Rroxscaffold_3G00233100

phosphatase 2c

Design Primers CRISPR Design

Basic Information

Type: gene
Biological Identity
rosa_roxburghii
GWHEROQ00000003
OG0004725
Physical Location & Seq
Forward (+)
18952127 .. 18960119
7993 bp
Loading structure...
UTR
Exon/CDS
Intron
Rroxscaffold_3G00233100.1

Sequence Viewer

Length: 2298 bp
ATGTTCGTTTTGATTTGCGCTGTGACAGAGTGCTCGGACGGTAGCATCGTCTTCCGATTTGGCGACGCGAGCGAGAAGGCTGCTTTTGATGAGATTAGTAGTCGTCTGGATGCGGAATTGAGTGAAGGATTGGTTGGGGAAAGCAATGTGGAAGAATCGGAGATTGATGCTTCTCCGGCGAGTTCACCTGAGGAGACTGCCTTCCTTGATTTGACCACGGCGGATGAAGTCCAAAGCTCAGCAGTGTTTTTGCATGAACTAATGGTCAAATTGGAAGGTGACAGCTTCCCTTCCACAGATTCACAAGGTCTATTCACAGAGGGTGAATCACACAGCAGGGATGTTTCAGGAGCATCAGCTCCTAGGAAATTGGAATTCCATACAATTGAAGGATCAGAGGGGTATCCTTCCACAATTTCAGAAGATGAATTGATAGAGGGTGAAGCACACAGAAGAGATGTCACAGGAGCATCAACTCCTATGCCAGTGGAATTGCATACATCTGAAGGCTCTGAGTGGTATCCTTCTGCAAAGTCAGAAGATGAATTAACAGAGGGTAAAGAAGATAACACATATGTCATAGGAGCATCAATTCCTACGTCAGTGGAATTGCATACATCTGAAGGCTCTGAGGGTTATCCTTCTACAAAGTCAGAAGATGAATTAACAGAGGGTAAAGAAGATAGAAGAGATGTCACAGGAGCATCAACTCCTGTGCCTGTGGAATTGCATACATCTGAAGGCTCTGAGGGGTATCCATCTACAAAATCAGAAGATGAAGTAACAGAGGGCAAAGAACACAGAAGAGATGTTACAGGAGCATCAACTTCTTTGCCAGTGGAATTGCATACATTTGAAGGTTCTGAGGGGTATCATTCTACAAAGTCAGAAGAGGAATCAACAGAGGGTAAAGAAGATAGAACAGATGTCACAGGAGCACCAACTTCAAGGACGGTTGAATTGCATGCTGTTGAAGGCTCTGCGGGATATCCCTCCACGAAGTCAGAAGATGAATTGACAGAGGGGAAAGCCCATCAAAGAGATGTCTCAGGAGCATCAACTGATAGGACAGAGGATATGCATACTGTTGAAGGTGCTGAGGAGAATGCTTCCATAAATTCAGAGGATGAATTTGTAGATGATAAAACACATAGAAGAGATGCTTCAGGAGAATTAATTCCTAGGATATTGGAATTTCATTCAGTTGAAGGCGCTTTGGGGTATCCTTTAACAAAATCAGAAGATGAATCAACAGGGGGTAAAGAACAAAGAGATGTTTCAGGAGCATCAACTCCTAGAACAGTGGAATTGCATACAGTTGAAGGTGCTCAGGGGTATGCTTCCATAAAGTTAGAAGATAAATTCGCAGAGGGTAAACTAGATAGAACAGATGTTTTAGAAGCAGCAACTCCCAGGACAGTGGAATTGCACACAGTTGAACGTGCTAAGGATACCCCTTCAAAAAAGTTGGAAGATGAATTCACAGAGGGTAAAGCACATATAAGAAATGTTCCGGGAGCATCAACTCCAAGGATAGTGGAATTGCATACAGTTGAAGGCGCTGAAGGGGAGGACATCTCTGCACTTCTACTATCGACTGGTGCTGCATTATTACCGCATCCCTCAAAGGCATTGACAGGTGGGGAGGATGCTTGCTTTGTCGCTTCTAGAAACTGGCTAGGTGTGGCTGATGGAGTTGGTCAGTGGTCACTGGAAGGGATTAATCCTGGGCTATATGCCCGTGAACTCATGGAGAACTGTGAAAGGTTTCTATCTGATAGCAAAGGTATACCATTGACTAAACCAGAAGACGTCCTTGTTGGAGGTGCTACACAAACAAAATCTCCTGGTTCATCTACAGTTTTAGTCGCTTACTTTGATGGTCAGGCCCTCCATGTGGCAAACATTGGTAACTCAGGATTTATCATAATCAGAAATGGTGCTGTTTTTAAGAGATCATCTCCAATGATTCATGGCTTCAATTTTCCATTACAAATTAGATACAGTATAGATTTGGATGAGGGAGATGTCATTGTCACTGCAACGGATGGCCTTTTTGACAACTTGTATGAGCAAGAAATCACTTCAATTGTAACCAAGTCATTCCAAAGGAGAATGAGACTTCAGGATATAGCAGAGTTCTTGGCCGCTAGCGCACAGGAGGTTGGGCAATCTACTTCTAGAAGAAGCCCATTCGCAGATGCAGTTGAGGCATCTGGGTATGTGGGATATACTGGTGGCAAGCTCGATGATGTGACAGTTATTGTGTCGCTTGTACAGAAGAAAAGCAGCTGTTAG
Functional Annotation
Gene Ontology
Molecular Function
Biological Process
Cellular Component
GO:0005575 GO:0005622 GO:0005623 GO:0005737 GO:0009507 GO:0009536 GO:0043226 GO:0043227 GO:0043229 GO:0043231 GO:0044424 GO:0044444 GO:0044464
KEGG Pathways
Metabolic & Signaling
No pathways identified.
Pfam Domains
Protein Families
PP2C SpoIIE

