyme Sites

1 / 10

Enzyme	Recognition Site	Cut Count	Positions (bp)
Acc65I	GGTACC	1 cut(s)	922
AccB1I	GGYRCC	1 cut(s)	922
AccB7I	CCANNNNNTGG	1 cut(s)	335
AccIII	TCCGGA	1 cut(s)	229
AciI	CCGC	7 cut(s)	103, 169, 404, 408, 452, 665, 707
AclWI	GGATC	1 cut(s)	480
AcsI	RAATTY	4 cut(s)	560, 789, 1058, 1070
AcuI	CTGAAG	1 cut(s)	1263
AfaI	GTAC	3 cut(s)	457, 855, 924
AfiI	CCNNNNNNNGG	5 cut(s)	335, 740, 814, 815, 1130
AgsI	TTSAA	4 cut(s)	304, 763, 1075, 1294
AhlI	ACTAGT	1 cut(s)	845
AjnI	CCWGG	2 cut(s)	687, 1105
AjuI	GAANNNNNNNTTGG	2 cut(s)	142, 174
AluBI	AGCT	6 cut(s)	30, 47, 227, 695, 1162, 1290
AluI	AGCT	6 cut(s)	30, 47, 227, 695, 1162, 1290
Alw21I	GWGCWC	1 cut(s)	420
Alw26I	GTCTC	1 cut(s)	828
AlwI	GGATC	1 cut(s)	480
Ama87I	CYCGRG	1 cut(s)	495
Aor13HI	TCCGGA	1 cut(s)	229
AoxI	GGCC	3 cut(s)	131, 959, 1190
ApeKI	GCWGC	3 cut(s)	100, 325, 653
ApoI	RAATTY	4 cut(s)	560, 789, 1058, 1070
Asp700I	GAANNNNTTC	1 cut(s)	536
Asp718I	GGTACC	1 cut(s)	922
AspLEI	GCGC	2 cut(s)	100, 679
AspS9I	GGNCC	4 cut(s)	290, 370, 685, 960
AsuC2I	CCSGG	1 cut(s)	260
AsuHPI	GGTGA	1 cut(s)	13
AsuII	TTCGAA	1 cut(s)	530
AsuNHI	GCTAGC	1 cut(s)	43
AvaI	CYCGRG	1 cut(s)	495
AvaII	GGWCC	3 cut(s)	290, 370, 685
BanI	GGYRCC	1 cut(s)	922
Bbv12I	GWGCWC	1 cut(s)	420
BbvI	GCAGC	3 cut(s)	112, 337, 665
BccI	CCATC	6 cut(s)	277, 329, 358, 761, 976, 1090
BcgI	CGANNNNNNTGC	2 cut(s)	164, 198
BciT130I	CCWGG	2 cut(s)	689, 1107
BciVI	GTATCC	1 cut(s)	925
BcnI	CCSGG	1 cut(s)	260
BcoDI	GTCTC	1 cut(s)	828
BcuI	ACTAGT	1 cut(s)	845
BfaI	CTAG	6 cut(s)	6, 44, 846, 1007, 1167, 1287
BfmI	CTRYAG	1 cut(s)	1125
BfuI	GTATCC	1 cut(s)	925
BisI	GCNGC	6 cut(s)	101, 326, 404, 654, 666, 708
BlsI	GCNGC	6 cut(s)	102, 327, 405, 655, 667, 709
Bme1390I	CCNGG	3 cut(s)	260, 689, 1107
Bme18I	GGWCC	3 cut(s)	290, 370, 685
BmeT110I	CYCGRG	1 cut(s)	495
BmgT120I	GGNCC	4 cut(s)	290, 370, 685, 960
BmiI	GGNNCC	1 cut(s)	924
BmrFI	CCNGG	3 cut(s)	260, 689, 1107
BmsI	GCATC	2 cut(s)	399, 1128
BmtI	GCTAGC	1 cut(s)	47
