Basic Information

Type: gene

Biological Identity

Species

rosa_multiflora

Chromosome

Rmu_sc0004741.1

Orthogroup ID

OG0004462

Physical Location & Seq

Genomic Coordinates Reverse (-)

34168 .. 37153

Gene Structure 2986 bp

Loading structure...

UTR

Exon/CDS

Intron

Transcript / Protein ID

Rmu_sc0004741.1_g000007.1.cds

Sequence Viewer

CDS Protein Promoter (2k)

Length: 1398 bp

atggtagcttctgatgagtctggaggccttggttatgaccacaattatgatagtaatgatgctaaacctcaaccaccaccatcacagcacgaaaatggggaatcatcaaaacagaattcgatattctcacttactcttgatgaaattcaaagcaagagtgggaagagttttgggtccatgaacatggatgaattccttgccaacatatggagtgctgatgaaaatcaagtttcttcagaacaacccaaccaaggtgatctggatgaccctgccaatgggaaagacacaacttcaaagcctgcctctctttcacggcagggatcattctccattcctaccccactctgcaagaaaactgtggatgaggtatggtcagagatcgacaaaactcgaccaccacccccgcctaacgacatgagtaatcttgatcatccccagagccaatcaaccataggagaaataactctggaagatttcctagtaaaagctggtgtagttcaagaatcaccttctcagaaccgaatggaaagtgacccaattcatccacaaggtggtgacgccaacaactctaatatggcaactagcagtagcatgtatgtgggtacaagttctaaaatggggaataatgtgattgcttcaggacatttgaatccccaaaatgctgtgccaaacaatctgtctggaaatagatatgcgattgctggatatccaatttttggacagaataaaatgattgtgggaaatgcatctttttgtaccagtaatgaaattgagaaacgtcacagtttgccagaatctagtgggacaaagaacaagaagaggatgattgatggtccactggaggtagcagtggagcggaggcaacgaaggatgatcaaaaatcgagagtcagcagctcggtctcgtgcaaggaaacaggcatacactgtggaactagaagctgaggtgaatcaacttgaagacgagaatgaaaagctgaagcaaacagtggagggaatcgaacagaggcgaaaacaagaggtaggtgatgggaaggaaacaatccacaaaagcgcaaaggttggcagcaaagttgaggacaatgaggaggacttggagggcggcgctgagaatttcagtggaatccatggcagccctgagattattgagggtctcctgatggtactcgtgggatccatggaagaagttgaagcgggccacgttcattccggcccagagaagcttctccaccatcggcaccgacctggatgcaggtcccagggtgcacaagatcttcttcttcggcttctgatctctctctctctctctcttcttcttcttcttcttcttcaatctgtgaattcatcctttcatccttcaatctgtgaattcgtccttgatgtgaagcacttgtgttaa

Functional Annotation

Gene Ontology

Molecular Function
Biological Process
Cellular Component

GO:0003674 GO:0003700 GO:0005488 GO:0005515 GO:0005575 GO:0005622 GO:0005623 GO:0005634 GO:0006355 GO:0008134 GO:0008150 GO:0009889 GO:0009891 GO:0009893 GO:0010468 GO:0010556 GO:0010557 GO:0010604 GO:0010628 GO:0019219 GO:0019222 GO:0031323 GO:0031325 GO:0031326 GO:0031328 GO:0033613 GO:0043226 GO:0043227 GO:0043229 GO:0043231 GO:0044424 GO:0044464 GO:0045893 GO:0045935 GO:0048518 GO:0048522 GO:0050789 GO:0050794 GO:0051171 GO:0051173 GO:0051252 GO:0051254 GO:0060255 GO:0065007 GO:0080090 GO:0140110 GO:1902680 GO:1903506 GO:1903508 GO:2000112 GO:2001141

KEGG Pathways

Metabolic & Signaling

ko04075 map04075

Pfam Domains

Protein Families

bZIP_1

Protein Analysis

465

Amino Acids

50.86

Weight (kDa)

5.29

Isoelectric Point (pI)

57.41

Instability Index

Protein Domains (Pfam)

No Pfam domains detected for this protein.

Hydrophobicity Profile (Kyte-Doolittle)

AI Structure Prediction Report

Calculating structure properties...

