Basic Information

Type: gene

Biological Identity

Species

pyrus_communis

Chromosome

Chr1

Orthogroup ID

OG0006055

Physical Location & Seq

Genomic Coordinates Forward (+)

15755994 .. 15757483

Gene Structure 1490 bp

Loading structure...

UTR

Exon/CDS

Intron

Transcript / Protein ID

pycom01g16040.4

Sequence Viewer

CDS Protein Promoter (2k)

Length: 1086 bp

ATGAAGGCACTTATTCTTGTTGGAGGTTTCGGGACGCGTCTGCGGCCATTGACACTCAGTTTCCCCAAGCCACTTGTTGAGTTTGCTAACAAACCCATGATACTGCATCAGATAGAGGCTCTTAAGGCAATTGGTGTGACCGAAGTGGTTCTGGCTATCAATTACCAACCAGAGGTGATGATGACTTTCTTGAAGGAGTTTGAGGAAAAGGTTGGCATCAAGATCACATGCTCACAAGAGACCGAACCCCTTGGCACTGCTGGGCCTCTGGCTCTTGCTAGGGACAAACTGATTGATGATTCTGGCGAGCCCTTCTTTGTCCTTAACAGTGATGTTATCAGCGAGTACCCGTTTAAGCTAATGATTGAATTCCATAAAGGCCATGGAGGAGAAGCTTCCATAATGGTGACCAAGGTGGATGAGCCATCGAAATATGGTGTGGTGGTTATGGAAGAGTCTACAGGGAAAGTTCAGAAATTTGTAGAGAAACCAAAGTTGTTTGTTGGTAACAAAATCAATGCTGGAATTTACCTGTTGAACCCCTCCGTTCTTGATAAAATTGAACTCAGACCAACCTCAATTGAGAAAGAAGTATTCCCGAAAATCGCAGCAGAGAATAAGCTCTTTGCAATGGTCCTTCCAGGGTTTTGGATGGACATTGGGCAACCAAGGGATTATATTACCGGCCTGAGGCTATACTTGGATTCGTTGAGGAAGAACTCTTCATCCAAGTTGGCCGTAGGCGCTCACATTGTGGGAAACGTTCTGGTGGATGAGACTGCCAAAATTGGCGTGGGGTGCCTGATTGGACCGGATGTTGCAATCGGTCCAGGCTGCGTTATCGAGTCAGGAGTCAGACTCTCTCGCTGCACGGTAATGCGTGGAGTCCGCATCAAGAACCACGCTTGCATTTCCAGCAGTATCATCGGATGGCACTCCACCGTTGGACAATGGGCTCGCGTAGAGAACATGACCATCCTTGGAGAAGATGTGCACGTGAGCGATGAGATTTACAGCAACGGAGGTGTGGTATTGCCACACAAAGAAATCAAGTCGAGCATTTTGAAGCCGGAAATTGTAATGTAG

Functional Annotation

Gene Ontology

Molecular Function
Biological Process
Cellular Component

GO:0000271 GO:0000302 GO:0001101 GO:0003674 GO:0003824 GO:0004475 GO:0005575 GO:0005622 GO:0005623 GO:0005634 GO:0005737 GO:0005829 GO:0005975 GO:0005976 GO:0005996 GO:0006073 GO:0006082 GO:0006732 GO:0006766 GO:0006767 GO:0006950 GO:0006952 GO:0006970 GO:0006979 GO:0008150 GO:0008152 GO:0008905 GO:0009058 GO:0009059 GO:0009108 GO:0009110 GO:0009250 GO:0009266 GO:0009408 GO:0009605 GO:0009607 GO:0009617 GO:0009628 GO:0009651 GO:0009719 GO:0009725 GO:0009753 GO:0009987 GO:0010033 GO:0010035 GO:0010193 GO:0016051 GO:0016053 GO:0016740 GO:0016772 GO:0016779 GO:0019752 GO:0019852 GO:0019853 GO:0030243 GO:0030244 GO:0033692 GO:0034637 GO:0034645 GO:0042221 GO:0042364 GO:0042493 GO:0042742 GO:0043170 GO:0043207 GO:0043226 GO:0043227 GO:0043229 GO:0043231 GO:0043436 GO:0044042 GO:0044237 GO:0044238 GO:0044249 GO:0044260 GO:0044262 GO:0044264 GO:0044281 GO:0044283 GO:0044424 GO:0044444 GO:0044464 GO:0046364 GO:0046394 GO:0046677 GO:0050896 GO:0051186 GO:0051188 GO:0051273 GO:0051274 GO:0051704 GO:0051707 GO:0060359 GO:0070568 GO:0071704 GO:0098542 GO:1901576 GO:1901698 GO:1901700