Protein Analysis

765

Amino Acids

82.26

Weight (kDa)

4.51

Isoelectric Point (pI)

42.5

Instability Index
Protein Domains (Pfam)
No Pfam domains detected for this protein.
Hydrophobicity Profile (Kyte-Doolittle)
AI Structure Prediction Report
Calculating structure properties...

Genomic Context

Gene Family Tree

Species Filter
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Publication-ready
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Restriction Enzyme Sites

1 / 10
Enzyme Recognition Site Cut Count Positions (bp)
AatII GACGTC 1 cut(s) 1815
AccI GTMKAC 1 cut(s) 1789
AccII CGCG 1 cut(s) 68
AciI CCGC 5 cut(s) 113, 221, 983, 1616, 2148
AclWI GGATC 1 cut(s) 400
AcoI YGGCCR 1 cut(s) 2145
AcsI RAATTY 6 cut(s) 374, 1117, 1130, 1193, 1361, 1478
AcuI CTGAAG 6 cut(s) 525, 642, 759, 1149, 1584, 2108
AcyI GRCGYC 1 cut(s) 1812
AfaI GTAC 1 cut(s) 2277
AfiI CCNNNNNNNGG 1 cut(s) 1897
AjnI CCWGG 3 cut(s) 1412, 1726, 1846
AjuI GAANNNNNNNTTGG 2 cut(s) 117, 149
AluBI AGCT 5 cut(s) 237, 285, 359, 2245, 2292
AluI AGCT 5 cut(s) 237, 285, 359, 2245, 2292
Alw21I GWGCWC 3 cut(s) 35, 940, 1330
Alw26I GTCTC 3 cut(s) 188, 1051, 2113
AlwI GGATC 1 cut(s) 400
AoxI GGCC 3 cut(s) 1887, 2050, 2145
ApeKI GCWGC 4 cut(s) 80, 1403, 1604, 2289
ApoI RAATTY 6 cut(s) 374, 1117, 1130, 1193, 1361, 1478
AseI ATTAAT 2 cut(s) 1175, 1722
Asp700I GAANNNNTTC 2 cut(s) 1176, 1767
AspA2I CCTAGG 2 cut(s) 362, 1181
AspLEI GCGC 4 cut(s) 20, 1214, 1562, 2156
AspS9I GGNCC 1 cut(s) 1888
AsuC2I CCSGG 1 cut(s) 1515
AsuHPI GGTGA 4 cut(s) 177, 290, 335, 452
AsuNHI GCTAGC 1 cut(s) 2150
AvrII CCTAGG 2 cut(s) 362, 1181
AxyI CCTNAGG 1 cut(s) 189
BarI GAAGNNNNNNTAC 2 cut(s) 796, 828
BbsI GAAGAC 2 cut(s) 43, 1815
Bbv12I GWGCWC 3 cut(s) 35, 940, 1330
BbvCI CCTCAGC 1 cut(s) 1098
BbvI GCAGC 3 cut(s) 67, 1415, 1591
BccI CCATC 5 cut(s) 766, 1041, 1685, 1874, 2042
BceAI ACGGC 1 cut(s) 234
BcgI CGANNNNNNTGC 2 cut(s) 62, 96
BciT130I CCWGG 3 cut(s) 1414, 1728, 1848
BciVI GTATCC 5 cut(s) 414, 531, 765, 1233, 1444
BcnI CCSGG 1 cut(s) 1515
BcoDI GTCTC 3 cut(s) 188, 1051, 2113
BfaI CTAG 8 cut(s) 363, 1182, 1296, 1379, 1668, 1679, 2151, 2181
