Rh4AG025300

PDDEXK-like family of unknown function

Design Primers CRISPR Design

Basic Information

Type: gene
Biological Identity
rosa_samantha
Chr4A
OG0013897
Physical Location & Seq
Forward (+)
4893276 .. 4894688
1413 bp
Loading structure...
UTR
Exon/CDS
Intron
Rh4AG025300.1

Sequence Viewer

Length: 1185 bp
ATGCCTTTTCCGATGAAGATCCAACCGATCGATATCGATTCCGAGGCGGTGAGAGAGCCAGTCCGGATCGACTCGGCTGCCAAGCCGGTGTTGAAATCCCGGCTCCGGCGGCTGTTTGACCGGCAGTTCCCGAGCGTGCTTAGAATCTCCTCGACGGAAAAACCTACCGTCGCCGGCGGCGATTTCAACAAGGACGGCGGAGGAGGAGGAGGAGCCGTCGCCGAGTTCGAGCCGAGCTCGGTTTGCCTTGCGAAGATGGTTCAGAATTTCCTCGAAGAAACCAACGAGAAGCAGCAGGCGACGAAATGTGGCCGGAATCGCTGCAATTGCTTTAACGGAAACAACAATGACAGCTCCGACGACGAGCTCGACGTCTTTGGCGGCGAGTCCTTCACCACCGGCGGTTCCTTTGGCGGCGACGCCTCTGATGTCCTCAAGAGTCTGATTCCATGCGCGAGTGTCCACGAGAGAAACCTCTTAGCCGACGCAGCCAAAATCGTCGAGACGAACAAGAATATCAAACAGAAAGACGAGCTCAGAAAGATCGTCACCGACGGCCTCTCGTCTCTCGGCTACGACTCCTCCATCTGCAAATCCAAGTGGGACAAATCACCGTCTTTTCCCGCCGGTGAATACGAGTACATCGATGTGATGGTGGAGGGGGACAGGTTGTTGATTGACATCGATTTCCGATCGGAATTCGAAGTGGCGCGGCCCACTGGGGCCTACAAGGCAGTCCTTCAATCGCTGCCGTTCATTTTCGTCGGCAAATCGGACCGTCTCGGCCAGATTGTCTCCATCGTAACGGAGGCGGCGAAGCAGAGCCTCAAGAAGAAAGGAATGCCGTTTCCGCCGTGGCGGAAAGCCGAGTACGTCAGAGCCAAGTGGCTCTCTCCTTGCACAACCAGAACCACCACCTCGCAGCTCCGGAACGACGTCGTTGCTCCGGTGCCGGAGAATGGGACCGTCGTCGTTTCCGACGACCTGAGCGAGTGCGGAGAACTCGAGTTGATCTTCGGCGAGGACGCCGTGAATTTAACCGATCCGTCGCCGGAAGTTGAGGCGAAAGTGACCGGAAATTCAGTGGCGGTGGCGTGGCAGCCGCCGGCGGTCAAGCCGAAGAGCGTTGAGAGAGGAGCGAGGATCGTGACCGGTTTGGCTTCGTTGCTAGCAGCAAAACCGTGA
Functional Annotation
Gene Ontology
Molecular Function
Biological Process
Cellular Component
No ontology terms assigned.
KEGG Pathways
Metabolic & Signaling
No pathways identified.
Pfam Domains
Protein Families
PDDEXK_6

Protein Analysis

394

Amino Acids

42.75

Weight (kDa)

5.37

Isoelectric Point (pI)

46.72

Instability Index
Protein Domains (Pfam)
Domain Name Pfam ID Position E-value Description
PDDEXK_6 PF04720 85 - 301 4.9e-75 PDDEXK-like family of unknown function
Hydrophobicity Profile (Kyte-Doolittle)
AI Structure Prediction Report
Calculating structure properties...