Bpu14I	TTCGAA	1 cut(s)	530
BpuEI	CTTGAG	1 cut(s)	1178
BpuMI	CCSGG	1 cut(s)	260
BsaBI	GATNNNNATC	1 cut(s)	936
BsaI	GGTCTC	1 cut(s)	828
BsaJI	CCNNGG	4 cut(s)	81, 259, 293, 784
BsaWI	WCCGGW	1 cut(s)	229
Bsc4I	CCNNNNNNNGG	5 cut(s)	335, 740, 814, 815, 1130
Bse118I	RCCGGY	2 cut(s)	679, 703
Bse3DI	GCAATG	1 cut(s)	447
Bse8I	GATNNNNATC	1 cut(s)	936
BseAI	TCCGGA	1 cut(s)	229
BseBI	CCWGG	2 cut(s)	689, 1107
BseDI	CCNNGG	4 cut(s)	81, 259, 293, 784
BseGI	GGATG	1 cut(s)	772
BseJI	GATNNNNATC	1 cut(s)	936
BseLI	CCNNNNNNNGG	5 cut(s)	335, 740, 814, 815, 1130
BseMI	GCAATG	1 cut(s)	447
BseMII	CTCAG	1 cut(s)	1244
BseRI	GAGGAG	1 cut(s)	67
BseXI	GCAGC	3 cut(s)	112, 337, 665
BseYI	CCCAGC	1 cut(s)	839
BsgI	GTGCAG	1 cut(s)	530
BshFI	GGCC	3 cut(s)	133, 961, 1192
BshNI	GGYRCC	1 cut(s)	922
BsiHKAI	GWGCWC	1 cut(s)	420
BsiHKCI	CYCGRG	1 cut(s)	495
BsiSI	CCGG	4 cut(s)	230, 259, 680, 704
BslFI	GGGAC	1 cut(s)	856
BslI	CCNNNNNNNGG	5 cut(s)	335, 740, 814, 815, 1130
BsmAI	GTCTC	1 cut(s)	828
BsmFI	GGGAC	1 cut(s)	856
BsmI	GAATGC	1 cut(s)	112
BsnI	GGCC	3 cut(s)	133, 961, 1192
Bso31I	GGTCTC	1 cut(s)	828
BsoBI	CYCGRG	1 cut(s)	495
Bsp119I	TTCGAA	1 cut(s)	530
Bsp1286I	GDGCHC	1 cut(s)	420
Bsp13I	TCCGGA	1 cut(s)	229
Bsp143I	GATC	4 cut(s)	163, 253, 472, 937
Bsp19I	CCATGG	2 cut(s)	81, 784
BspACI	CCGC	7 cut(s)	103, 169, 404, 408, 452, 665, 707
BspANI	GGCC	3 cut(s)	133, 961, 1192
BspCNI	CTCAG	1 cut(s)	1245
BspEI	TCCGGA	1 cut(s)	229
BspLI	GGNNCC	1 cut(s)	924
BspMAI	CTGCAG	1 cut(s)	1129
BspOI	GCTAGC	1 cut(s)	47
BspPI	GGATC	1 cut(s)	480
BspT104I	TTCGAA	1 cut(s)	530
BspT107I	GGYRCC	1 cut(s)	922
BspTNI	GGTCTC	1 cut(s)	828
BsrDI	GCAATG	1 cut(s)	447
BsrFI	RCCGGY	2 cut(s)	679, 703
BssAI	RCCGGY	2 cut(s)	679, 703
BssECI	CCNNGG	4 cut(s)	81, 259, 293, 784
BssMI	GATC	4 cut(s)	163, 253, 472, 937
BssT1I	CCWWGG	2 cut(s)	81, 784
Bst2UI	CCWGG	2 cut(s)	689, 1107
Bst4CI	ACNGT	1 cut(s)	114
Bst6I	CTCTTC	2 cut(s)	248, 1175
BstAPI	GCANNNNNTGC	2 cut(s)	627, 674
BstBI	TTCGAA	1 cut(s)	530