KEGG Pathways

Metabolic & Signaling

ko00051 ko00520 ko01100 ko01110 map00051 map00520 map01100 map01110

Pfam Domains

Protein Families

Hexapep NTP_transferase

Protein Analysis

362

Amino Acids

39.54

Weight (kDa)

6.73

Isoelectric Point (pI)

35.74

Instability Index

Protein Domains (Pfam)

No Pfam domains detected for this protein.

Hydrophobicity Profile (Kyte-Doolittle)

AI Structure Prediction Report

Calculating structure properties...

Genomic Context

Gene Family Tree

Species Filter

Style Settings

Font Size (px)

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Align Labels (Cladogram)

Image

Tree File

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Publication-ready

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Orthologous Genes (Group: OG0006055)

Species	Orthologous Gene IDs
arabidopsis_thaliana	AT2G39770 AT2G39770 AT2G39770 AT3G55590
fragaria_vesca	FvH4_7g23380 FvH4_7g23380 FvH4_7g23380 FvH4_7g23380 FvH4_7g23380
malus_domestica	MD01G1139900.v1.1 MD07G1204200.v1.1
prunus_persica	Prupe.2G240500_v2.0.a1
pyrus_communis	pycom01g16040 pycom07g19080
rosa_chinensis	RchiOBHm_Chr1g0368511
rosa_laevigata	RLG00000027207
rosa_multiflora	Rmu_co8389653.1_g000001 Rmu_sc0006908.1_g000002 Rmu_sc0011394.1_g000010
rosa_roxburghii	Rroxscaffold_4G00288480
rosa_rugosa	Rorug01G0342400 Rorug01G0342500 Rorug01G0342500
rosa_samantha	Rh1AG349800 Rh1BG312900 Rh1CG327700 Rh1DG344400