BfmI CTRYAG 1 cut(s) 1857
BfoI RGCGCY 2 cut(s) 1215, 1563
BfuI GTATCC 5 cut(s) 414, 531, 765, 1233, 1444
BisI GCNGC 5 cut(s) 81, 1404, 1605, 2148, 2290
BlnI CCTAGG 2 cut(s) 362, 1181
BlpI GCTNAGC 1 cut(s) 238
BlsI GCNGC 5 cut(s) 82, 1405, 1606, 2149, 2291
Bme1390I CCNGG 4 cut(s) 1414, 1515, 1728, 1848
BmgT120I GGNCC 1 cut(s) 1888
BmrFI CCNGG 4 cut(s) 1414, 1515, 1728, 1848
BmtI GCTAGC 1 cut(s) 2154
BpiI GAAGAC 2 cut(s) 43, 1815
BplI GAGNNNNNCTC 4 cut(s) 1562, 1594, 1945, 1977
Bpu10I CCTNAGC 3 cut(s) 1098, 1329, 1446
Bpu1102I GCTNAGC 1 cut(s) 238
BpuMI CCSGG 1 cut(s) 1515
BsaHI GRCGYC 1 cut(s) 1812
BsaJI CCNNGG 6 cut(s) 216, 362, 1181, 1412, 1529, 1727
BsaXI ACNNNNNCTCC 2 cut(s) 1828, 1858
Bsc4I CCNNNNNNNGG 1 cut(s) 1897
Bse1I ACTGG 6 cut(s) 485, 836, 1603, 1679, 1716, 2239
Bse21I CCTNAGG 1 cut(s) 189
Bse3DI GCAATG 1 cut(s) 151
BseBI CCWGG 3 cut(s) 1414, 1728, 1848
BseDI CCNNGG 6 cut(s) 216, 362, 1181, 1412, 1529, 1727
BseGI GGATG 8 cut(s) 115, 229, 346, 1132, 1618, 1654, 2023, 2053
BseLI CCNNNNNNNGG 1 cut(s) 1897
BseMI GCAATG 1 cut(s) 151
BseNI ACTGG 6 cut(s) 485, 836, 1603, 1679, 1716, 2239
BseRI GAGGAG 2 cut(s) 206, 1115
BseXI GCAGC 3 cut(s) 67, 1415, 1591
BsgI GTGCAG 1 cut(s) 1566
Bsh1236I CGCG 1 cut(s) 68
BshFI GGCC 3 cut(s) 1889, 2052, 2147
BsiHKAI GWGCWC 3 cut(s) 35, 940, 1330
BsiSI CCGG 2 cut(s) 176, 1514
BslI CCNNNNNNNGG 1 cut(s) 1897
BsmAI GTCTC 3 cut(s) 188, 1051, 2113
BsmI GAATGC 1 cut(s) 1111
BsnI GGCC 3 cut(s) 1889, 2052, 2147
Bsp1286I GDGCHC 3 cut(s) 35, 940, 1330
Bsp1407I TGTACA 1 cut(s) 2275
Bsp143I GATC 2 cut(s) 392, 1955
Bsp1720I GCTNAGC 1 cut(s) 238
BspACI CCGC 5 cut(s) 113, 221, 983, 1616, 2148
BspANI GGCC 3 cut(s) 1889, 2052, 2147
BspFNI CGCG 1 cut(s) 68
BspOI GCTAGC 1 cut(s) 2154
BspPI GGATC 1 cut(s) 400
BsrDI GCAATG 1 cut(s) 151
BsrGI TGTACA 1 cut(s) 2275
BsrI ACTGG 6 cut(s) 485, 836, 1603, 1679, 1716, 2239
BssECI CCNNGG 6 cut(s) 216, 362, 1181, 1412, 1529, 1727
BssMI GATC 2 cut(s) 392, 1955
BssNAI GTATAC 1 cut(s) 1790
BssNI GRCGYC 1 cut(s) 1812
BssT1I CCWWGG 3 cut(s) 362, 1181, 1529
Bst1107I GTATAC 1 cut(s) 1790
Bst2UI CCWGG 3 cut(s) 1414, 1728, 1848
