FvH4_3g06240

Putative S-adenosyl-L-methionine-dependent methyltransferase

Design Primers CRISPR Design

Basic Information

Type: gene
Biological Identity
fragaria_vesca
Fvb3
OG0014427
Physical Location & Seq
Forward (+)
3584151 .. 3587430
3280 bp
Loading structure...
UTR
Exon/CDS
Intron
FvH4_3g06240.t2

Sequence Viewer

Length: 1098 bp
ATGGGTTTCACAATGGGTCTTAACTTGCTTCTGCTTATGGCCATGGTCGCCACCAACATCCTCTCCCTCTACCACCTCTCCTCCACCATCCAATCCCCCACCCCTCCCAACACCGCCGTCCCCGACCACCTCCTCCGCCAGCTCCAAACCATACGCGCCACCATCAACCACCTCACGCGCCACCACCCTCCTCCCCCACGCGCCGCCGCCGCCTCCTCCGCCGACCTCCTCCTCTACTCCCGCCTCTCCCCGATCGCCTCCTCCTGCCACAACCACCCCGACCTCCTCAACAAGTACATGACCTACACTCCCTTCTCCCTCTGCCCCGCCGACGCCGACGACGTCGCCGAGAGCCTCATCCTCCGCGGCTGCCACCCCCTCCCCCGCCGGAGATGCTTCTCCAGAACGCCGCAGAAGCCCACGTCATCGCTCTCCCACGACGCTTTCCCTCCCTCTCTCCCGGATCAGAATGTGATATGGGACAAGTACGCCTGCAAAAGCTTCAGCTGCCTCAACCGCCGTGGCCTCAGCTCCGGCTTCGACTTCAGCCACGAGCTCTCCAACTTCATGACCTACAAGTCCGAGCTGGACCTCCCCATCCCGCAGTTCTTCCAGATCGCCAAGGCCGCCGGCGCCGTCCTCCGCCTCGGGCTGGACGTCGGCGGCGGCACCGGGACCTTCGCCGCCAGGATGAAGCTCTACAATGTGACAGTTGTCACCACCACCATGAATTTCGGCAGTCCGAATAACGAGGCGGTGGCGCTGAGGGGGCTGGTGCCGCTGCACGTGCCGCTGCAGCAGCGGTTCCCCGTGTTTGACGGGGTGTTGGATCTGGTGCGGTGCGGGCACGCGGTGAACAGGTGGATTCCGGTGACGGCGTTGGAGTTTTTGTTGTACGATGCGGATAGGGTGTTGAGGGTAGGAGGGTACTTGTGGTTGGATCATTTCTTTAGCAAGGGGGTGGATCTTGAGAAGGTTTTCGGGCCGTTGATTGGGAAGTTGGGGTACCGGAGAGTGAAGTGGGCGACGGCGAGTAAGAATGAGAAGAATGGGGAGGTTTACTTGACTGCTCTTCTGCAAAAGCCTGTCTTGAGATGA
Functional Annotation
Gene Ontology
Molecular Function
Biological Process
Cellular Component
No ontology terms assigned.
KEGG Pathways
Metabolic & Signaling
No pathways identified.
Pfam Domains
Protein Families
Methyltransf_11 Methyltransf_29

Protein Analysis

366

Amino Acids

40.48

Weight (kDa)

9.39

Isoelectric Point (pI)

49.1

Instability Index
Protein Domains (Pfam)
No Pfam domains detected for this protein.
Hydrophobicity Profile (Kyte-Doolittle)
AI Structure Prediction Report
Calculating structure properties...

Genomic Context

Gene Family Tree

Species Filter
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Publication-ready
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Orthologous Genes (Group: OG0014427)

Species Orthologous Gene IDs
arabidopsis_thaliana AT1G29790 AT1G29790
fragaria_vesca FvH4_3g06240 FvH4_3g06240
malus_domestica MD05G1290600.v1.1 MD16G1203100.v1.1
prunus_persica Prupe.4G075100_v2.0.a1
pyrus_communis pycom10g22250
rosa_chinensis RchiOBHm_Chr5g0013071
rosa_laevigata RLG00000031983
rosa_multiflora Rmu_sc0003892.1_g000015
rosa_rugosa Rorug05G0007000
rosa_samantha Rh5AG101000 Rh5BG097600 Rh5CG109700 Rh5DG096600
rosa_wichuraiana Rw5G008830