Genomic Context

Gene Family Tree

Species Filter
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Publication-ready
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Orthologous Genes (Group: OG0013897)

Species Orthologous Gene IDs
arabidopsis_thaliana AT3G22970 AT3G22970
fragaria_vesca FvH4_4g01830
malus_domestica MD13G1202800.v1.1
prunus_persica Prupe.1G024900_v2.0.a1
pyrus_communis pycom13g17580 pycom16g17110
rosa_chinensis RchiOBHm_Chr4g0388891
rosa_laevigata RLG00000010016
rosa_multiflora Rmu_sc0000697.1_g000004
rosa_roxburghii Rroxscaffold_5G00335200
rosa_rugosa Rorug03G0320800
rosa_samantha Rh4AG025300 Rh4BG018500 Rh4CG028300 Rh4DG019000
rosa_wichuraiana Rw4G001820

Restriction Enzyme Sites

1 / 10
Enzyme Recognition Site Cut Count Positions (bp)
AatII GACGTC 2 cut(s) 375, 939
AccB1I GGYRCC 1 cut(s) 949
AccII CGCG 2 cut(s) 455, 712
AccIII TCCGGA 2 cut(s) 63, 927
AclWI GGATC 4 cut(s) 13, 74, 1037, 1151
AcoI YGGCCR 2 cut(s) 310, 784
AcsI RAATTY 4 cut(s) 265, 698, 1033, 1078
AcyI GRCGYC 4 cut(s) 372, 420, 936, 1026
AfaI GTAC 2 cut(s) 641, 872
AfiI CCNNNNNNNGG 2 cut(s) 105, 959
AgeI ACCGGT 1 cut(s) 1151
AgsI TTSAA 3 cut(s) 94, 187, 743
AluBI AGCT 5 cut(s) 237, 354, 367, 535, 925
AluI AGCT 5 cut(s) 237, 354, 367, 535, 925
Alw21I GWGCWC 3 cut(s) 239, 369, 537
Alw26I GTCTC 4 cut(s) 497, 570, 785, 799
AlwI GGATC 4 cut(s) 13, 74, 1037, 1151
Ama87I CYCGRG 2 cut(s) 130, 1004
Aor13HI TCCGGA 2 cut(s) 63, 927
AoxI GGCC 5 cut(s) 310, 556, 713, 723, 784
ApeKI GCWGC 8 cut(s) 77, 292, 321, 488, 748, 922, 1099, 1172
ApoI RAATTY 4 cut(s) 265, 698, 1033, 1078
AsiGI ACCGGT 1 cut(s) 1151
AspLEI GCGC 2 cut(s) 455, 712
AspS9I GGNCC 4 cut(s) 714, 723, 775, 963
AsuC2I CCSGG 1 cut(s) 100
AsuHPI GGTGA 5 cut(s) 61, 385, 541, 603, 641
AsuII TTCGAA 1 cut(s) 702
AsuNHI GCTAGC 1 cut(s) 1168
AvaI CYCGRG 2 cut(s) 130, 1004
AvaII GGWCC 2 cut(s) 775, 963
BanI GGYRCC 1 cut(s) 949
BanII GRGCYC 3 cut(s) 239, 369, 537
BauI CACGAG 1 cut(s) 464
Bbv12I GWGCWC 3 cut(s) 239, 369, 537
BbvI GCAGC 7 cut(s) 64, 304, 308, 500, 735, 934, 1111
BccI CCATC 4 cut(s) 250, 593, 646, 806
BceAI ACGGC 7 cut(s) 200, 211, 571, 736, 829, 838, 1013
BcgI CGANNNNNNTGC 4 cut(s) 59, 93, 923, 957
BcnI CCSGG 1 cut(s) 100
BcoDI GTCTC 4 cut(s) 497, 570, 785, 799
BfaI CTAG 1 cut(s) 1169
BglI GCCNNNNNGGC 1 cut(s) 731
Bme1390I CCNGG 1 cut(s) 100
Bme18I GGWCC 2 cut(s) 775, 963