Bst6I CTCTTC 5 cut(s) 448, 682, 799, 885, 1150
BstACI GRCGYC 1 cut(s) 1812
BstAUI TGTACA 1 cut(s) 2275
BstC8I GCNNGC 5 cut(s) 70, 966, 1654, 2152, 2243
BstDSI CCRYGG 1 cut(s) 216
BstF5I GGATG 8 cut(s) 115, 229, 346, 1132, 1618, 1654, 2023, 2053
BstFNI CGCG 1 cut(s) 68
BstH2I RGCGCY 2 cut(s) 1215, 1563
BstHHI GCGC 4 cut(s) 20, 1214, 1562, 2156
BstKTI GATC 2 cut(s) 395, 1958
BstMAI GTCTC 3 cut(s) 188, 1051, 2113
BstMBI GATC 2 cut(s) 392, 1955
BstMWI GCNNNNNNNGC 4 cut(s) 69, 176, 2153, 2210
BstNI CCWGG 3 cut(s) 1414, 1728, 1848
BstNSI RCATGY 1 cut(s) 968
BstSCI CCNGG 4 cut(s) 1412, 1513, 1726, 1846
BstSFI CTRYAG 1 cut(s) 1857
BstUI CGCG 1 cut(s) 68
BstV1I GCAGC 3 cut(s) 67, 1415, 1591
BstV2I GAAGAC 2 cut(s) 43, 1815
BstXI CCANNNNNNTGG 1 cut(s) 1420
BstZ17I GTATAC 1 cut(s) 1790
Bsu36I CCTNAGG 1 cut(s) 189
BsuI GTATCC 5 cut(s) 414, 531, 765, 1233, 1444
BsuRI GGCC 3 cut(s) 1889, 2052, 2147
BtgI CCRYGG 1 cut(s) 216
BtsCI GGATG 8 cut(s) 115, 229, 346, 1132, 1618, 1654, 2023, 2053
BtsI GCAGTG 2 cut(s) 249, 2037
BtsIMutI CAGTG 9 cut(s) 249, 492, 609, 843, 1308, 1425, 1709, 1709, 2037
Cac8I GCNNGC 5 cut(s) 70, 966, 1654, 2152, 2243
CfoI GCGC 4 cut(s) 20, 1214, 1562, 2156
Cfr13I GGNCC 1 cut(s) 1888
CseI GACGC 1 cut(s) 74
Csp6I GTAC 1 cut(s) 2276
CspCI CAANNNNNGTGG 2 cut(s) 1518, 1553
CviAII CATG 5 cut(s) 254, 965, 1750, 1895, 1973
CviQI GTAC 1 cut(s) 2276
DpnI GATC 2 cut(s) 394, 1957
DpnII GATC 2 cut(s) 392, 1955
EaeI YGGCCR 1 cut(s) 2145
Eam1104I CTCTTC 5 cut(s) 448, 682, 799, 885, 1150
EarI CTCTTC 5 cut(s) 448, 682, 799, 885, 1150
EciI GGCGGA 1 cut(s) 236
Eco130I CCWWGG 3 cut(s) 362, 1181, 1529
Eco32I GATATC 1 cut(s) 989
Eco57I CTGAAG 6 cut(s) 525, 642, 759, 1149, 1584, 2108
Eco81I CCTNAGG 1 cut(s) 189
EcoO109I RGGNCCY 1 cut(s) 1888
EcoRI GAATTC 2 cut(s) 374, 1478
EcoRII CCWGG 3 cut(s) 1412, 1726, 1846
EcoRV GATATC 1 cut(s) 989
EcoT14I CCWWGG 3 cut(s) 362, 1181, 1529
EcoT22I ATGCAT 1 cut(s) 1083
ErhI CCWWGG 3 cut(s) 362, 1181, 1529
FaeI CATG 5 cut(s) 257, 968, 1753, 1898, 1976
FalI AAGNNNNNCTT 4 cut(s) 1147, 1179, 1800, 1832
FatI CATG 5 cut(s) 253, 964, 1749, 1894, 1972