BstC8I	GCNNGC	4 cut(s)	45, 408, 1125, 1274
BstDEI	CTNAG	3 cut(s)	739, 948, 1253
BstDSI	CCRYGG	3 cut(s)	81, 293, 784
BstF5I	GGATG	1 cut(s)	772
BstHHI	GCGC	2 cut(s)	100, 679
BstKTI	GATC	4 cut(s)	166, 256, 475, 940
BstMAI	GTCTC	1 cut(s)	828
BstMBI	GATC	4 cut(s)	163, 253, 472, 937
BstMWI	GCNNNNNNNGC	6 cut(s)	440, 449, 627, 662, 674, 1296
BstNI	CCWGG	2 cut(s)	689, 1107
BstNSI	RCATGY	1 cut(s)	403
BstSCI	CCNGG	3 cut(s)	258, 687, 1105
BstSFI	CTRYAG	1 cut(s)	1125
BstV1I	GCAGC	3 cut(s)	112, 337, 665
BsuI	GTATCC	1 cut(s)	925
BsuRI	GGCC	3 cut(s)	133, 961, 1192
BtgI	CCRYGG	3 cut(s)	81, 293, 784
BtsCI	GGATG	1 cut(s)	772
BtsIMutI	CAGTG	2 cut(s)	217, 393
Cac8I	GCNNGC	4 cut(s)	45, 408, 1125, 1274
CfoI	GCGC	2 cut(s)	100, 679
Cfr10I	RCCGGY	2 cut(s)	679, 703
Cfr13I	GGNCC	4 cut(s)	290, 370, 685, 960
Csp6I	GTAC	3 cut(s)	456, 854, 923
CviAII	CATG	6 cut(s)	82, 322, 400, 521, 785, 878
CviQI	GTAC	3 cut(s)	456, 854, 923
DdeI	CTNAG	3 cut(s)	739, 948, 1253
DpnI	GATC	4 cut(s)	165, 255, 474, 939
DpnII	GATC	4 cut(s)	163, 253, 472, 937
Eam1104I	CTCTTC	2 cut(s)	248, 1175
EarI	CTCTTC	2 cut(s)	248, 1175
Eco130I	CCWWGG	2 cut(s)	81, 784
Eco147I	AGGCCT	1 cut(s)	133
Eco31I	GGTCTC	1 cut(s)	828
Eco32I	GATATC	1 cut(s)	73
Eco47I	GGWCC	3 cut(s)	290, 370, 685
Eco57I	CTGAAG	1 cut(s)	1263
Eco88I	CYCGRG	1 cut(s)	495
EcoO109I	RGGNCCY	1 cut(s)	370
EcoRI	GAATTC	1 cut(s)	1058
EcoRII	CCWGG	2 cut(s)	687, 1105
EcoRV	GATATC	1 cut(s)	73
EcoT14I	CCWWGG	2 cut(s)	81, 784
ErhI	CCWWGG	2 cut(s)	81, 784
FaeI	CATG	6 cut(s)	85, 325, 403, 524, 788, 881
FaqI	GGGAC	1 cut(s)	856
FatI	CATG	6 cut(s)	81, 321, 399, 520, 784, 877
FauI	CCCGC	2 cut(s)	415, 445
FauNDI	CATATG	2 cut(s)	331, 412
Fnu4HI	GCNGC	6 cut(s)	101, 326, 404, 654, 666, 708
FokI	GGATG	1 cut(s)	779
Fsp4HI	GCNGC	6 cut(s)	101, 326, 404, 654, 666, 708
FspBI	CTAG	6 cut(s)	6, 44, 846, 1007, 1167, 1287
GlaI	GCGC	2 cut(s)	99, 678
GluI	GCNGC	6 cut(s)	101, 326, 404, 654, 666, 708
GsaI	CCCAGC	1 cut(s)	843
HaeIII	GGCC	3 cut(s)	133, 961, 1192