Genomic Context

Gene Family Tree

Species Filter

Style Settings

Font Size (px)

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Align Labels (Cladogram)

Image

Tree File

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Publication-ready

iTOL Interactive Tree Of Life · Most popular EvolView Flexible annotations MEGA Desktop phylogenetics suite FigTree Desktop tree editor

Orthologous Genes (Group: OG0004462)

Species	Orthologous Gene IDs
arabidopsis_thaliana	AT3G44460
fragaria_vesca	FvH4_3g24780 FvH4_3g24800
malus_domestica	MD03G1143300.v1.1
prunus_persica	Prupe.6G128900_v2.0.a1 Prupe.6G128900_v2.0.a1 Prupe.6G128900_v2.0.a1 Prupe.6G128900_v2.0.a1
rosa_chinensis	RchiOBHm_Chr5g0044361 RchiOBHm_Chr5g0044371
rosa_laevigata	RLG00000034268 RLG00000034269
rosa_multiflora	Rmu_sc0004741.1_g000007 Rmu_sc0004741.1_g000008
rosa_roxburghii	Rroxscaffold_1G00036580 Rroxscaffold_1G00036600
rosa_rugosa	Rorug05G0213700 Rorug05G0213700 Rorug05G0213700 Rorug05G0213700 Rorug05G0213700 Rorug05G0213700
rosa_samantha	Rh5AG297900 Rh5AG298100 Rh5BG305300 Rh5BG305500 Rh5CG332300 Rh5CG332500 Rh5DG314500 Rh5DG314700
rosa_wichuraiana	Rw5G027580 Rw5G027590