Restriction Enzyme Sites

1 / 10
Enzyme Recognition Site Cut Count Positions (bp)
AasI GACNNNNNNGTC 1 cut(s) 577
AatII GACGTC 2 cut(s) 345, 660
Acc65I GGTACC 1 cut(s) 1005
AccB1I GGYRCC 4 cut(s) 632, 668, 775, 1005
AccII CGCG 5 cut(s) 156, 178, 201, 366, 851
AclWI GGATC 4 cut(s) 471, 837, 948, 972
AcoI YGGCCR 1 cut(s) 39
AcsI RAATTY 1 cut(s) 730
AcuI CTGAAG 2 cut(s) 487, 529
AcvI CACGTG 1 cut(s) 787
AcyI GRCGYC 4 cut(s) 333, 342, 633, 657
AdeI CACNNNGTG 1 cut(s) 853
AfaI GTAC 5 cut(s) 296, 488, 896, 929, 1007
AfiI CCNNNNNNNGG 2 cut(s) 652, 992
AjiI CACGTC 1 cut(s) 423
AjnI CCWGG 1 cut(s) 686
AluBI AGCT 7 cut(s) 142, 501, 507, 531, 556, 586, 697
AluI AGCT 7 cut(s) 142, 501, 507, 531, 556, 586, 697
Alw21I GWGCWC 1 cut(s) 558
AlwI GGATC 4 cut(s) 471, 837, 948, 972
Ama87I CYCGRG 1 cut(s) 647
AoxI GGCC 4 cut(s) 39, 523, 624, 983
ApeKI GCWGC 6 cut(s) 369, 507, 781, 793, 796, 799
ApoI RAATTY 1 cut(s) 730
Asp700I GAANNNNTTC 1 cut(s) 977
Asp718I GGTACC 1 cut(s) 1005
AspLEI GCGC 5 cut(s) 158, 180, 203, 635, 763
AspS9I GGNCC 3 cut(s) 589, 675, 983
AsuC2I CCSGG 2 cut(s) 461, 673
AsuHPI GGTGA 3 cut(s) 709, 865, 883
AvaI CYCGRG 1 cut(s) 647
AvaII GGWCC 2 cut(s) 589, 675
BaeGI GKGCMC 1 cut(s) 849
BalI TGGCCA 1 cut(s) 41
BanI GGYRCC 4 cut(s) 632, 668, 775, 1005
BanII GRGCYC 1 cut(s) 558
BarI GAAGNNNNNNTAC 2 cut(s) 989, 1021
BauI CACGAG 1 cut(s) 551
BbrPI CACGTG 1 cut(s) 787
Bbv12I GWGCWC 1 cut(s) 558
BbvCI CCTCAGC 2 cut(s) 527, 764
BbvI GCAGC 6 cut(s) 356, 494, 768, 780, 808, 811
BccI CCATC 3 cut(s) 95, 170, 605
BceAI ACGGC 6 cut(s) 101, 504, 620, 891, 970, 1044
BciT130I CCWGG 1 cut(s) 688
BcnI CCSGG 2 cut(s) 461, 673
BfmI CTRYAG 1 cut(s) 794
BfoI RGCGCY 2 cut(s) 636, 764
Bme1390I CCNGG 3 cut(s) 461, 673, 688
Bme18I GGWCC 2 cut(s) 589, 675
BmeT110I CYCGRG 1 cut(s) 647
BmgBI CACGTC 1 cut(s) 423
BmgT120I GGNCC 3 cut(s) 589, 675, 983
BmiI GGNNCC 6 cut(s) 634, 670, 676, 777, 806, 1007
BmrFI CCNGG 3 cut(s) 461, 673, 688
BmsI GCATC 2 cut(s) 383, 889
BoxI GACNNNNGTC 1 cut(s) 713
BpmI CTGGAG 1 cut(s) 385
Bpu10I CCTNAGC 2 cut(s) 527, 764
BpuEI CTTGAG 1 cut(s) 989
BpuMI CCSGG 2 cut(s) 461, 673
BsaAI YACGTR 1 cut(s) 787
BsaHI GRCGYC 4 cut(s) 333, 342, 633, 657
BsaJI CCNNGG 5 cut(s) 42, 364, 520, 621, 646