HapII	CCGG	4 cut(s)	230, 259, 680, 704
HhaI	GCGC	2 cut(s)	100, 679
Hin1II	CATG	6 cut(s)	85, 325, 403, 524, 788, 881
Hin6I	GCGC	2 cut(s)	98, 677
HinP1I	GCGC	2 cut(s)	98, 677
HindIII	AAGCTT	1 cut(s)	28
HinfI	GANTC	4 cut(s)	14, 237, 604, 1234
HpaII	CCGG	4 cut(s)	230, 259, 680, 704
HphI	GGTGA	1 cut(s)	13
Hpy166II	GTNNAC	1 cut(s)	589
Hpy188I	TCNGA	3 cut(s)	138, 163, 1000
Hpy188III	TCNNGA	5 cut(s)	230, 470, 497, 749, 1252
Hpy8I	GTNNAC	1 cut(s)	589
Hpy99I	CGWCG	1 cut(s)	186
HpyAV	CCTTC	4 cut(s)	547, 745, 813, 1036
HpyCH4III	ACNGT	1 cut(s)	114
HpyF10VI	GCNNNNNNNGC	6 cut(s)	440, 449, 627, 662, 674, 1296
HpyF3I	CTNAG	3 cut(s)	739, 948, 1253
Hsp92II	CATG	6 cut(s)	85, 325, 403, 524, 788, 881
HspAI	GCGC	2 cut(s)	98, 677
Kpn2I	TCCGGA	1 cut(s)	229
KpnI	GGTACC	1 cut(s)	926
Kzo9I	GATC	4 cut(s)	163, 253, 472, 937
LmnI	GCTCC	4 cut(s)	232, 700, 1036, 1199
Lsp1109I	GCAGC	3 cut(s)	112, 337, 665
LweI	GCATC	2 cut(s)	399, 1128
MaeI	CTAG	6 cut(s)	6, 44, 846, 1007, 1167, 1287
MaeIII	GTNAC	2 cut(s)	19, 52
MalI	GATC	4 cut(s)	165, 255, 474, 939
MboI	GATC	4 cut(s)	163, 253, 472, 937
MboII	GAAGA	6 cut(s)	235, 455, 532, 863, 1192, 1256
MhlI	GDGCHC	1 cut(s)	420
MluCI	AATT	9 cut(s)	501, 525, 560, 716, 789, 911, 980, 1058, 1070
MroI	TCCGGA	1 cut(s)	229
MroXI	GAANNNNTTC	1 cut(s)	536
MseI	TTAA	1 cut(s)	719
MslI	CAYNNNNRTG	2 cut(s)	398, 897
MspA1I	CMGCKG	1 cut(s)	103
MspI	CCGG	4 cut(s)	230, 259, 680, 704
MspR9I	CCNGG	3 cut(s)	260, 689, 1107
Mva1269I	GAATGC	1 cut(s)	112
MvaI	CCWGG	2 cut(s)	689, 1107
MwoI	GCNNNNNNNGC	6 cut(s)	440, 449, 627, 662, 674, 1296
NciI	CCSGG	1 cut(s)	260
NcoI	CCATGG	2 cut(s)	81, 784
NdeI	CATATG	2 cut(s)	331, 412
NdeII	GATC	4 cut(s)	163, 253, 472, 937
NheI	GCTAGC	1 cut(s)	43
NlaIII	CATG	6 cut(s)	85, 325, 403, 524, 788, 881
NlaIV	GGNNCC	1 cut(s)	924
NmuCI	GTSAC	1 cut(s)	19
NspI	RCATGY	1 cut(s)	403
NspV	TTCGAA	1 cut(s)	530
PceI	AGGCCT	1 cut(s)	133
PctI	GAATGC	1 cut(s)	112
PdmI	GAANNNNTTC	1 cut(s)	536
PfeI	GAWTC	4 cut(s)	14, 237, 604, 1234
