FvH4_6g12650

expressed protein

Design Primers CRISPR Design

Basic Information

Type: gene
Biological Identity
fragaria_vesca
Fvb6
OG0011807
Physical Location & Seq
Reverse (-)
7649350 .. 7652099
2750 bp
Loading structure...
UTR
Exon/CDS
Intron
FvH4_6g12650.t1

Sequence Viewer

Length: 1197 bp
ATGGCTGATGAGTTCGACCACGCCGCGTTTTGGTTACCCCAGGAGATTCTGACCGACTCTGACGGCGAAATCAGCCCCAAGTTTGGCTCCAGTAGCCGGAAGATGAGCTCGTTCGGCTCTGACATTGATGCTGACTCCAAGCCGGCTCTCTTTCCTTTCGAGTTTCCTTACTCCGCGTTTGGGGCTTCCACGGTGGAGTCCAGCGAGACTGAGAGCGACGAGGAGGAGTATCTTGCTGGATTGACTCGCCACCTGGCTCAGTCCTCTCTCGACGATGAGTTCAAGCGCTATGACTCGGTTCTTGGCTCCAAGAACAGCTCCAAGCCGTGGGTTTCGTCTGGCTCACCGCAATCGACTCTGTGCGCGGCCGGGAGTAGTTGCAGCTGCGGCGGACAAGGCTCGAGCCGGGGAAGCCCGAACGCTCAGTCGCCGCCGGCCACTTGGGATCTGCTACATGCGGCTGCAGGGGAAGTGGCCAAGATGAGCATCCGGGAAGAAGCTATGCGTTTCAACCAAAAAGGCAGAGGCATCTTGGGTGCTCCAAGAGAGACCTCTCCGGTCGCTGTCTCTGATTTCGGGTACTACCTCCAGCACCATCAGCAGTCGCTTTCCCAGAAGCACTTGCAGTTTGAGCGGTTGAGACAGCAACAGATTCTGAAGAACAACCTCTCGGCGGCGATTGCTCAGTCTCAGGCGAACCAACGATTCCAACATCTGATGGTCCAGAACAGGGTCAGGAACACCAACAATGGGCCTCTGGGTCTTTCGCCGTCGGCTTGGCCTCCACTGCAACAAGCTCAACGGCAACAGTTTCAGCCCAATGGGCCTAGATTCGGAGGTGCTGCTGTCGTCGGAGCAGCCAAGGAGTCTACTGGTACTGGTGTGTTCTTGCCCCGCCAAATCGGAGCCCCAGCTGAGTTTCCTCGCGGCAAGAAGCAACAACCCAAACAAGCTAGCACAACGGTTTTCGTCCCAGCAAAGGTAGTGCATGCATTGAAGTTGAACGTTGAGGAAGTGTGCGCACCGTCACAGGCTCAAGTCCATATCCCTCTCAACGGAAGAAACACTCCTGACTATGAACTTGCTCTTCGGCTTCGGAATGACAGAATTGCCATGGCTCAGCAGAATCGCAGCATGATGAGGCCACAGGCACCAGTGAACCATGAGATTAGGCTTCCTCAGGAGTGGACTTACTAG
Functional Annotation
Gene Ontology
Molecular Function
Biological Process
Cellular Component
No ontology terms assigned.
KEGG Pathways
Metabolic & Signaling
No pathways identified.
Pfam Domains
Protein Families
No domains found.

Protein Analysis

399

Amino Acids

43.59

Weight (kDa)

8.29

Isoelectric Point (pI)

64.21

Instability Index
Protein Domains (Pfam)
No Pfam domains detected for this protein.
Hydrophobicity Profile (Kyte-Doolittle)
AI Structure Prediction Report
Calculating structure properties...