BsaWI WCCGGW 2 cut(s) 868, 1008
Bsc4I CCNNNNNNNGG 2 cut(s) 652, 992
Bse118I RCCGGY 1 cut(s) 629
BseBI CCWGG 1 cut(s) 688
BseDI CCNNGG 5 cut(s) 42, 364, 520, 621, 646
BseGI GGATG 5 cut(s) 57, 87, 357, 597, 696
BseLI CCNNNNNNNGG 2 cut(s) 652, 992
BseMII CTCAG 2 cut(s) 541, 755
BseRI GAGGAG 8 cut(s) 70, 122, 180, 205, 218, 221, 250, 275
BseSI GKGCMC 1 cut(s) 849
BseXI GCAGC 6 cut(s) 356, 494, 768, 780, 808, 811
BsgI GTGCAG 1 cut(s) 767
Bsh1236I CGCG 5 cut(s) 156, 178, 201, 366, 851
Bsh1285I CGRYCG 1 cut(s) 255
BshFI GGCC 4 cut(s) 41, 525, 626, 985
BshNI GGYRCC 4 cut(s) 632, 668, 775, 1005
BsiEI CGRYCG 1 cut(s) 255
BsiHKAI GWGCWC 1 cut(s) 558
BsiHKCI CYCGRG 1 cut(s) 647
BsiSI CCGG 7 cut(s) 388, 461, 534, 630, 672, 869, 1009
BslFI GGGAC 3 cut(s) 104, 494, 688
BslI CCNNNNNNNGG 2 cut(s) 652, 992
BsmFI GGGAC 3 cut(s) 104, 494, 688
BsnI GGCC 4 cut(s) 41, 525, 626, 985
BsoBI CYCGRG 1 cut(s) 647
Bsp1286I GDGCHC 2 cut(s) 558, 849
Bsp143I GATC 6 cut(s) 252, 463, 615, 829, 940, 964
Bsp19I CCATGG 1 cut(s) 42
BspANI GGCC 4 cut(s) 41, 525, 626, 985
BspCNI CTCAG 2 cut(s) 540, 756
BspFNI CGCG 5 cut(s) 156, 178, 201, 366, 851
BspHI TCATGA 1 cut(s) 567
BspLI GGNNCC 6 cut(s) 634, 670, 676, 777, 806, 1007
BspMAI CTGCAG 1 cut(s) 798
BspPI GGATC 4 cut(s) 471, 837, 948, 972
BspQI GCTCTTC 1 cut(s) 1077
BspT107I GGYRCC 4 cut(s) 632, 668, 775, 1005
BsrFI RCCGGY 1 cut(s) 629
BssAI RCCGGY 1 cut(s) 629
BssECI CCNNGG 5 cut(s) 42, 364, 520, 621, 646
BssMI GATC 6 cut(s) 252, 463, 615, 829, 940, 964
BssNI GRCGYC 4 cut(s) 333, 342, 633, 657
BssSI CACGAG 1 cut(s) 551
BssT1I CCWWGG 2 cut(s) 42, 621
Bst2BI CACGAG 1 cut(s) 551
Bst2UI CCWGG 1 cut(s) 688
Bst4CI ACNGT 1 cut(s) 712
Bst6I CTCTTC 1 cut(s) 1077
BstACI GRCGYC 4 cut(s) 333, 342, 633, 657
BstBAI YACGTR 1 cut(s) 787
BstC8I GCNNGC 5 cut(s) 140, 493, 631, 845, 849
BstDEI CTNAG 2 cut(s) 527, 764
BstDSI CCRYGG 3 cut(s) 42, 364, 520
BstF5I GGATG 5 cut(s) 57, 87, 357, 597, 696
BstFNI CGCG 5 cut(s) 156, 178, 201, 366, 851
BstH2I RGCGCY 2 cut(s) 636, 764
BstHHI GCGC 5 cut(s) 158, 180, 203, 635, 763
BstKTI GATC 6 cut(s) 255, 466, 618, 832, 943, 967
BstMBI GATC 6 cut(s) 252, 463, 615, 829, 940, 964
BstMCI CGRYCG 1 cut(s) 255
BstNI CCWGG 1 cut(s) 688
BstPAI GACNNNNGTC 1 cut(s) 713
BstSCI CCNGG 3 cut(s) 459, 671, 686