PflMI	CCANNNNNTGG	1 cut(s)	335
PfoI	TCCNGGA	1 cut(s)	687
PkrI	GCNGC	6 cut(s)	102, 327, 405, 655, 667, 709
PpuMI	RGGWCCY	1 cut(s)	370
Psp5II	RGGWCCY	1 cut(s)	370
Psp6I	CCWGG	2 cut(s)	687, 1105
PspFI	CCCAGC	1 cut(s)	839
PspGI	CCWGG	2 cut(s)	687, 1105
PspN4I	GGNNCC	1 cut(s)	924
PspPI	GGNCC	4 cut(s)	290, 370, 685, 960
PspPPI	RGGWCCY	1 cut(s)	370
PstI	CTGCAG	1 cut(s)	1129
RsaI	GTAC	3 cut(s)	457, 855, 924
RsaNI	GTAC	3 cut(s)	456, 854, 923
RseI	CAYNNNNRTG	2 cut(s)	398, 897
SaqAI	TTAA	1 cut(s)	719
SatI	GCNGC	6 cut(s)	101, 326, 404, 654, 666, 708
Sau3AI	GATC	4 cut(s)	163, 253, 472, 937
Sau96I	GGNCC	4 cut(s)	290, 370, 685, 960
ScrFI	CCNGG	3 cut(s)	260, 689, 1107
SduI	GDGCHC	1 cut(s)	420
SfaNI	GCATC	2 cut(s)	399, 1128
SfcI	CTRYAG	1 cut(s)	1125
SfuI	TTCGAA	1 cut(s)	530
SgrAI	CRCCGGYG	2 cut(s)	679, 703
SinI	GGWCC	3 cut(s)	290, 370, 685
SmiMI	CAYNNNNRTG	2 cut(s)	398, 897
SmlI	CTYRAG	1 cut(s)	1157
SmoI	CTYRAG	1 cut(s)	1157
SpeI	ACTAGT	1 cut(s)	845
Sse9I	AATT	9 cut(s)	501, 525, 560, 716, 789, 911, 980, 1058, 1070
SseBI	AGGCCT	1 cut(s)	133
SsiI	CCGC	7 cut(s)	103, 169, 404, 408, 452, 665, 707
SspMI	CTAG	6 cut(s)	6, 44, 846, 1007, 1167, 1287
StuI	AGGCCT	1 cut(s)	133
StyD4I	CCNGG	3 cut(s)	258, 687, 1105
StyI	CCWWGG	2 cut(s)	81, 784
TaaI	ACNGT	1 cut(s)	114
TaqI	TCGA	2 cut(s)	184, 530
TasI	AATT	9 cut(s)	501, 525, 560, 716, 789, 911, 980, 1058, 1070
TauI	GCSGC	3 cut(s)	406, 668, 710
TfiI	GAWTC	4 cut(s)	14, 237, 604, 1234
Tru1I	TTAA	1 cut(s)	719
Tru9I	TTAA	1 cut(s)	719
TscAI	CASTG	2 cut(s)	217, 400
TseFI	GTSAC	1 cut(s)	19
TseI	GCWGC	3 cut(s)	100, 325, 653
Tsp45I	GTSAC	1 cut(s)	19
TspDTI	ATGAA	2 cut(s)	455, 924
TspGWI	ACGGA	1 cut(s)	282
TspRI	CASTG	2 cut(s)	217, 400
Van91I	CCANNNNNTGG	1 cut(s)	335
VpaK11BI	GGWCC	3 cut(s)	290, 370, 685
XapI	RAATTY	4 cut(s)	560, 789, 1058, 1070
XceI	RCATGY	1 cut(s)	403
XmnI	GAANNNNTTC	1 cut(s)	536
XspI	CTAG	6 cut(s)	6, 44, 846, 1007, 1167, 1287

Using CommOnly database (standard laboratory enzymes). Scanned on CDS sequence.