BmeT110I CYCGRG 2 cut(s) 130, 1004
BmgT120I GGNCC 4 cut(s) 714, 723, 775, 963
BmiI GGNNCC 6 cut(s) 104, 214, 406, 724, 951, 964
BmrFI CCNGG 1 cut(s) 100
BmrI ACTGGG 1 cut(s) 729
BmtI GCTAGC 1 cut(s) 1172
BmuI ACTGGG 1 cut(s) 729
BoxI GACNNNNGTC 1 cut(s) 968
BplI GAGNNNNNCTC 2 cut(s) 221, 253
Bpu10I CCTNAGC 1 cut(s) 986
Bpu14I TTCGAA 1 cut(s) 702
BpuEI CTTGAG 2 cut(s) 419, 812
BpuMI CCSGG 1 cut(s) 100
Bsa29I ATCGAT 4 cut(s) 30, 36, 645, 684
BsaBI GATNNNNATC 3 cut(s) 17, 32, 680
BsaHI GRCGYC 4 cut(s) 372, 420, 936, 1026
BsaJI CCNNGG 2 cut(s) 42, 854
BsaWI WCCGGW 5 cut(s) 63, 927, 946, 1073, 1151
BsaXI ACNNNNNCTCC 4 cut(s) 201, 231, 566, 596
Bsc4I CCNNNNNNNGG 2 cut(s) 105, 959
Bse118I RCCGGY 7 cut(s) 85, 120, 173, 398, 626, 1105, 1151
Bse1I ACTGG 2 cut(s) 59, 724
Bse8I GATNNNNATC 3 cut(s) 17, 32, 680
BseAI TCCGGA 2 cut(s) 63, 927
BseCI ATCGAT 4 cut(s) 30, 36, 645, 684
BseDI CCNNGG 2 cut(s) 42, 854
BseJI GATNNNNATC 3 cut(s) 17, 32, 680
BseLI CCNNNNNNNGG 2 cut(s) 105, 959
BseMII CTCAG 2 cut(s) 550, 977
BseNI ACTGG 2 cut(s) 59, 724
BseRI GAGGAG 7 cut(s) 139, 216, 219, 222, 225, 571, 1149
BseXI GCAGC 7 cut(s) 64, 304, 308, 500, 735, 934, 1111
Bsh1236I CGCG 2 cut(s) 455, 712
Bsh1285I CGRYCG 2 cut(s) 30, 695
BshFI GGCC 5 cut(s) 312, 558, 715, 725, 786
BshNI GGYRCC 1 cut(s) 949
BshTI ACCGGT 1 cut(s) 1151
BshVI ATCGAT 4 cut(s) 30, 36, 645, 684
BsiEI CGRYCG 2 cut(s) 30, 695
BsiHKAI GWGCWC 3 cut(s) 239, 369, 537
BsiHKCI CYCGRG 2 cut(s) 130, 1004
BslFI GGGAC 3 cut(s) 617, 677, 976
BslI CCNNNNNNNGG 2 cut(s) 105, 959
BsmAI GTCTC 4 cut(s) 497, 570, 785, 799
BsmBI CGTCTC 3 cut(s) 497, 570, 785
BsmFI GGGAC 3 cut(s) 617, 677, 976
BsmI GAATGC 1 cut(s) 846
BsnI GGCC 5 cut(s) 312, 558, 715, 725, 786
BsoBI CYCGRG 2 cut(s) 130, 1004
Bsp119I TTCGAA 1 cut(s) 702
Bsp1286I GDGCHC 3 cut(s) 239, 369, 537
Bsp13I TCCGGA 2 cut(s) 63, 927
Bsp143I GATC 8 cut(s) 18, 27, 66, 543, 692, 1011, 1042, 1143
BspANI GGCC 5 cut(s) 312, 558, 715, 725, 786
BspCNI CTCAG 2 cut(s) 549, 978
BspDI ATCGAT 4 cut(s) 30, 36, 645, 684
BspEI TCCGGA 2 cut(s) 63, 927
BspFNI CGCG 2 cut(s) 455, 712
BspLI GGNNCC 6 cut(s) 104, 214, 406, 724, 951, 964
BspOI GCTAGC 1 cut(s) 1172
BspPI GGATC 4 cut(s) 13, 74, 1037, 1151
BspQI GCTCTTC 1 cut(s) 1115
BspT104I TTCGAA 1 cut(s) 702
BspT107I GGYRCC 1 cut(s) 949
BsrFI RCCGGY 7 cut(s) 85, 120, 173, 398, 626, 1105, 1151