BstSFI CTRYAG 1 cut(s) 794
BstSLI GKGCMC 1 cut(s) 849
BstUI CGCG 5 cut(s) 156, 178, 201, 366, 851
BstV1I GCAGC 6 cut(s) 356, 494, 768, 780, 808, 811
BstX2I RGATCY 2 cut(s) 829, 964
BstYI RGATCY 2 cut(s) 829, 964
BsuRI GGCC 4 cut(s) 41, 525, 626, 985
BtgI CCRYGG 3 cut(s) 42, 364, 520
BtgZI GCGATG 1 cut(s) 411
BtrI CACGTC 1 cut(s) 423
BtsCI GGATG 5 cut(s) 57, 87, 357, 597, 696
Cac8I GCNNGC 5 cut(s) 140, 493, 631, 845, 849
CciI TCATGA 1 cut(s) 567
CfoI GCGC 5 cut(s) 158, 180, 203, 635, 763
Cfr10I RCCGGY 1 cut(s) 629
Cfr13I GGNCC 3 cut(s) 589, 675, 983
Cfr42I CCGCGG 1 cut(s) 367
CseI GACGC 2 cut(s) 341, 449
Csp6I GTAC 5 cut(s) 295, 487, 895, 928, 1006
CspCI CAANNNNNGTGG 2 cut(s) 502, 537
CviAII CATG 4 cut(s) 43, 298, 568, 727
CviQI GTAC 5 cut(s) 295, 487, 895, 928, 1006
DdeI CTNAG 2 cut(s) 527, 764
DinI GGCGCC 1 cut(s) 634
DpnI GATC 6 cut(s) 254, 465, 617, 831, 942, 966
DpnII GATC 6 cut(s) 252, 463, 615, 829, 940, 964
DraIII CACNNNGTG 1 cut(s) 853
DrdI GACNNNNNNGTC 1 cut(s) 577
DseDI GACNNNNNNGTC 1 cut(s) 577
EaeI YGGCCR 1 cut(s) 39
Eam1104I CTCTTC 1 cut(s) 1077
EarI CTCTTC 1 cut(s) 1077
EciI GGCGGA 3 cut(s) 125, 208, 632
Ecl136II GAGCTC 1 cut(s) 556
Eco130I CCWWGG 2 cut(s) 42, 621
Eco24I GRGCYC 1 cut(s) 558
Eco47I GGWCC 2 cut(s) 589, 675
Eco53kI GAGCTC 1 cut(s) 556
Eco57I CTGAAG 2 cut(s) 487, 529
Eco72I CACGTG 1 cut(s) 787
Eco88I CYCGRG 1 cut(s) 647
EcoICRI GAGCTC 1 cut(s) 556
EcoO109I RGGNCCY 1 cut(s) 675
EcoRII CCWGG 1 cut(s) 686
EcoT14I CCWWGG 2 cut(s) 42, 621
EcoT38I GRGCYC 1 cut(s) 558
EgeI GGCGCC 1 cut(s) 634
EheI GGCGCC 1 cut(s) 634
ErhI CCWWGG 2 cut(s) 42, 621
FaeI CATG 4 cut(s) 46, 301, 571, 730
FaiI YATR 7 cut(s) 38, 44, 152, 299, 478, 569, 728
FalI AAGNNNNNCTT 1 cut(s) 1073
FaqI GGGAC 3 cut(s) 104, 494, 688
FatI CATG 4 cut(s) 42, 297, 567, 726
FauI CCCGC 5 cut(s) 248, 334, 392, 609, 836
FokI GGATG 5 cut(s) 44, 74, 344, 584, 703
FriOI GRGCYC 1 cut(s) 558
GlaI GCGC 5 cut(s) 157, 179, 202, 634, 762
GsuI CTGGAG 1 cut(s) 385
HaeII RGCGCY 2 cut(s) 636, 764
HaeIII GGCC 4 cut(s) 41, 525, 626, 985
HapII CCGG 7 cut(s) 388, 461, 534, 630, 672, 869, 1009
HgaI GACGC 2 cut(s) 341, 449
HhaI GCGC 5 cut(s) 158, 180, 203, 635, 763
Hin1I GRCGYC 4 cut(s) 333, 342, 633, 657