Restriction Enzyme Sites

1 / 10

Enzyme	Recognition Site	Cut Count	Positions (bp)
Acc36I	ACCTGC	1 cut(s)	1242
AccB1I	GGYRCC	1 cut(s)	1236
AccB7I	CCANNNNNTGG	3 cut(s)	207, 551, 716
AccBSI	CCGCTC	1 cut(s)	856
AciI	CCGC	4 cut(s)	404, 856, 1101, 1193
AclWI	GGATC	3 cut(s)	328, 1167, 1180
AcsI	RAATTY	6 cut(s)	115, 144, 191, 1111, 1339, 1367
AcuI	CTGAAG	3 cut(s)	219, 621, 998
AcyI	GRCGYC	1 cut(s)	558
AdeI	CACNNNGTG	1 cut(s)	551
AfaI	GTAC	3 cut(s)	604, 757, 1164
AfiI	CCNNNNNNNGG	5 cut(s)	207, 251, 275, 551, 716
AgsI	TTSAA	8 cut(s)	149, 294, 500, 649, 959, 1190, 1331, 1359
AjnI	CCWGG	2 cut(s)	1243, 1257
AluBI	AGCT	6 cut(s)	8, 488, 896, 941, 976, 1222
AluI	AGCT	6 cut(s)	8, 488, 896, 941, 976, 1222
Alw21I	GWGCWC	1 cut(s)	1267
Alw26I	GTCTC	2 cut(s)	906, 1157
Alw44I	GTGCAC	1 cut(s)	1263
AlwI	GGATC	3 cut(s)	328, 1167, 1180
AoxI	GGCC	3 cut(s)	25, 1195, 1210
ApaLI	GTGCAC	1 cut(s)	1263
ApeKI	GCWGC	3 cut(s)	893, 1065, 1131
ApoI	RAATTY	6 cut(s)	115, 144, 191, 1111, 1339, 1367
ArsI	GACNNNNNNTTYG	2 cut(s)	662, 694
AspLEI	GCGC	2 cut(s)	1055, 1106
AspS9I	GGNCC	5 cut(s)	174, 833, 1195, 1211, 1254
AsuHPI	GGTGA	5 cut(s)	266, 498, 566, 958, 1037
AvaII	GGWCC	3 cut(s)	174, 833, 1254
BaeGI	GKGCMC	1 cut(s)	1267
BamHI	GGATCC	1 cut(s)	1172
BanI	GGYRCC	1 cut(s)	1236
BauI	CACGAG	2 cut(s)	903, 1166
BbsI	GAAGAC	1 cut(s)	966
Bbv12I	GWGCWC	1 cut(s)	1267
BbvCI	CCTCAGC	1 cut(s)	942
BbvI	GCAGC	3 cut(s)	905, 1077, 1143
BccI	CCATC	5 cut(s)	88, 824, 1022, 1153, 1239
BceAI	ACGGC	1 cut(s)	329
BcgI	CGANNNNNNTGC	2 cut(s)	1230, 1264
BciT130I	CCWGG	2 cut(s)	1245, 1259
BclI	TGATCA	2 cut(s)	427, 873
BcoDI	GTCTC	2 cut(s)	906, 1157
BfaI	CTAG	4 cut(s)	479, 582, 798, 935
BfoI	RGCGCY	1 cut(s)	1107
BfuAI	ACCTGC	1 cut(s)	1242
BglII	AGATCT	1 cut(s)	1270
BisI	GCNGC	4 cut(s)	894, 1066, 1102, 1132
BlsI	GCNGC	4 cut(s)	895, 1067, 1103, 1133
Bme1390I	CCNGG	2 cut(s)	1245, 1259
Bme18I	GGWCC	3 cut(s)	174, 833, 1254
BmgT120I	GGNCC	5 cut(s)	174, 833, 1195, 1211, 1254
BmiI	GGNNCC	4 cut(s)	175, 1174, 1238, 1256
BmrFI	CCNGG	2 cut(s)	1245, 1259
BmsI	GCATC	3 cut(s)	49, 755, 1238
BpiI	GAAGAC	1 cut(s)	966
BpmI	CTGGAG	2 cut(s)	42, 860
Bpu10I	CCTNAGC	1 cut(s)	942
BsaBI	GATNNNNATC	1 cut(s)	222
BsaHI	GRCGYC	1 cut(s)	558
BsaI	GGTCTC	2 cut(s)	906, 1157
BsaJI	CCNNGG	6 cut(s)	28, 250, 1126, 1176, 1257, 1258
Bsc4I	CCNNNNNNNGG	5 cut(s)	207, 251, 275, 551, 716
Bse1I	ACTGG	2 cut(s)	759, 843
Bse8I	GATNNNNATC	1 cut(s)	222
BseBI	CCWGG	2 cut(s)	1245, 1259
BseDI	CCNNGG	6 cut(s)	28, 250, 1126, 1176, 1257, 1258
BseJI	GATNNNNATC	1 cut(s)	222
BseLI	CCNNNNNNNGG	5 cut(s)	207, 251, 275, 551, 716