Genomic Context

Gene Family Tree

Species Filter
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Publication-ready
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Orthologous Genes (Group: OG0011807)

Species Orthologous Gene IDs
arabidopsis_thaliana AT3G54000 AT3G54000 AT3G54000
fragaria_vesca FvH4_6g12650
malus_domestica MD04G1140200.v1.1 MD12G1155400.v1.1 MD15G1149300.v1.1
prunus_persica Prupe.6G265300_v2.0.a1
pyrus_communis pycom04g12780 pycom12g14750
rosa_chinensis RchiOBHm_Chr3g0464501
rosa_laevigata RLG00000024691
rosa_multiflora Rmu_sc0003074.1_g000005
rosa_roxburghii Rroxscaffold_6G00415610
rosa_rugosa Rorug03G0072700
rosa_samantha Rh3BG134900 Rh3CG137400 Rh3DG136200
rosa_wichuraiana Rw3G011030

Restriction Enzyme Sites

1 / 10
Enzyme Recognition Site Cut Count Positions (bp)
Acc16I TGCGCA 1 cut(s) 1021
AccB1I GGYRCC 1 cut(s) 1150
AccB7I CCANNNNNTGG 1 cut(s) 327
AccBSI CCGCTC 1 cut(s) 634
AccI GTMKAC 1 cut(s) 869
AccII CGCG 4 cut(s) 26, 176, 365, 927
AclI AACGTT 1 cut(s) 1005
AclWI GGATC 1 cut(s) 453
AcoI YGGCCR 3 cut(s) 366, 435, 474
AcuI CTGAAG 1 cut(s) 677
AfaI GTAC 2 cut(s) 581, 877
AfeI AGCGCT 1 cut(s) 287
AgsI TTSAA 4 cut(s) 283, 511, 997, 1003
AjnI CCWGG 2 cut(s) 39, 252
AluBI AGCT 7 cut(s) 108, 318, 384, 500, 797, 914, 953
AluI AGCT 7 cut(s) 108, 318, 384, 500, 797, 914, 953
Alw21I GWGCWC 2 cut(s) 110, 541
Alw26I GTCTC 5 cut(s) 200, 542, 571, 634, 693
AlwI GGATC 1 cut(s) 453
AlwNI CAGNNNCTG 1 cut(s) 655
Ama87I CYCGRG 1 cut(s) 400
Aor51HI AGCGCT 1 cut(s) 287
AoxI GGCC 7 cut(s) 366, 435, 474, 752, 779, 824, 1142
ApeKI GCWGC 6 cut(s) 381, 384, 461, 842, 857, 1131
ArsI GACNNNNNNTTYG 2 cut(s) 410, 442
AspLEI GCGC 3 cut(s) 288, 365, 1022
AspS9I GGNCC 3 cut(s) 721, 752, 824
AsuC2I CCSGG 3 cut(s) 370, 407, 491
AsuHPI GGTGA 1 cut(s) 336
AsuNHI GCTAGC 1 cut(s) 953
AvaI CYCGRG 1 cut(s) 400
AvaII GGWCC 1 cut(s) 721
AxyI CCTNAGG 1 cut(s) 1179
BaeI ACNNNNGTAYC 2 cut(s) 867, 900
BalI TGGCCA 1 cut(s) 476
BanI GGYRCC 1 cut(s) 1150
BanII GRGCYC 2 cut(s) 110, 910
Bbv12I GWGCWC 2 cut(s) 110, 541
BbvI GCAGC 6 cut(s) 371, 393, 448, 829, 869, 1143
BccI CCATC 2 cut(s) 603, 712
BceAI ACGGC 4 cut(s) 79, 310, 754, 818
BciT130I CCWGG 2 cut(s) 41, 254
BcnI CCSGG 3 cut(s) 370, 407, 491
BcoDI GTCTC 5 cut(s) 200, 542, 571, 634, 693
BfaI CTAG 3 cut(s) 828, 954, 1195
BfmI CTRYAG 1 cut(s) 462
BfoI RGCGCY 1 cut(s) 289