Hin1II CATG 4 cut(s) 46, 301, 571, 730
Hin6I GCGC 5 cut(s) 156, 178, 201, 633, 761
HinP1I GCGC 5 cut(s) 156, 178, 201, 633, 761
HindIII AAGCTT 1 cut(s) 499
HinfI GANTC 1 cut(s) 865
HpaII CCGG 7 cut(s) 388, 461, 534, 630, 672, 869, 1009
HphI GGTGA 3 cut(s) 709, 865, 883
Hpy166II GTNNAC 2 cut(s) 856, 1060
Hpy188I TCNGA 3 cut(s) 468, 583, 744
Hpy188III TCNNGA 5 cut(s) 402, 568, 613, 968, 1090
Hpy8I GTNNAC 2 cut(s) 856, 1060
Hpy99I CGWCG 7 cut(s) 335, 341, 344, 347, 443, 662, 1030
HpyAV CCTTC 3 cut(s) 322, 688, 967
HpyCH4III ACNGT 1 cut(s) 712
HpyCH4IV ACGT 4 cut(s) 342, 422, 657, 786
HpyCH4V TGCA 4 cut(s) 495, 784, 796, 1078
HpyF3I CTNAG 2 cut(s) 527, 764
HpySE526I ACGT 4 cut(s) 342, 422, 657, 786
Hsp92I GRCGYC 4 cut(s) 333, 342, 633, 657
Hsp92II CATG 4 cut(s) 46, 301, 571, 730
HspAI GCGC 5 cut(s) 156, 178, 201, 633, 761
KasI GGCGCC 1 cut(s) 632
KpnI GGTACC 1 cut(s) 1009
KroI GCCGGC 1 cut(s) 629
KroNI GCCGGC 1 cut(s) 631
KspI CCGCGG 1 cut(s) 367
Kzo9I GATC 6 cut(s) 252, 463, 615, 829, 940, 964
LguI GCTCTTC 1 cut(s) 1077
LmnI GCTCC 2 cut(s) 147, 536
Lsp1109I GCAGC 6 cut(s) 356, 494, 768, 780, 808, 811
LweI GCATC 2 cut(s) 383, 889
MaeII ACGT 4 cut(s) 342, 422, 657, 786
MaeIII GTNAC 3 cut(s) 706, 715, 871
MalI GATC 6 cut(s) 254, 465, 617, 831, 942, 966
MboI GATC 6 cut(s) 252, 463, 615, 829, 940, 964
MboII GAAGA 3 cut(s) 601, 1057, 1064
MflI RGATCY 2 cut(s) 829, 964
MhlI GDGCHC 2 cut(s) 558, 849
MlsI TGGCCA 1 cut(s) 41
MluCI AATT 1 cut(s) 730
MluNI TGGCCA 1 cut(s) 41
Mly113I GGCGCC 1 cut(s) 633
MmeI TCCRAC 4 cut(s) 585, 807, 861, 918
Mox20I TGGCCA 1 cut(s) 41
MreI CGCCGGCG 1 cut(s) 629
MroNI GCCGGC 1 cut(s) 629
MroXI GAANNNNTTC 1 cut(s) 977
MscI TGGCCA 1 cut(s) 41
MseI TTAA 1 cut(s) 21
MslI CAYNNNNRTG 1 cut(s) 725
Msp20I TGGCCA 1 cut(s) 41
MspA1I CMGCKG 5 cut(s) 366, 507, 781, 793, 802
MspI CCGG 7 cut(s) 388, 461, 534, 630, 672, 869, 1009
MspR9I CCNGG 3 cut(s) 461, 673, 688
MvaI CCWGG 1 cut(s) 688
MvnI CGCG 5 cut(s) 156, 178, 201, 366, 851
NaeI GCCGGC 1 cut(s) 631
NarI GGCGCC 1 cut(s) 633
NciI CCSGG 2 cut(s) 461, 673
NcoI CCATGG 1 cut(s) 42
NdeII GATC 6 cut(s) 252, 463, 615, 829, 940, 964
NgoMIV GCCGGC 1 cut(s) 629
NlaIII CATG 4 cut(s) 46, 301, 571, 730
NlaIV GGNNCC 6 cut(s) 634, 670, 676, 777, 806, 1007