BseMII	CTCAG	4 cut(s)	527, 933, 1098, 1128
BseNI	ACTGG	2 cut(s)	759, 843
BseRI	GAGGAG	1 cut(s)	1100
BseSI	GKGCMC	1 cut(s)	1267
BseXI	GCAGC	3 cut(s)	905, 1077, 1143
BshFI	GGCC	3 cut(s)	27, 1197, 1212
BshNI	GGYRCC	1 cut(s)	1236
BsiHKAI	GWGCWC	1 cut(s)	1267
BsiSI	CCGG	1 cut(s)	1209
BslFI	GGGAC	2 cut(s)	817, 1240
BslI	CCNNNNNNNGG	5 cut(s)	207, 251, 275, 551, 716
BsmAI	GTCTC	2 cut(s)	906, 1157
BsmFI	GGGAC	2 cut(s)	817, 1240
BsnI	GGCC	3 cut(s)	27, 1197, 1212
Bso31I	GGTCTC	2 cut(s)	906, 1157
Bsp1286I	GDGCHC	1 cut(s)	1267
Bsp143I	GATC	8 cut(s)	256, 320, 378, 427, 873, 1172, 1270, 1290
Bsp19I	CCATGG	2 cut(s)	1126, 1176
BspACI	CCGC	4 cut(s)	404, 856, 1101, 1193
BspANI	GGCC	3 cut(s)	27, 1197, 1212
BspCNI	CTCAG	4 cut(s)	526, 934, 1099, 1129
BspLI	GGNNCC	4 cut(s)	175, 1174, 1238, 1256
BspMI	ACCTGC	1 cut(s)	1242
BspPI	GGATC	3 cut(s)	328, 1167, 1180
BspT107I	GGYRCC	1 cut(s)	1236
BspTNI	GGTCTC	2 cut(s)	906, 1157
BsrBI	CCGCTC	1 cut(s)	856
BsrI	ACTGG	2 cut(s)	759, 843
BssECI	CCNNGG	6 cut(s)	28, 250, 1126, 1176, 1257, 1258
BssMI	GATC	8 cut(s)	256, 320, 378, 427, 873, 1172, 1270, 1290
BssNI	GRCGYC	1 cut(s)	558
BssSI	CACGAG	2 cut(s)	903, 1166
BssT1I	CCWWGG	4 cut(s)	28, 250, 1126, 1176
Bst2BI	CACGAG	2 cut(s)	903, 1166
Bst2UI	CCWGG	2 cut(s)	1245, 1259
Bst4CI	ACNGT	4 cut(s)	358, 785, 928, 988
Bst6I	CTCTTC	3 cut(s)	158, 812, 1314
BstACI	GRCGYC	1 cut(s)	558
BstC8I	GCNNGC	2 cut(s)	300, 1195
BstDEI	CTNAG	4 cut(s)	513, 942, 1107, 1137
BstDSI	CCRYGG	2 cut(s)	1126, 1176
BstH2I	RGCGCY	1 cut(s)	1107
BstHHI	GCGC	2 cut(s)	1055, 1106
BstKTI	GATC	8 cut(s)	259, 323, 381, 430, 876, 1175, 1273, 1293
BstMAI	GTCTC	2 cut(s)	906, 1157
BstMBI	GATC	8 cut(s)	256, 320, 378, 427, 873, 1172, 1270, 1290
BstNI	CCWGG	2 cut(s)	1245, 1259
BstNSI	RCATGY	1 cut(s)	595
BstSCI	CCNGG	2 cut(s)	1243, 1257
BstSLI	GKGCMC	1 cut(s)	1267
BstV1I	GCAGC	3 cut(s)	905, 1077, 1143
BstV2I	GAAGAC	1 cut(s)	966
BstX2I	RGATCY	2 cut(s)	1172, 1270
BstXI	CCANNNNNNTGG	1 cut(s)	184
BstYI	RGATCY	2 cut(s)	1172, 1270
BsuRI	GGCC	3 cut(s)	27, 1197, 1212
BtgI	CCRYGG	2 cut(s)	1126, 1176
BtsI	GCAGTG	1 cut(s)	855
BtsIMutI	CAGTG	5 cut(s)	836, 855, 924, 993, 1123
BveI	ACCTGC	1 cut(s)	1242
Cac8I	GCNNGC	2 cut(s)	300, 1195
CfoI	GCGC	2 cut(s)	1055, 1106
Cfr13I	GGNCC	5 cut(s)	174, 833, 1195, 1211, 1254
CseI	GACGC	1 cut(s)	566
Csp6I	GTAC	3 cut(s)	603, 756, 1163
CviAII	CATG	6 cut(s)	178, 184, 415, 592, 1127, 1177
CviQI	GTAC	3 cut(s)	603, 756, 1163
DdeI	CTNAG	4 cut(s)	513, 942, 1107, 1137
DpnI	GATC	8 cut(s)	258, 322, 380, 429, 875, 1174, 1272, 1292