BglI GCCNNNNNGGC 1 cut(s) 823
BlpI GCTNAGC 1 cut(s) 1119
Bme1390I CCNGG 5 cut(s) 41, 254, 370, 407, 491
Bme18I GGWCC 1 cut(s) 721
BmeT110I CYCGRG 1 cut(s) 400
BmgT120I GGNCC 3 cut(s) 721, 752, 824
BmiI GGNNCC 4 cut(s) 88, 307, 907, 1152
BmrFI CCNGG 5 cut(s) 41, 254, 370, 407, 491
BmsI GCATC 3 cut(s) 118, 495, 537
BmtI GCTAGC 1 cut(s) 957
BpmI CTGGAG 2 cut(s) 73, 572
Bpu1102I GCTNAGC 1 cut(s) 1119
BpuEI CTTGAG 1 cut(s) 1020
BpuMI CCSGG 3 cut(s) 370, 407, 491
BsaBI GATNNNNATC 1 cut(s) 485
BsaI GGTCTC 1 cut(s) 542
BsaJI CCNNGG 6 cut(s) 39, 189, 326, 406, 861, 1113
BsaWI WCCGGW 1 cut(s) 556
Bse118I RCCGGY 2 cut(s) 142, 433
Bse1I ACTGG 4 cut(s) 90, 877, 883, 1154
Bse21I CCTNAGG 1 cut(s) 1179
Bse8I GATNNNNATC 1 cut(s) 485
BseBI CCWGG 2 cut(s) 41, 254
BseDI CCNNGG 6 cut(s) 39, 189, 326, 406, 861, 1113
BseGI GGATG 1 cut(s) 486
BseJI GATNNNNATC 1 cut(s) 485
BseMII CTCAG 8 cut(s) 201, 272, 437, 698, 704, 906, 1133, 1193
BseNI ACTGG 4 cut(s) 90, 877, 883, 1154
BseRI GAGGAG 2 cut(s) 236, 239
BseX3I CGGCCG 1 cut(s) 366
BseXI GCAGC 6 cut(s) 371, 393, 448, 829, 869, 1143
BseYI CCCAGC 2 cut(s) 910, 973
Bsh1236I CGCG 4 cut(s) 26, 176, 365, 927
Bsh1285I CGRYCG 2 cut(s) 369, 561
BshFI GGCC 7 cut(s) 368, 437, 476, 754, 781, 826, 1144
BshNI GGYRCC 1 cut(s) 1150
BsiEI CGRYCG 2 cut(s) 369, 561
BsiHKAI GWGCWC 2 cut(s) 110, 541
BsiHKCI CYCGRG 1 cut(s) 400
BsiSI CCGG 7 cut(s) 97, 143, 369, 406, 434, 490, 557
BslFI GGGAC 1 cut(s) 956
BsmAI GTCTC 5 cut(s) 200, 542, 571, 634, 693
BsmFI GGGAC 1 cut(s) 956
BsnI GGCC 7 cut(s) 368, 437, 476, 754, 781, 826, 1144
Bso31I GGTCTC 1 cut(s) 542
BsoBI CYCGRG 1 cut(s) 400
Bsp1286I GDGCHC 3 cut(s) 110, 541, 910
Bsp143I GATC 1 cut(s) 445
Bsp1720I GCTNAGC 1 cut(s) 1119
Bsp19I CCATGG 1 cut(s) 1113
BspANI GGCC 7 cut(s) 368, 437, 476, 754, 781, 826, 1144
BspCNI CTCAG 8 cut(s) 202, 271, 436, 697, 703, 907, 1132, 1192
BspFNI CGCG 4 cut(s) 26, 176, 365, 927
BspLI GGNNCC 4 cut(s) 88, 307, 907, 1152
BspMAI CTGCAG 1 cut(s) 466
BspOI GCTAGC 1 cut(s) 957
BspPI GGATC 1 cut(s) 453
BspQI GCTCTTC 1 cut(s) 1092
BspT107I GGYRCC 1 cut(s) 1150
BspTNI GGTCTC 1 cut(s) 542
BsrBI CCGCTC 1 cut(s) 634
BsrFI RCCGGY 2 cut(s) 142, 433
BsrI ACTGG 4 cut(s) 90, 877, 883, 1154
BssAI RCCGGY 2 cut(s) 142, 433
BssECI CCNNGG 6 cut(s) 39, 189, 326, 406, 861, 1113
BssMI GATC 1 cut(s) 445
BssT1I CCWWGG 2 cut(s) 861, 1113