FauI CCCGC 1 cut(s) 976
FauNDI CATATG 1 cut(s) 574
FblI GTMKAC 1 cut(s) 1789
Fnu4HI GCNGC 5 cut(s) 81, 1404, 1605, 2148, 2290
FokI GGATG 8 cut(s) 122, 236, 353, 1139, 1605, 1661, 2030, 2060
Fsp4HI GCNGC 5 cut(s) 81, 1404, 1605, 2148, 2290
FspBI CTAG 8 cut(s) 363, 1182, 1296, 1379, 1668, 1679, 2151, 2181
GlaI GCGC 4 cut(s) 19, 1213, 1561, 2155
GluI GCNGC 5 cut(s) 81, 1404, 1605, 2148, 2290
HaeII RGCGCY 2 cut(s) 1215, 1563
HaeIII GGCC 3 cut(s) 1889, 2052, 2147
HapII CCGG 2 cut(s) 176, 1514
HgaI GACGC 1 cut(s) 74
HhaI GCGC 4 cut(s) 20, 1214, 1562, 2156
Hin1I GRCGYC 1 cut(s) 1812
Hin1II CATG 5 cut(s) 257, 968, 1753, 1898, 1976
Hin6I GCGC 4 cut(s) 18, 1212, 1560, 2154
HinP1I GCGC 4 cut(s) 18, 1212, 1560, 2154
HinfI GANTC 6 cut(s) 155, 299, 326, 896, 1247, 1969
HpaII CCGG 2 cut(s) 176, 1514
HphI GGTGA 4 cut(s) 177, 290, 335, 452
Hpy166II GTNNAC 4 cut(s) 185, 1376, 1745, 1790
Hpy188III TCNNGA 9 cut(s) 107, 348, 1050, 1167, 1281, 1668, 1917, 2126, 2181
Hpy8I GTNNAC 4 cut(s) 185, 1376, 1745, 1790
Hpy99I CGWCG 1 cut(s) 68
HpyCH4IV ACGT 3 cut(s) 599, 1441, 1812
HpyF10VI GCNNNNNNNGC 4 cut(s) 69, 176, 2153, 2210
HpySE526I ACGT 3 cut(s) 599, 1441, 1812
Hsp92I GRCGYC 1 cut(s) 1812
Hsp92II CATG 5 cut(s) 257, 968, 1753, 1898, 1976
HspAI GCGC 4 cut(s) 18, 1212, 1560, 2154
Kzo9I GATC 2 cut(s) 392, 1955
Lsp1109I GCAGC 3 cut(s) 67, 1415, 1591
MaeI CTAG 8 cut(s) 363, 1182, 1296, 1379, 1668, 1679, 2151, 2181
MaeII ACGT 3 cut(s) 599, 1441, 1812
MalI GATC 2 cut(s) 394, 1957
MboI GATC 2 cut(s) 392, 1955
MfeI CAATTG 2 cut(s) 384, 2088
MhlI GDGCHC 3 cut(s) 35, 940, 1330
MmeI TCCRAC 2 cut(s) 1449, 1801
Mph1103I ATGCAT 1 cut(s) 1083
MroXI GAANNNNTTC 2 cut(s) 1176, 1767
MseI TTAA 6 cut(s) 548, 665, 1175, 1229, 1722, 1950
MspA1I CMGCKG 1 cut(s) 2292
MspI CCGG 2 cut(s) 176, 1514
MspR9I CCNGG 4 cut(s) 1414, 1515, 1728, 1848
MunI CAATTG 2 cut(s) 384, 2088
Mva1269I GAATGC 1 cut(s) 1111
MvaI CCWGG 3 cut(s) 1414, 1728, 1848
MvnI CGCG 1 cut(s) 68
MwoI GCNNNNNNNGC 4 cut(s) 69, 176, 2153, 2210
NciI CCSGG 1 cut(s) 1515
NdeI CATATG 1 cut(s) 574
NdeII GATC 2 cut(s) 392, 1955
NheI GCTAGC 1 cut(s) 2150
NlaIII CATG 5 cut(s) 257, 968, 1753, 1898, 1976