DpnII	GATC	8 cut(s)	256, 320, 378, 427, 873, 1172, 1270, 1290
DraIII	CACNNNGTG	1 cut(s)	551
Eam1104I	CTCTTC	3 cut(s)	158, 812, 1314
EarI	CTCTTC	3 cut(s)	158, 812, 1314
Eco130I	CCWWGG	4 cut(s)	28, 250, 1126, 1176
Eco147I	AGGCCT	1 cut(s)	27
Eco31I	GGTCTC	2 cut(s)	906, 1157
Eco32I	GATATC	1 cut(s)	707
Eco47I	GGWCC	3 cut(s)	174, 833, 1254
Eco57I	CTGAAG	3 cut(s)	219, 621, 998
EcoO109I	RGGNCCY	1 cut(s)	1254
EcoRI	GAATTC	4 cut(s)	115, 191, 1339, 1367
EcoRII	CCWGG	2 cut(s)	1243, 1257
EcoRV	GATATC	1 cut(s)	707
EcoT14I	CCWWGG	4 cut(s)	28, 250, 1126, 1176
EcoT22I	ATGCAT	1 cut(s)	748
ErhI	CCWWGG	4 cut(s)	28, 250, 1126, 1176
FaeI	CATG	6 cut(s)	181, 187, 418, 595, 1130, 1180
FalI	AAGNNNNNCTT	2 cut(s)	1260, 1292
FaqI	GGGAC	2 cut(s)	817, 1240
FatI	CATG	6 cut(s)	177, 183, 414, 591, 1126, 1176
FauI	CCCGC	2 cut(s)	411, 1186
FauNDI	CATATG	1 cut(s)	206
FbaI	TGATCA	2 cut(s)	427, 873
Fnu4HI	GCNGC	4 cut(s)	894, 1066, 1102, 1132
Fsp4HI	GCNGC	4 cut(s)	894, 1066, 1102, 1132
FspBI	CTAG	4 cut(s)	479, 582, 798, 935
GlaI	GCGC	2 cut(s)	1054, 1105
GluI	GCNGC	4 cut(s)	894, 1066, 1102, 1132
GsuI	CTGGAG	2 cut(s)	42, 860
HaeII	RGCGCY	1 cut(s)	1107
HaeIII	GGCC	3 cut(s)	27, 1197, 1212
HapII	CCGG	1 cut(s)	1209
HgaI	GACGC	1 cut(s)	566
HhaI	GCGC	2 cut(s)	1055, 1106
Hin1I	GRCGYC	1 cut(s)	558
Hin1II	CATG	6 cut(s)	181, 187, 418, 595, 1130, 1180
Hin6I	GCGC	2 cut(s)	1053, 1104
HinP1I	GCGC	2 cut(s)	1053, 1104
HindIII	AAGCTT	1 cut(s)	1220
HinfI	GANTC	9 cut(s)	17, 101, 503, 649, 794, 887, 949, 996, 1122
HpaII	CCGG	1 cut(s)	1209
HphI	GGTGA	5 cut(s)	266, 498, 566, 958, 1037
Hpy166II	GTNNAC	2 cut(s)	836, 1265
Hpy188I	TCNGA	5 cut(s)	13, 238, 376, 516, 1290
Hpy8I	GTNNAC	2 cut(s)	836, 1265
HpyAV	CCTTC	4 cut(s)	519, 861, 1027, 1365
HpyCH4III	ACNGT	4 cut(s)	358, 785, 928, 988
HpyCH4IV	ACGT	2 cut(s)	778, 1200
HpyCH4V	TGCA	5 cut(s)	348, 746, 908, 1251, 1265
HpyF3I	CTNAG	4 cut(s)	513, 942, 1107, 1137
HpySE526I	ACGT	2 cut(s)	778, 1200
Hsp92I	GRCGYC	1 cut(s)	558
Hsp92II	CATG	6 cut(s)	181, 187, 418, 595, 1130, 1180
HspAI	GCGC	2 cut(s)	1053, 1104
Ksp22I	TGATCA	2 cut(s)	427, 873
Kzo9I	GATC	8 cut(s)	256, 320, 378, 427, 873, 1172, 1270, 1290
LmnI	GCTCC	1 cut(s)	853
Lsp1109I	GCAGC	3 cut(s)	905, 1077, 1143
LweI	GCATC	3 cut(s)	49, 755, 1238
MaeI	CTAG	4 cut(s)	479, 582, 798, 935
MaeII	ACGT	2 cut(s)	778, 1200
MaeIII	GTNAC	3 cut(s)	530, 554, 779
MalI	GATC	8 cut(s)	258, 322, 380, 429, 875, 1174, 1272, 1292
MbiI	CCGCTC	1 cut(s)	856
MboI	GATC	8 cut(s)	256, 320, 378, 427, 873, 1172, 1270, 1290
MflI	RGATCY	2 cut(s)	1172, 1270
MhlI	GDGCHC	1 cut(s)	1267