BsrI ACTGG 2 cut(s) 59, 724
BssAI RCCGGY 7 cut(s) 85, 120, 173, 398, 626, 1105, 1151
BssECI CCNNGG 2 cut(s) 42, 854
BssMI GATC 8 cut(s) 18, 27, 66, 543, 692, 1011, 1042, 1143
BssNI GRCGYC 4 cut(s) 372, 420, 936, 1026
BssSI CACGAG 1 cut(s) 464
Bst2BI CACGAG 1 cut(s) 464
Bst4CI ACNGT 5 cut(s) 169, 615, 779, 967, 1182
Bst6I CTCTTC 1 cut(s) 1115
BstACI GRCGYC 4 cut(s) 372, 420, 936, 1026
BstBI TTCGAA 1 cut(s) 702
BstC8I GCNNGC 5 cut(s) 137, 175, 297, 1107, 1170
BstDEI CTNAG 4 cut(s) 140, 478, 536, 986
BstDSI CCRYGG 1 cut(s) 854
BstFNI CGCG 2 cut(s) 455, 712
BstHHI GCGC 2 cut(s) 455, 712
BstKTI GATC 8 cut(s) 21, 30, 69, 546, 695, 1014, 1045, 1146
BstMAI GTCTC 4 cut(s) 497, 570, 785, 799
BstMBI GATC 8 cut(s) 18, 27, 66, 543, 692, 1011, 1042, 1143
BstMCI CGRYCG 2 cut(s) 30, 695
BstMWI GCNNNNNNNGC 7 cut(s) 109, 243, 318, 327, 488, 731, 850
BstPAI GACNNNNGTC 1 cut(s) 968
BstSCI CCNGG 1 cut(s) 98
BstUI CGCG 2 cut(s) 455, 712
BstV1I GCAGC 7 cut(s) 64, 304, 308, 500, 735, 934, 1111
BstX2I RGATCY 1 cut(s) 18
BstYI RGATCY 1 cut(s) 18
Bsu15I ATCGAT 4 cut(s) 30, 36, 645, 684
BsuRI GGCC 5 cut(s) 312, 558, 715, 725, 786
BsuTUI ATCGAT 4 cut(s) 30, 36, 645, 684
BtgI CCRYGG 1 cut(s) 854
BtsIMutI CAGTG 2 cut(s) 717, 1089
Cac8I GCNNGC 5 cut(s) 137, 175, 297, 1107, 1170
CfoI GCGC 2 cut(s) 455, 712
Cfr10I RCCGGY 7 cut(s) 85, 120, 173, 398, 626, 1105, 1151
Cfr13I GGNCC 4 cut(s) 714, 723, 775, 963
ClaI ATCGAT 4 cut(s) 30, 36, 645, 684
CpoI CGGWCCG 1 cut(s) 775
CseI GACGC 3 cut(s) 428, 494, 1034
Csp6I GTAC 2 cut(s) 640, 871
CspAI ACCGGT 1 cut(s) 1151
CspI CGGWCCG 1 cut(s) 775
CviAII CATG 1 cut(s) 450
CviQI GTAC 2 cut(s) 640, 871
DdeI CTNAG 4 cut(s) 140, 478, 536, 986
DpnI GATC 8 cut(s) 20, 29, 68, 545, 694, 1013, 1044, 1145
DpnII GATC 8 cut(s) 18, 27, 66, 543, 692, 1011, 1042, 1143
EaeI YGGCCR 2 cut(s) 310, 784
Eam1104I CTCTTC 1 cut(s) 1115
EarI CTCTTC 1 cut(s) 1115
EciI GGCGGA 3 cut(s) 213, 840, 874
Ecl136II GAGCTC 3 cut(s) 237, 367, 535
Eco24I GRGCYC 3 cut(s) 239, 369, 537
Eco32I GATATC 1 cut(s) 34
Eco47I GGWCC 2 cut(s) 775, 963
Eco53kI GAGCTC 3 cut(s) 237, 367, 535
Eco88I CYCGRG 2 cut(s) 130, 1004
EcoICRI GAGCTC 3 cut(s) 237, 367, 535
EcoO109I RGGNCCY 1 cut(s) 723
EcoRI GAATTC 1 cut(s) 698
EcoRV GATATC 1 cut(s) 34
EcoT38I GRGCYC 3 cut(s) 239, 369, 537
Esp3I CGTCTC 3 cut(s) 497, 570, 785
FaeI CATG 1 cut(s) 453
FaiI YATR 1 cut(s) 451