Restriction Enzyme Sites

1 / 10

Enzyme	Recognition Site	Cut Count	Positions (bp)
AccB1I	GGYRCC	1 cut(s)	798
AccI	GTMKAC	1 cut(s)	458
AccII	CGCG	2 cut(s)	37, 960
AciI	CCGC	2 cut(s)	43, 889
AclI	AACGTT	1 cut(s)	762
AcoI	YGGCCR	2 cut(s)	44, 735
AcsI	RAATTY	3 cut(s)	368, 476, 525
AcvI	CACGTG	1 cut(s)	997
AdeI	CACNNNGTG	1 cut(s)	754
AfaI	GTAC	1 cut(s)	347
AfiI	CCNNNNNNNGG	2 cut(s)	172, 690
AflII	CTTAAG	1 cut(s)	122
AflIII	ACRYGT	1 cut(s)	35
AgsI	TTSAA	5 cut(s)	193, 368, 538, 563, 1066
AjnI	CCWGG	2 cut(s)	640, 829
AloI	GAACNNNNNNTCC	2 cut(s)	710, 742
AluBI	AGCT	3 cut(s)	358, 395, 622
AluI	AGCT	3 cut(s)	358, 395, 622
Alw21I	GWGCWC	1 cut(s)	996
Alw26I	GTCTC	2 cut(s)	233, 770
Alw44I	GTGCAC	1 cut(s)	992
AoxI	GGCC	5 cut(s)	44, 263, 379, 685, 735
ApaLI	GTGCAC	1 cut(s)	992
ApeKI	GCWGC	3 cut(s)	608, 834, 867
ApoI	RAATTY	3 cut(s)	368, 476, 525
Asp700I	GAANNNNTTC	1 cut(s)	147
AspLEI	GCGC	1 cut(s)	746
AspS9I	GGNCC	4 cut(s)	263, 634, 809, 827
AsuHPI	GGTGA	2 cut(s)	187, 418
AvaII	GGWCC	3 cut(s)	634, 809, 827
AxyI	CCTNAGG	1 cut(s)	689
BaeGI	GKGCMC	1 cut(s)	996
BanI	GGYRCC	1 cut(s)	798
BanII	GRGCYC	2 cut(s)	312, 958
BbrPI	CACGTG	1 cut(s)	997
Bbv12I	GWGCWC	1 cut(s)	996
BbvI	GCAGC	3 cut(s)	620, 821, 854
BccI	CCATC	4 cut(s)	433, 646, 924, 983
BceAI	ACGGC	1 cut(s)	722
BcgI	CGANNNNNNTGC	2 cut(s)	907, 941
BciT130I	CCWGG	2 cut(s)	642, 831
BcoDI	GTCTC	2 cut(s)	233, 770
BfaI	CTAG	1 cut(s)	279
BfmI	CTRYAG	1 cut(s)	459
BfoI	RGCGCY	1 cut(s)	747
BfrI	CTTAAG	1 cut(s)	122
BisI	GCNGC	4 cut(s)	44, 609, 835, 868
BlsI	GCNGC	4 cut(s)	45, 610, 836, 869
Bme1390I	CCNGG	2 cut(s)	642, 831
Bme18I	GGWCC	3 cut(s)	634, 809, 827
BmgT120I	GGNCC	4 cut(s)	263, 634, 809, 827
BmiI	GGNNCC	1 cut(s)	800
BmrFI	CCNGG	2 cut(s)	642, 831
BmsI	GCATC	3 cut(s)	115, 225, 900
BplI	GAGNNNNNCTC	2 cut(s)	843, 875
BsaAI	YACGTR	1 cut(s)	997
BsaI	GGTCTC	1 cut(s)	233
BsaJI	CCNNGG	6 cut(s)	250, 382, 411, 641, 668, 979
BsaWI	WCCGGW	1 cut(s)	811
BsaXI	ACNNNNNCTCC	4 cut(s)	975, 1005, 1014, 1044
Bsc4I	CCNNNNNNNGG	2 cut(s)	172, 690
Bse118I	RCCGGY	1 cut(s)	683
Bse21I	CCTNAGG	1 cut(s)	689
Bse3DI	GCAATG	1 cut(s)	636
BseBI	CCWGG	2 cut(s)	642, 831
BseDI	CCNNGG	6 cut(s)	250, 382, 411, 641, 668, 979
BseGI	GGATG	7 cut(s)	424, 657, 725, 778, 820, 935, 975
BseLI	CCNNNNNNNGG	2 cut(s)	172, 690
BseMI	GCAATG	1 cut(s)	636
BseMII	CTCAG	3 cut(s)	70, 580, 680
BseRI	GAGGAG	1 cut(s)	402
BseSI	GKGCMC	1 cut(s)	996
BseXI	GCAGC	3 cut(s)	620, 821, 854
BseYI	CCCAGC	1 cut(s)	260