MlyI	GAGTC	2 cut(s)	26, 896
Mph1103I	ATGCAT	1 cut(s)	748
MseI	TTAA	1 cut(s)	1396
MslI	CAYNNNNRTG	4 cut(s)	45, 93, 182, 596
MspI	CCGG	1 cut(s)	1209
MspR9I	CCNGG	2 cut(s)	1245, 1259
MvaI	CCWGG	2 cut(s)	1245, 1259
NcoI	CCATGG	2 cut(s)	1126, 1176
NdeI	CATATG	1 cut(s)	206
NdeII	GATC	8 cut(s)	256, 320, 378, 427, 873, 1172, 1270, 1290
NlaIII	CATG	6 cut(s)	181, 187, 418, 595, 1130, 1180
NlaIV	GGNNCC	4 cut(s)	175, 1174, 1238, 1256
NmuCI	GTSAC	3 cut(s)	530, 554, 779
NsiI	ATGCAT	1 cut(s)	748
NspI	RCATGY	1 cut(s)	595
PasI	CCCWGGG	1 cut(s)	1258
PceI	AGGCCT	1 cut(s)	27
PfeI	GAWTC	7 cut(s)	101, 503, 649, 794, 949, 996, 1122
PflMI	CCANNNNNTGG	3 cut(s)	207, 551, 716
PkrI	GCNGC	4 cut(s)	895, 1067, 1103, 1133
PleI	GAGTC	2 cut(s)	25, 895
PpsI	GAGTC	2 cut(s)	25, 895
PpuMI	RGGWCCY	1 cut(s)	1254
Psp5II	RGGWCCY	1 cut(s)	1254
Psp6I	CCWGG	2 cut(s)	1243, 1257
PspGI	CCWGG	2 cut(s)	1243, 1257
PspN4I	GGNNCC	4 cut(s)	175, 1174, 1238, 1256
PspPI	GGNCC	5 cut(s)	174, 833, 1195, 1211, 1254
PspPPI	RGGWCCY	1 cut(s)	1254
PsuI	RGATCY	2 cut(s)	1172, 1270
RsaI	GTAC	3 cut(s)	604, 757, 1164
RsaNI	GTAC	3 cut(s)	603, 756, 1163
RseI	CAYNNNNRTG	4 cut(s)	45, 93, 182, 596
SaqAI	TTAA	1 cut(s)	1396
SatI	GCNGC	4 cut(s)	894, 1066, 1102, 1132
Sau3AI	GATC	8 cut(s)	256, 320, 378, 427, 873, 1172, 1270, 1290
Sau96I	GGNCC	5 cut(s)	174, 833, 1195, 1211, 1254
SchI	GAGTC	2 cut(s)	26, 896
ScrFI	CCNGG	2 cut(s)	1245, 1259
SduI	GDGCHC	1 cut(s)	1267
SfaNI	GCATC	3 cut(s)	49, 755, 1238
SinI	GGWCC	3 cut(s)	174, 833, 1254
SmiMI	CAYNNNNRTG	4 cut(s)	45, 93, 182, 596
SseBI	AGGCCT	1 cut(s)	27
SsiI	CCGC	4 cut(s)	404, 856, 1101, 1193
SspMI	CTAG	4 cut(s)	479, 582, 798, 935
StuI	AGGCCT	1 cut(s)	27
StyD4I	CCNGG	2 cut(s)	1243, 1257
StyI	CCWWGG	4 cut(s)	28, 250, 1126, 1176
TaaI	ACNGT	4 cut(s)	358, 785, 928, 988
TaiI	ACGT	2 cut(s)	781, 1203
TaqI	TCGA	5 cut(s)	119, 381, 391, 883, 999
TaqII	GACCGA	1 cut(s)	888
TauI	GCSGC	1 cut(s)	1104
TfiI	GAWTC	7 cut(s)	101, 503, 649, 794, 949, 996, 1122
Tru1I	TTAA	1 cut(s)	1396
Tru9I	TTAA	1 cut(s)	1396
TscAI	CASTG	5 cut(s)	843, 855, 931, 993, 1123
TseFI	GTSAC	3 cut(s)	530, 554, 779
TseI	GCWGC	3 cut(s)	893, 1065, 1131
Tsp45I	GTSAC	3 cut(s)	530, 554, 779
TspRI	CASTG	5 cut(s)	843, 855, 931, 993, 1123
Van91I	CCANNNNNTGG	3 cut(s)	207, 551, 716
VneI	GTGCAC	1 cut(s)	1263
VpaK11BI	GGWCC	3 cut(s)	174, 833, 1254
XapI	RAATTY	6 cut(s)	115, 144, 191, 1111, 1339, 1367
XceI	RCATGY	1 cut(s)	595
XspI	CTAG	4 cut(s)	479, 582, 798, 935
Zsp2I	ATGCAT	1 cut(s)	748

Using CommOnly database (standard laboratory enzymes). Scanned on CDS sequence.