Bst2UI CCWGG 2 cut(s) 41, 254
Bst4CI ACNGT 4 cut(s) 193, 810, 964, 1026
Bst6I CTCTTC 1 cut(s) 1092
BstC8I GCNNGC 4 cut(s) 144, 435, 955, 990
BstDEI CTNAG 8 cut(s) 210, 258, 423, 684, 690, 915, 1119, 1179
BstDSI CCRYGG 3 cut(s) 189, 326, 1113
BstEII GGTNACC 1 cut(s) 33
BstF5I GGATG 1 cut(s) 486
BstFNI CGCG 4 cut(s) 26, 176, 365, 927
BstH2I RGCGCY 1 cut(s) 289
BstHHI GCGC 3 cut(s) 288, 365, 1022
BstKTI GATC 1 cut(s) 448
BstMAI GTCTC 5 cut(s) 200, 542, 571, 634, 693
BstMBI GATC 1 cut(s) 445
BstMCI CGRYCG 2 cut(s) 369, 561
BstNI CCWGG 2 cut(s) 41, 254
BstNSI RCATGY 2 cut(s) 458, 992
BstPI GGTNACC 1 cut(s) 33
BstSCI CCNGG 5 cut(s) 39, 252, 368, 405, 489
BstSFI CTRYAG 1 cut(s) 462
BstUI CGCG 4 cut(s) 26, 176, 365, 927
BstV1I GCAGC 6 cut(s) 371, 393, 448, 829, 869, 1143
BstX2I RGATCY 1 cut(s) 445
BstYI RGATCY 1 cut(s) 445
BstZI CGGCCG 1 cut(s) 366
Bsu36I CCTNAGG 1 cut(s) 1179
BsuRI GGCC 7 cut(s) 368, 437, 476, 754, 781, 826, 1144
BtgI CCRYGG 3 cut(s) 189, 326, 1113
BtsCI GGATG 1 cut(s) 486
BtsI GCAGTG 1 cut(s) 785
BtsIMutI CAGTG 2 cut(s) 785, 1161
Cac8I GCNNGC 4 cut(s) 144, 435, 955, 990
CaiI CAGNNNCTG 1 cut(s) 655
CfoI GCGC 3 cut(s) 288, 365, 1022
Cfr10I RCCGGY 2 cut(s) 142, 433
Cfr13I GGNCC 3 cut(s) 721, 752, 824
Csp6I GTAC 2 cut(s) 580, 876
CviAII CATG 5 cut(s) 455, 989, 1114, 1135, 1163
CviQI GTAC 2 cut(s) 580, 876
DdeI CTNAG 8 cut(s) 210, 258, 423, 684, 690, 915, 1119, 1179
DpnI GATC 1 cut(s) 447
DpnII GATC 1 cut(s) 445
EaeI YGGCCR 3 cut(s) 366, 435, 474
EagI CGGCCG 1 cut(s) 366
Eam1104I CTCTTC 1 cut(s) 1092
EarI CTCTTC 1 cut(s) 1092
EciI GGCGGA 1 cut(s) 405
Ecl136II GAGCTC 1 cut(s) 108
EclXI CGGCCG 1 cut(s) 366
Eco130I CCWWGG 2 cut(s) 861, 1113
Eco24I GRGCYC 2 cut(s) 110, 910
Eco31I GGTCTC 1 cut(s) 542
Eco47I GGWCC 1 cut(s) 721
Eco47III AGCGCT 1 cut(s) 287
Eco52I CGGCCG 1 cut(s) 366
Eco53kI GAGCTC 1 cut(s) 108
Eco57I CTGAAG 1 cut(s) 677
Eco81I CCTNAGG 1 cut(s) 1179
Eco88I CYCGRG 1 cut(s) 400
Eco91I GGTNACC 1 cut(s) 33
EcoICRI GAGCTC 1 cut(s) 108
EcoO65I GGTNACC 1 cut(s) 33
EcoRII CCWGG 2 cut(s) 39, 252
EcoT14I CCWWGG 2 cut(s) 861, 1113
EcoT22I ATGCAT 1 cut(s) 994
EcoT38I GRGCYC 2 cut(s) 110, 910
ErhI CCWWGG 2 cut(s) 861, 1113
FaeI CATG 5 cut(s) 458, 992, 1117, 1138, 1166
FaiI YATR 9 cut(s) 291, 456, 503, 990, 1044, 1077, 1115, 1136, 1164
FaqI GGGAC 1 cut(s) 956
FatI CATG 5 cut(s) 454, 988, 1113, 1134, 1162