BsgI	GTGCAG	1 cut(s)	853
Bsh1236I	CGCG	2 cut(s)	37, 960
BshFI	GGCC	5 cut(s)	46, 265, 381, 687, 737
BshNI	GGYRCC	1 cut(s)	798
BsiHKAI	GWGCWC	1 cut(s)	996
BsiSI	CCGG	3 cut(s)	684, 812, 1070
BslFI	GGGAC	2 cut(s)	46, 296
BslI	CCNNNNNNNGG	2 cut(s)	172, 690
BsmAI	GTCTC	2 cut(s)	233, 770
BsmFI	GGGAC	2 cut(s)	46, 296
BsnI	GGCC	5 cut(s)	46, 265, 381, 687, 737
Bso31I	GGTCTC	1 cut(s)	233
Bsp1286I	GDGCHC	3 cut(s)	312, 958, 996
Bsp143I	GATC	1 cut(s)	222
Bsp19I	CCATGG	1 cut(s)	382
BspACI	CCGC	2 cut(s)	43, 889
BspANI	GGCC	5 cut(s)	46, 265, 381, 687, 737
BspCNI	CTCAG	3 cut(s)	69, 579, 681
BspFNI	CGCG	2 cut(s)	37, 960
BspLI	GGNNCC	1 cut(s)	800
BspT107I	GGYRCC	1 cut(s)	798
BspTI	CTTAAG	1 cut(s)	122
BspTNI	GGTCTC	1 cut(s)	233
BsrDI	GCAATG	1 cut(s)	636
BsrFI	RCCGGY	1 cut(s)	683
BssAI	RCCGGY	1 cut(s)	683
BssECI	CCNNGG	6 cut(s)	250, 382, 411, 641, 668, 979
BssMI	GATC	1 cut(s)	222
BssT1I	CCWWGG	5 cut(s)	250, 382, 411, 668, 979
Bst2UI	CCWGG	2 cut(s)	642, 831
Bst4CI	ACNGT	3 cut(s)	329, 874, 943
Bst6I	CTCTTC	2 cut(s)	447, 727
BstAFI	CTTAAG	1 cut(s)	122
BstBAI	YACGTR	1 cut(s)	997
BstC8I	GCNNGC	3 cut(s)	308, 907, 958
BstDEI	CTNAG	3 cut(s)	56, 566, 689
BstDSI	CCRYGG	1 cut(s)	382
BstEII	GGTNACC	1 cut(s)	406
BstF5I	GGATG	7 cut(s)	424, 657, 725, 778, 820, 935, 975
BstFNI	CGCG	2 cut(s)	37, 960
BstH2I	RGCGCY	1 cut(s)	747
BstHHI	GCGC	1 cut(s)	746
BstKTI	GATC	1 cut(s)	225
BstMAI	GTCTC	2 cut(s)	233, 770
BstMBI	GATC	1 cut(s)	222
BstMWI	GCNNNNNNNGC	5 cut(s)	43, 125, 743, 798, 915
BstNI	CCWGG	2 cut(s)	642, 831
BstNSI	RCATGY	1 cut(s)	231
BstPI	GGTNACC	1 cut(s)	406
BstSCI	CCNGG	2 cut(s)	640, 829
BstSFI	CTRYAG	1 cut(s)	459
BstSLI	GKGCMC	1 cut(s)	996
BstUI	CGCG	2 cut(s)	37, 960
BstV1I	GCAGC	3 cut(s)	620, 821, 854
BstXI	CCANNNNNNTGG	1 cut(s)	648
Bsu36I	CCTNAGG	1 cut(s)	689
BsuRI	GGCC	5 cut(s)	46, 265, 381, 687, 737
BtgI	CCRYGG	1 cut(s)	382
BtgZI	GCGATG	1 cut(s)	1017
BtsCI	GGATG	7 cut(s)	424, 657, 725, 778, 820, 935, 975
BtsI	GCAGTG	1 cut(s)	255
BtsIMutI	CAGTG	2 cut(s)	255, 334
Cac8I	GCNNGC	3 cut(s)	308, 907, 958
CfoI	GCGC	1 cut(s)	746
Cfr10I	RCCGGY	1 cut(s)	683
Cfr13I	GGNCC	4 cut(s)	263, 634, 809, 827
CseI	GACGC	2 cut(s)	26, 43
Csp6I	GTAC	1 cut(s)	346
CviAII	CATG	4 cut(s)	97, 228, 383, 970
CviQI	GTAC	1 cut(s)	346
DdeI	CTNAG	3 cut(s)	56, 566, 689
DpnI	GATC	1 cut(s)	224
DpnII	GATC	1 cut(s)	222
DraIII	CACNNNGTG	1 cut(s)	754
EaeI	YGGCCR	2 cut(s)	44, 735
Eam1104I	CTCTTC	2 cut(s)	447, 727
EarI	CTCTTC	2 cut(s)	447, 727