NmuCI GTSAC 8 cut(s) 22, 278, 460, 694, 928, 1707, 2035, 2254
NsiI ATGCAT 1 cut(s) 1083
NspI RCATGY 1 cut(s) 968
PaeI GCATGC 1 cut(s) 968
PcsI WCGNNNNNNNCGW 1 cut(s) 45
PctI GAATGC 1 cut(s) 1111
PdmI GAANNNNTTC 2 cut(s) 1176, 1767
PfeI GAWTC 6 cut(s) 155, 299, 326, 896, 1247, 1969
PfoI TCCNGGA 1 cut(s) 1513
PkrI GCNGC 5 cut(s) 82, 1405, 1606, 2149, 2291
PshBI ATTAAT 2 cut(s) 1175, 1722
Psp6I CCWGG 3 cut(s) 1412, 1726, 1846
PspGI CCWGG 3 cut(s) 1412, 1726, 1846
PspPI GGNCC 1 cut(s) 1888
PvuII CAGCTG 1 cut(s) 2292
RsaI GTAC 1 cut(s) 2277
RsaNI GTAC 1 cut(s) 2276
SaqAI TTAA 6 cut(s) 548, 665, 1175, 1229, 1722, 1950
SatI GCNGC 5 cut(s) 81, 1404, 1605, 2148, 2290
Sau3AI GATC 2 cut(s) 392, 1955
Sau96I GGNCC 1 cut(s) 1888
ScrFI CCNGG 4 cut(s) 1414, 1515, 1728, 1848
SduI GDGCHC 3 cut(s) 35, 940, 1330
SfcI CTRYAG 1 cut(s) 1857
SphI GCATGC 1 cut(s) 968
SsiI CCGC 5 cut(s) 113, 221, 983, 1616, 2148
SspMI CTAG 8 cut(s) 363, 1182, 1296, 1379, 1668, 1679, 2151, 2181
StyD4I CCNGG 4 cut(s) 1412, 1513, 1726, 1846
StyI CCWWGG 3 cut(s) 362, 1181, 1529
TaiI ACGT 3 cut(s) 602, 1444, 1815
TaqI TCGA 2 cut(s) 1595, 2247
TatI WGTACW 1 cut(s) 2275
TauI GCSGC 1 cut(s) 2150
TfiI GAWTC 6 cut(s) 155, 299, 326, 896, 1247, 1969
Tru1I TTAA 6 cut(s) 548, 665, 1175, 1229, 1722, 1950
Tru9I TTAA 6 cut(s) 548, 665, 1175, 1229, 1722, 1950
TscAI CASTG 9 cut(s) 249, 492, 609, 843, 1308, 1425, 1709, 1716, 2044
TseFI GTSAC 8 cut(s) 22, 278, 460, 694, 928, 1707, 2035, 2254
TseI GCWGC 4 cut(s) 80, 1403, 1604, 2289
Tsp45I GTSAC 8 cut(s) 22, 278, 460, 694, 928, 1707, 2035, 2254
TspGWI ACGGA 1 cut(s) 2060
TspRI CASTG 9 cut(s) 249, 492, 609, 843, 1308, 1425, 1709, 1716, 2044
VspI ATTAAT 2 cut(s) 1175, 1722
XapI RAATTY 6 cut(s) 374, 1117, 1130, 1193, 1361, 1478
XbaI TCTAGA 2 cut(s) 1667, 2180
XceI RCATGY 1 cut(s) 968
XmaJI CCTAGG 2 cut(s) 362, 1181
XmiI GTMKAC 1 cut(s) 1789
XmnI GAANNNNTTC 2 cut(s) 1176, 1767
XspI CTAG 8 cut(s) 363, 1182, 1296, 1379, 1668, 1679, 2151, 2181
ZraI GACGTC 1 cut(s) 1813
Zsp2I ATGCAT 1 cut(s) 1083
Using CommOnly database (standard laboratory enzymes). Scanned on CDS sequence.