FauI CCCGC 1 cut(s) 902
FblI GTMKAC 1 cut(s) 869
FokI GGATG 1 cut(s) 473
FriOI GRGCYC 2 cut(s) 110, 910
FspAI RTGCGCAY 1 cut(s) 1021
FspBI CTAG 3 cut(s) 828, 954, 1195
FspI TGCGCA 1 cut(s) 1021
GlaI GCGC 3 cut(s) 287, 364, 1021
GsaI CCCAGC 2 cut(s) 914, 977
GsuI CTGGAG 2 cut(s) 73, 572
HaeII RGCGCY 1 cut(s) 289
HaeIII GGCC 7 cut(s) 368, 437, 476, 754, 781, 826, 1144
HapII CCGG 7 cut(s) 97, 143, 369, 406, 434, 490, 557
HhaI GCGC 3 cut(s) 288, 365, 1022
Hin1II CATG 5 cut(s) 458, 992, 1117, 1138, 1166
Hin6I GCGC 3 cut(s) 286, 363, 1020
HinP1I GCGC 3 cut(s) 286, 363, 1020
HpaII CCGG 7 cut(s) 97, 143, 369, 406, 434, 490, 557
HphI GGTGA 1 cut(s) 336
Hpy166II GTNNAC 3 cut(s) 870, 1159, 1188
Hpy188III TCNNGA 5 cut(s) 269, 724, 736, 1070, 1181
Hpy8I GTNNAC 3 cut(s) 870, 1159, 1188
Hpy99I CGWCG 4 cut(s) 221, 275, 775, 854
HpyCH4III ACNGT 4 cut(s) 193, 810, 964, 1026
HpyCH4IV ACGT 1 cut(s) 1005
HpyCH4V TGCA 6 cut(s) 381, 464, 625, 790, 988, 992
HpyF3I CTNAG 8 cut(s) 210, 258, 423, 684, 690, 915, 1119, 1179
HpySE526I ACGT 1 cut(s) 1005
Hsp92II CATG 5 cut(s) 458, 992, 1117, 1138, 1166
HspAI GCGC 3 cut(s) 286, 363, 1020
KroI GCCGGC 2 cut(s) 142, 433
KroNI GCCGGC 2 cut(s) 144, 435
Kzo9I GATC 1 cut(s) 445
LguI GCTCTTC 1 cut(s) 1092
LmnI GCTCC 6 cut(s) 92, 311, 323, 544, 854, 905
Lsp1109I GCAGC 6 cut(s) 371, 393, 448, 829, 869, 1143
LweI GCATC 3 cut(s) 118, 495, 537
MaeI CTAG 3 cut(s) 828, 954, 1195
MaeII ACGT 1 cut(s) 1005
MaeIII GTNAC 2 cut(s) 33, 1026
MalI GATC 1 cut(s) 447
MbiI CCGCTC 1 cut(s) 634
MboI GATC 1 cut(s) 445
MboII GAAGA 5 cut(s) 112, 506, 670, 1071, 1079
MflI RGATCY 1 cut(s) 445
MhlI GDGCHC 3 cut(s) 110, 541, 910
MlsI TGGCCA 1 cut(s) 476
MluCI AATT 1 cut(s) 1107
MluNI TGGCCA 1 cut(s) 476
MlyI GAGTC 7 cut(s) 50, 128, 206, 238, 287, 349, 875
MmeI TCCRAC 2 cut(s) 733, 832
Mox20I TGGCCA 1 cut(s) 476
Mph1103I ATGCAT 1 cut(s) 994
MroNI GCCGGC 2 cut(s) 142, 433
MscI TGGCCA 1 cut(s) 476
Msp20I TGGCCA 1 cut(s) 476
MspA1I CMGCKG 2 cut(s) 384, 914
MspI CCGG 7 cut(s) 97, 143, 369, 406, 434, 490, 557
MspR9I CCNGG 5 cut(s) 41, 254, 370, 407, 491
MvaI CCWGG 2 cut(s) 41, 254
MvnI CGCG 4 cut(s) 26, 176, 365, 927
NaeI GCCGGC 2 cut(s) 144, 435
NciI CCSGG 3 cut(s) 370, 407, 491
NcoI CCATGG 1 cut(s) 1113
NdeII GATC 1 cut(s) 445
NgoMIV GCCGGC 2 cut(s) 142, 433
NheI GCTAGC 1 cut(s) 953
NlaIII CATG 5 cut(s) 458, 992, 1117, 1138, 1166