FaqI GGGAC 3 cut(s) 617, 677, 976
FatI CATG 1 cut(s) 449
FauI CCCGC 1 cut(s) 631
FriOI GRGCYC 3 cut(s) 239, 369, 537
FspBI CTAG 1 cut(s) 1169
GlaI GCGC 2 cut(s) 454, 711
HaeIII GGCC 5 cut(s) 312, 558, 715, 725, 786
HgaI GACGC 3 cut(s) 428, 494, 1034
HhaI GCGC 2 cut(s) 455, 712
Hin1I GRCGYC 4 cut(s) 372, 420, 936, 1026
Hin1II CATG 1 cut(s) 453
Hin6I GCGC 2 cut(s) 453, 710
HinP1I GCGC 2 cut(s) 453, 710
HinfI GANTC 8 cut(s) 38, 71, 144, 316, 386, 439, 445, 578
HphI GGTGA 5 cut(s) 61, 385, 541, 603, 641
Hpy166II GTNNAC 1 cut(s) 463
Hpy188III TCNNGA 7 cut(s) 64, 130, 436, 502, 829, 928, 1147
Hpy8I GTNNAC 1 cut(s) 463
HpyAV CCTTC 2 cut(s) 400, 749
HpyCH4III ACNGT 5 cut(s) 169, 615, 779, 967, 1182
HpyCH4IV ACGT 3 cut(s) 372, 873, 936
HpyCH4V TGCA 3 cut(s) 324, 591, 900
HpyF10VI GCNNNNNNNGC 7 cut(s) 109, 243, 318, 327, 488, 731, 850
HpyF3I CTNAG 4 cut(s) 140, 478, 536, 986
HpySE526I ACGT 3 cut(s) 372, 873, 936
Hsp92I GRCGYC 4 cut(s) 372, 420, 936, 1026
Hsp92II CATG 1 cut(s) 453
HspAI GCGC 2 cut(s) 453, 710
Kpn2I TCCGGA 2 cut(s) 63, 927
KroI GCCGGC 2 cut(s) 173, 1105
KroNI GCCGGC 2 cut(s) 175, 1107
Kzo9I GATC 8 cut(s) 18, 27, 66, 543, 692, 1011, 1042, 1143
LguI GCTCTTC 1 cut(s) 1115
LmnI GCTCC 6 cut(s) 108, 212, 359, 930, 949, 1136
Lsp1109I GCAGC 7 cut(s) 64, 304, 308, 500, 735, 934, 1111
MaeI CTAG 1 cut(s) 1169
MaeII ACGT 3 cut(s) 372, 873, 936
MaeIII GTNAC 4 cut(s) 547, 802, 1069, 1147
MalI GATC 8 cut(s) 20, 29, 68, 545, 694, 1013, 1044, 1145
MboI GATC 8 cut(s) 18, 27, 66, 543, 692, 1011, 1042, 1143
MboII GAAGA 6 cut(s) 28, 265, 287, 844, 1006, 1132
MfeI CAATTG 1 cut(s) 325
MflI RGATCY 1 cut(s) 18
MhlI GDGCHC 3 cut(s) 239, 369, 537
MluCI AATT 5 cut(s) 265, 325, 698, 1033, 1078
MlyI GAGTC 4 cut(s) 65, 395, 448, 572
MmeI TCCRAC 3 cut(s) 46, 381, 1002
MreI CGCCGGCG 2 cut(s) 173, 1105
MroI TCCGGA 2 cut(s) 63, 927
MroNI GCCGGC 2 cut(s) 173, 1105
MseI TTAA 2 cut(s) 333, 1037
MslI CAYNNNNRTG 1 cut(s) 647
MspR9I CCNGG 1 cut(s) 100
MunI CAATTG 1 cut(s) 325
Mva1269I GAATGC 1 cut(s) 846
MvnI CGCG 2 cut(s) 455, 712
MwoI GCNNNNNNNGC 7 cut(s) 109, 243, 318, 327, 488, 731, 850
NaeI GCCGGC 2 cut(s) 175, 1107
NciI CCSGG 1 cut(s) 100
NdeII GATC 8 cut(s) 18, 27, 66, 543, 692, 1011, 1042, 1143
NgoMIV GCCGGC 2 cut(s) 173, 1105
NheI GCTAGC 1 cut(s) 1168
NlaIII CATG 1 cut(s) 453
NlaIV GGNNCC 6 cut(s) 104, 214, 406, 724, 951, 964