Eco130I	CCWWGG	5 cut(s)	250, 382, 411, 668, 979
Eco24I	GRGCYC	2 cut(s)	312, 958
Eco31I	GGTCTC	1 cut(s)	233
Eco47I	GGWCC	3 cut(s)	634, 809, 827
Eco72I	CACGTG	1 cut(s)	997
Eco81I	CCTNAGG	1 cut(s)	689
Eco91I	GGTNACC	1 cut(s)	406
EcoO65I	GGTNACC	1 cut(s)	406
EcoRI	GAATTC	1 cut(s)	368
EcoRII	CCWGG	2 cut(s)	640, 829
EcoT14I	CCWWGG	5 cut(s)	250, 382, 411, 668, 979
EcoT38I	GRGCYC	2 cut(s)	312, 958
ErhI	CCWWGG	5 cut(s)	250, 382, 411, 668, 979
FaeI	CATG	4 cut(s)	100, 231, 386, 973
FaqI	GGGAC	2 cut(s)	46, 296
FatI	CATG	4 cut(s)	96, 227, 382, 969
FblI	GTMKAC	1 cut(s)	458
Fnu4HI	GCNGC	4 cut(s)	44, 609, 835, 868
FokI	GGATG	7 cut(s)	431, 664, 712, 785, 827, 942, 962
FriOI	GRGCYC	2 cut(s)	312, 958
Fsp4HI	GCNGC	4 cut(s)	44, 609, 835, 868
FspBI	CTAG	1 cut(s)	279
GlaI	GCGC	1 cut(s)	745
GluI	GCNGC	4 cut(s)	44, 609, 835, 868
GsaI	CCCAGC	1 cut(s)	264
HaeII	RGCGCY	1 cut(s)	747
HaeIII	GGCC	5 cut(s)	46, 265, 381, 687, 737
HapII	CCGG	3 cut(s)	684, 812, 1070
HgaI	GACGC	2 cut(s)	26, 43
HhaI	GCGC	1 cut(s)	746
Hin1II	CATG	4 cut(s)	100, 231, 386, 973
Hin6I	GCGC	1 cut(s)	744
HinP1I	GCGC	1 cut(s)	744
HindIII	AAGCTT	1 cut(s)	393
HinfI	GANTC	7 cut(s)	299, 455, 704, 845, 852, 858, 885
HpaII	CCGG	3 cut(s)	684, 812, 1070
HphI	GGTGA	2 cut(s)	187, 418
Hpy166II	GTNNAC	2 cut(s)	459, 994
Hpy188I	TCNGA	5 cut(s)	111, 474, 569, 857, 929
Hpy188III	TCNNGA	7 cut(s)	31, 190, 220, 551, 598, 849, 895
Hpy8I	GTNNAC	2 cut(s)	459, 994
HpyAV	CCTTC	3 cut(s)	187, 322, 647
HpyCH4III	ACNGT	3 cut(s)	329, 874, 943
HpyCH4IV	ACGT	2 cut(s)	762, 996
HpyCH4V	TGCA	6 cut(s)	106, 629, 821, 870, 909, 994
HpyF10VI	GCNNNNNNNGC	5 cut(s)	43, 125, 743, 798, 915
HpyF3I	CTNAG	3 cut(s)	56, 566, 689
HpySE526I	ACGT	2 cut(s)	762, 996
Hsp92II	CATG	4 cut(s)	100, 231, 386, 973
HspAI	GCGC	1 cut(s)	744
Kzo9I	GATC	1 cut(s)	222
Lsp1109I	GCAGC	3 cut(s)	620, 821, 854
LweI	GCATC	3 cut(s)	115, 225, 900
MaeI	CTAG	1 cut(s)	279
MaeII	ACGT	2 cut(s)	762, 996
MaeIII	GTNAC	3 cut(s)	136, 406, 506
MalI	GATC	1 cut(s)	224
MboI	GATC	1 cut(s)	222
MboII	GAAGA	4 cut(s)	464, 714, 727, 998
MfeI	CAATTG	2 cut(s)	129, 579
MhlI	GDGCHC	3 cut(s)	312, 958, 996
MluCI	AATT	9 cut(s)	129, 160, 368, 476, 525, 558, 579, 786, 1074
MluI	ACGCGT	1 cut(s)	35
MlyI	GAGTC	5 cut(s)	464, 852, 854, 861, 894
MmeI	TCCRAC	1 cut(s)	925
MroXI	GAANNNNTTC	1 cut(s)	147
MseI	TTAA	3 cut(s)	123, 324, 354
MslI	CAYNNNNRTG	2 cut(s)	404, 875
MspCI	CTTAAG	1 cut(s)	122
MspI	CCGG	3 cut(s)	684, 812, 1070