NlaIV GGNNCC 4 cut(s) 88, 307, 907, 1152
NmeAIII GCCGAG 1 cut(s) 650
NmuCI GTSAC 1 cut(s) 1026
NsbI TGCGCA 1 cut(s) 1021
NsiI ATGCAT 1 cut(s) 994
NspI RCATGY 2 cut(s) 458, 992
PaeI GCATGC 1 cut(s) 992
PaeR7I CTCGAG 1 cut(s) 400
PciSI GCTCTTC 1 cut(s) 1092
PdiI GCCGGC 2 cut(s) 144, 435
PfeI GAWTC 5 cut(s) 46, 652, 705, 831, 1126
PflMI CCANNNNNTGG 1 cut(s) 327
PfoI TCCNGGA 1 cut(s) 489
PleI GAGTC 7 cut(s) 50, 128, 205, 238, 287, 349, 874
PpsI GAGTC 7 cut(s) 50, 128, 205, 238, 287, 349, 874
Psp124BI GAGCTC 1 cut(s) 110
Psp1406I AACGTT 1 cut(s) 1005
Psp6I CCWGG 2 cut(s) 39, 252
PspEI GGTNACC 1 cut(s) 33
PspFI CCCAGC 2 cut(s) 910, 973
PspGI CCWGG 2 cut(s) 39, 252
PspN4I GGNNCC 4 cut(s) 88, 307, 907, 1152
PspPI GGNCC 3 cut(s) 721, 752, 824
PspXI VCTCGAGB 1 cut(s) 400
PstI CTGCAG 1 cut(s) 466
PstNI CAGNNNCTG 1 cut(s) 655
PsuI RGATCY 1 cut(s) 445
PvuII CAGCTG 2 cut(s) 384, 914
RsaI GTAC 2 cut(s) 581, 877
RsaNI GTAC 2 cut(s) 580, 876
SacI GAGCTC 1 cut(s) 110
SapI GCTCTTC 1 cut(s) 1092
Sau3AI GATC 1 cut(s) 445
Sau96I GGNCC 3 cut(s) 721, 752, 824
SchI GAGTC 7 cut(s) 50, 128, 206, 238, 287, 349, 875
ScrFI CCNGG 5 cut(s) 41, 254, 370, 407, 491
SduI GDGCHC 3 cut(s) 110, 541, 910
SfaNI GCATC 3 cut(s) 118, 495, 537
SfcI CTRYAG 1 cut(s) 462
Sfr274I CTCGAG 1 cut(s) 400
SinI GGWCC 1 cut(s) 721
SlaI CTCGAG 1 cut(s) 400
SmlI CTYRAG 2 cut(s) 400, 1035
SmoI CTYRAG 2 cut(s) 400, 1035
SphI GCATGC 1 cut(s) 992
Sse9I AATT 1 cut(s) 1107
SspMI CTAG 3 cut(s) 828, 954, 1195
SstI GAGCTC 1 cut(s) 110
StyD4I CCNGG 5 cut(s) 39, 252, 368, 405, 489
StyI CCWWGG 2 cut(s) 861, 1113
TaaI ACNGT 4 cut(s) 193, 810, 964, 1026
TaiI ACGT 1 cut(s) 1008
TaqI TCGA 5 cut(s) 15, 159, 270, 353, 401
TaqII GACCGA 1 cut(s) 68
TasI AATT 1 cut(s) 1107
TauI GCSGC 7 cut(s) 26, 368, 390, 433, 461, 677, 930
TfiI GAWTC 5 cut(s) 46, 652, 705, 831, 1126
TscAI CASTG 2 cut(s) 792, 1161
TseFI GTSAC 1 cut(s) 1026
TseI GCWGC 6 cut(s) 381, 384, 461, 842, 857, 1131
Tsp45I GTSAC 1 cut(s) 1026
TspDTI ATGAA 1 cut(s) 1092
TspGWI ACGGA 1 cut(s) 1071
TspRI CASTG 2 cut(s) 792, 1161
Van91I CCANNNNNTGG 1 cut(s) 327
VpaK11BI GGWCC 1 cut(s) 721
XceI RCATGY 2 cut(s) 458, 992
XhoI CTCGAG 1 cut(s) 400
XmiI GTMKAC 1 cut(s) 869
XspI CTAG 3 cut(s) 828, 954, 1195
Zsp2I ATGCAT 1 cut(s) 994
Using CommOnly database (standard laboratory enzymes). Scanned on CDS sequence.