NmeAIII GCCGAG 1 cut(s) 373
NmuCI GTSAC 3 cut(s) 706, 715, 871
PagI TCATGA 1 cut(s) 567
PciSI GCTCTTC 1 cut(s) 1077
PcsI WCGNNNNNNNCGW 3 cut(s) 339, 345, 654
PdiI GCCGGC 1 cut(s) 631
PdmI GAANNNNTTC 1 cut(s) 977
PfeI GAWTC 1 cut(s) 865
PflFI GACNNNGTC 1 cut(s) 341
PfoI TCCNGGA 1 cut(s) 459
Ple19I CGATCG 1 cut(s) 255
PluTI GGCGCC 1 cut(s) 636
PmaCI CACGTG 1 cut(s) 787
PmlI CACGTG 1 cut(s) 787
Ppu21I YACGTR 1 cut(s) 787
PpuMI RGGWCCY 1 cut(s) 675
PshAI GACNNNNGTC 1 cut(s) 713
Psp124BI GAGCTC 1 cut(s) 558
Psp5II RGGWCCY 1 cut(s) 675
Psp6I CCWGG 1 cut(s) 686
PspCI CACGTG 1 cut(s) 787
PspGI CCWGG 1 cut(s) 686
PspN4I GGNNCC 6 cut(s) 634, 670, 676, 777, 806, 1007
PspPI GGNCC 3 cut(s) 589, 675, 983
PspPPI RGGWCCY 1 cut(s) 675
PstI CTGCAG 1 cut(s) 798
PsuI RGATCY 2 cut(s) 829, 964
PsyI GACNNNGTC 1 cut(s) 341
PvuI CGATCG 1 cut(s) 255
PvuII CAGCTG 1 cut(s) 507
RsaI GTAC 5 cut(s) 296, 488, 896, 929, 1007
RsaNI GTAC 5 cut(s) 295, 487, 895, 928, 1006
RseI CAYNNNNRTG 1 cut(s) 725
SacI GAGCTC 1 cut(s) 558
SacII CCGCGG 1 cut(s) 367
SapI GCTCTTC 1 cut(s) 1077
SaqAI TTAA 1 cut(s) 21
Sau3AI GATC 6 cut(s) 252, 463, 615, 829, 940, 964
Sau96I GGNCC 3 cut(s) 589, 675, 983
ScrFI CCNGG 3 cut(s) 461, 673, 688
SduI GDGCHC 2 cut(s) 558, 849
SfaNI GCATC 2 cut(s) 383, 889
SfcI CTRYAG 1 cut(s) 794
SfoI GGCGCC 1 cut(s) 634
Sfr303I CCGCGG 1 cut(s) 367
SgrAI CRCCGGYG 1 cut(s) 629
SgrBI CCGCGG 1 cut(s) 367
SinI GGWCC 2 cut(s) 589, 675
SmiMI CAYNNNNRTG 1 cut(s) 725
SmlI CTYRAG 2 cut(s) 968, 1090
SmoI CTYRAG 2 cut(s) 968, 1090
Sse9I AATT 1 cut(s) 730
SspDI GGCGCC 1 cut(s) 632
SstI GAGCTC 1 cut(s) 558
StyD4I CCNGG 3 cut(s) 459, 671, 686
StyI CCWWGG 2 cut(s) 42, 621
TaaI ACNGT 1 cut(s) 712
TaiI ACGT 4 cut(s) 345, 425, 660, 789
TaqI TCGA 1 cut(s) 540
TasI AATT 1 cut(s) 730
TatI WGTACW 1 cut(s) 294
TfiI GAWTC 1 cut(s) 865
Tru1I TTAA 1 cut(s) 21
Tru9I TTAA 1 cut(s) 21
TseFI GTSAC 3 cut(s) 706, 715, 871
TseI GCWGC 6 cut(s) 369, 507, 781, 793, 796, 799
Tsp45I GTSAC 3 cut(s) 706, 715, 871
TspDTI ATGAA 3 cut(s) 556, 707, 743
Tth111I GACNNNGTC 1 cut(s) 341
VpaK11BI GGWCC 2 cut(s) 589, 675
XapI RAATTY 1 cut(s) 730
XmnI GAANNNNTTC 1 cut(s) 977
ZraI GACGTC 2 cut(s) 343, 658
Using CommOnly database (standard laboratory enzymes). Scanned on CDS sequence.