MspR9I	CCNGG	2 cut(s)	642, 831
MunI	CAATTG	2 cut(s)	129, 579
MvaI	CCWGG	2 cut(s)	642, 831
MvnI	CGCG	2 cut(s)	37, 960
MwoI	GCNNNNNNNGC	5 cut(s)	43, 125, 743, 798, 915
NcoI	CCATGG	1 cut(s)	382
NdeII	GATC	1 cut(s)	222
NlaIII	CATG	4 cut(s)	100, 231, 386, 973
NlaIV	GGNNCC	1 cut(s)	800
NmuCI	GTSAC	2 cut(s)	136, 406
NspI	RCATGY	1 cut(s)	231
PdmI	GAANNNNTTC	1 cut(s)	147
PfeI	GAWTC	2 cut(s)	299, 704
PkrI	GCNGC	4 cut(s)	45, 610, 836, 869
PleI	GAGTC	5 cut(s)	463, 852, 853, 860, 893
PmaCI	CACGTG	1 cut(s)	997
PmlI	CACGTG	1 cut(s)	997
PpsI	GAGTC	5 cut(s)	463, 852, 853, 860, 893
Ppu21I	YACGTR	1 cut(s)	997
Psp1406I	AACGTT	1 cut(s)	762
Psp6I	CCWGG	2 cut(s)	640, 829
PspCI	CACGTG	1 cut(s)	997
PspEI	GGTNACC	1 cut(s)	406
PspFI	CCCAGC	1 cut(s)	260
PspGI	CCWGG	2 cut(s)	640, 829
PspN4I	GGNNCC	1 cut(s)	800
PspPI	GGNCC	4 cut(s)	263, 634, 809, 827
RsaI	GTAC	1 cut(s)	347
RsaNI	GTAC	1 cut(s)	346
RseI	CAYNNNNRTG	2 cut(s)	404, 875
SaqAI	TTAA	3 cut(s)	123, 324, 354
SatI	GCNGC	4 cut(s)	44, 609, 835, 868
Sau3AI	GATC	1 cut(s)	222
Sau96I	GGNCC	4 cut(s)	263, 634, 809, 827
SchI	GAGTC	5 cut(s)	464, 852, 854, 861, 894
ScrFI	CCNGG	2 cut(s)	642, 831
SduI	GDGCHC	3 cut(s)	312, 958, 996
SfaNI	GCATC	3 cut(s)	115, 225, 900
SfcI	CTRYAG	1 cut(s)	459
SinI	GGWCC	3 cut(s)	634, 809, 827
SmiMI	CAYNNNNRTG	2 cut(s)	404, 875
SmlI	CTYRAG	1 cut(s)	122
SmoI	CTYRAG	1 cut(s)	122
Sse9I	AATT	9 cut(s)	129, 160, 368, 476, 525, 558, 579, 786, 1074
SsiI	CCGC	2 cut(s)	43, 889
SspMI	CTAG	1 cut(s)	279
StyD4I	CCNGG	2 cut(s)	640, 829
StyI	CCWWGG	5 cut(s)	250, 382, 411, 668, 979
TaaI	ACNGT	3 cut(s)	329, 874, 943
TaiI	ACGT	2 cut(s)	765, 999
TaqI	TCGA	3 cut(s)	428, 843, 1055
TaqII	GACCGA	3 cut(s)	155, 257, 815
TasI	AATT	9 cut(s)	129, 160, 368, 476, 525, 558, 579, 786, 1074
TauI	GCSGC	1 cut(s)	46
TfiI	GAWTC	2 cut(s)	299, 704
Tru1I	TTAA	3 cut(s)	123, 324, 354
Tru9I	TTAA	3 cut(s)	123, 324, 354
TscAI	CASTG	2 cut(s)	262, 334
TseFI	GTSAC	2 cut(s)	136, 406
TseI	GCWGC	3 cut(s)	608, 834, 867
Tsp45I	GTSAC	2 cut(s)	136, 406
TspDTI	ATGAA	2 cut(s)	17, 714
TspGWI	ACGGA	2 cut(s)	535, 1035
TspRI	CASTG	2 cut(s)	262, 334
Vha464I	CTTAAG	1 cut(s)	122
VneI	GTGCAC	1 cut(s)	992
VpaK11BI	GGWCC	3 cut(s)	634, 809, 827
XapI	RAATTY	3 cut(s)	368, 476, 525
XceI	RCATGY	1 cut(s)	231
XcmI	CCANNNNNNNNNTGG	2 cut(s)	380, 790
XmiI	GTMKAC	1 cut(s)	458
XmnI	GAANNNNTTC	1 cut(s)	147
XspI	CTAG	1 cut(s)	279

Using CommOnly database (standard laboratory enzymes). Scanned on CDS sequence.