NmeAIII GCCGAG 6 cut(s) 53, 247, 258, 549, 762, 892
NmuCI GTSAC 3 cut(s) 547, 1069, 1147
NspV TTCGAA 1 cut(s) 702
PaeR7I CTCGAG 1 cut(s) 1004
PciSI GCTCTTC 1 cut(s) 1115
PcsI WCGNNNNNNNCGW 9 cut(s) 177, 225, 366, 369, 642, 812, 975, 978, 987
PctI GAATGC 1 cut(s) 846
PdiI GCCGGC 2 cut(s) 175, 1107
PfeI GAWTC 4 cut(s) 38, 144, 316, 445
PinAI ACCGGT 1 cut(s) 1151
Ple19I CGATCG 2 cut(s) 30, 695
PleI GAGTC 4 cut(s) 65, 394, 447, 572
PpsI GAGTC 4 cut(s) 65, 394, 447, 572
PshAI GACNNNNGTC 1 cut(s) 968
Psp124BI GAGCTC 3 cut(s) 239, 369, 537
PspN4I GGNNCC 6 cut(s) 104, 214, 406, 724, 951, 964
PspPI GGNCC 4 cut(s) 714, 723, 775, 963
PspXI VCTCGAGB 1 cut(s) 1004
PsuI RGATCY 1 cut(s) 18
PvuI CGATCG 2 cut(s) 30, 695
RsaI GTAC 2 cut(s) 641, 872
RsaNI GTAC 2 cut(s) 640, 871
RseI CAYNNNNRTG 1 cut(s) 647
Rsr2I CGGWCCG 1 cut(s) 775
RsrII CGGWCCG 1 cut(s) 775
SacI GAGCTC 3 cut(s) 239, 369, 537
SapI GCTCTTC 1 cut(s) 1115
SaqAI TTAA 2 cut(s) 333, 1037
Sau3AI GATC 8 cut(s) 18, 27, 66, 543, 692, 1011, 1042, 1143
Sau96I GGNCC 4 cut(s) 714, 723, 775, 963
SchI GAGTC 4 cut(s) 65, 395, 448, 572
ScrFI CCNGG 1 cut(s) 100
SduI GDGCHC 3 cut(s) 239, 369, 537
Sfr274I CTCGAG 1 cut(s) 1004
SfuI TTCGAA 1 cut(s) 702
SgrAI CRCCGGYG 4 cut(s) 173, 398, 626, 1105
SinI GGWCC 2 cut(s) 775, 963
SlaI CTCGAG 1 cut(s) 1004
SmiMI CAYNNNNRTG 1 cut(s) 647
SmlI CTYRAG 3 cut(s) 434, 827, 1004
SmoI CTYRAG 3 cut(s) 434, 827, 1004
Sse9I AATT 5 cut(s) 265, 325, 698, 1033, 1078
SspMI CTAG 1 cut(s) 1169
SstI GAGCTC 3 cut(s) 239, 369, 537
StyD4I CCNGG 1 cut(s) 98
TaaI ACNGT 5 cut(s) 169, 615, 779, 967, 1182
TaiI ACGT 3 cut(s) 375, 876, 939
TasI AATT 5 cut(s) 265, 325, 698, 1033, 1078
TatI WGTACW 1 cut(s) 639
TauI GCSGC 7 cut(s) 112, 180, 384, 417, 715, 815, 1105
TfiI GAWTC 4 cut(s) 38, 144, 316, 445
Tru1I TTAA 2 cut(s) 333, 1037
Tru9I TTAA 2 cut(s) 333, 1037
TscAI CASTG 2 cut(s) 724, 1089
TseFI GTSAC 3 cut(s) 547, 1069, 1147
TseI GCWGC 8 cut(s) 77, 292, 321, 488, 748, 922, 1099, 1172
Tsp45I GTSAC 3 cut(s) 547, 1069, 1147
TspDTI ATGAA 2 cut(s) 29, 745
TspGWI ACGGA 4 cut(s) 170, 351, 821, 1035
TspRI CASTG 2 cut(s) 724, 1089
VpaK11BI GGWCC 2 cut(s) 775, 963
XapI RAATTY 4 cut(s) 265, 698, 1033, 1078
XhoI CTCGAG 1 cut(s) 1004
XspI CTAG 1 cut(s) 1169
ZraI GACGTC 2 cut(s) 373, 937
Using CommOnly database (standard laboratory enzymes). Scanned on CDS sequence.