Basic Information

Type: gene

Biological Identity

Species

fragaria_vesca

Chromosome

Fvb2

Orthogroup ID

OG0003187

Physical Location & Seq

Genomic Coordinates Forward (+)

25851652 .. 25855668

Gene Structure 4017 bp

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UTR

Exon/CDS

Intron

Transcript / Protein ID

FvH4_2g34930.t2

Sequence Viewer

CDS Protein Promoter (2k)

Length: 1611 bp

ATGCAAGGGAGTAAAGAAAGGAATATCACAAAGGCAATCAGCCTTGGAAAAGCGTCAAGAAGAGAGAGAAAATTGGCTTTGCAACAAGATGTTGATAAGCTGAAGAAGAAGCTTAGACATGAAGAGAATGTACACAGAGCTTTGCAGAGGGCATTCAATAGACCCCTGGGAGCTCTACCTCGTCTACCACCTTATCTGCCTCCTTATACATTAGAGCTTCTTGCTGAGGTTGCTGTTTTGGAAGAGGAAGTTGTTCGGCTTGAAGAACAAGTAGTGCATTTTAGGAAGGACTTGTATCAGGAAGCTGTCAACATTTCGACTTCCAAGAGGAGCTTAGAAACCTCAGCTGAACTATGTGATTCAAACCCTAAAAAGAATCATAAATTTCATGGTAGTATAGTGGACACTGATACCGCGGTGAAGCATTCGCCATCCCCTTCTGATCATAAGCAAGAGAATCAATCATGTAATCCCTCAATGAAGAACAACAAGAAATCCTTGATCCATAGTAATGCTCAGGTTAGGACTCCAGCGAAAAGACCTCCAGTTGATCCGAAAATAGCACAAAAGCGTTTAGATTCTCCAAAACTACAGTTAGAAGTTAGAGTAACTGAACAAGAGAGCACAGAAGCAAGACTTTCTAGTATTCCAGAAAAAAAGCCATCCGGAGATGACAGCCCAAACAAAATATCTGAAAATATTCTGAAGTGCTTATCAAGCATTTTCATGAGAATGAGTTCAGCAAAGGGCATTACGGAAAACCAGCCTTCCTTTTCAACATTAGGAATCCAACAAAGCAATGAAAAACCAGAATTTTGGGATCCTTATGGTATCTGTTCAGAATTTGGACGGAGAGATATTGGTCCATATAAGCAATTACATGCAGTTGAAGCCCGCTCTATTAATCCTAATCGAACAGCAAGCTCTTTGTTCCTGCTCCGAAGATTGAAATTACTGCTAGGGAAACTTGCTTCTGTAAATTTGAAAAGCCTCGGTCATCAAGAGAAGCTTGCATTCTGGATTAATATCTACAATTCCTGCATGATGAATGCTTTCTTAGAACATGGAATACCAGAGAGGCCTGAAATAATTGTCGCGCTAATGCAGAAGGCAACTATAAATGTTGGGGGGCACTTGCTCAGTGCAATAACTATAGAACATTTTATCTTGAGGCTGCCATATCACTCAAAATATACCTTCTCCAAGGGTGCAAAGAATGATGAGAATACAGCAAGAAGCATTTTCGCATTGGAATTATCTGAACCGCTGGTAACATTTGCTCTATCATGTGGAAGCTGGTCCTCCCCTGCTGTAAGAGTTTACACTGCATCCCAAGTTGACAATGAACTAGAAATCGCAAAAAGAGAGTACTTGCAGGCTGCAGTTGGAATTTCATCGAGCAAGTTTGCAATTCCAAAGCTTTTGGATTGGTACCTACTTGATTACGCAAAGGATTTAGAGTCGTTGCTAGACTGGATCTGCCTTCAGCTACCTAGTGAACTAGGAAAAGAAGCAATCAAGTTTCTTGAAGGAGCCAAAACCGATCCTCATTCACAGTTTGTCCAAATCATGCCCTATGAGTTCAGCTTTAGGTATCTTCTACACACATGA

Functional Annotation

Gene Ontology

Molecular Function
Biological Process
Cellular Component

No ontology terms assigned.

KEGG Pathways

Metabolic & Signaling

No pathways identified.

Pfam Domains

Protein Families

DUF547 Lzipper-MIP1

Protein Analysis

537

Amino Acids

60.63

Weight (kDa)

9.13

Isoelectric Point (pI)

54.48

Instability Index

Protein Domains (Pfam)

Domain Name	Pfam ID	Position	E-value	Description
Lzipper-MIP1	PF14389	47 - 128	3e-23	Leucine-zipper of ternary complex factor MIP1
DUF547	PF04784	356 - 485	8.5e-40	Protein of unknown function, DUF547

Hydrophobicity Profile (Kyte-Doolittle)

AI Structure Prediction Report

Calculating structure properties...

Genomic Context

Gene Family Tree

Species Filter

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Align Labels (Cladogram)

Image

Tree File

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Publication-ready

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Orthologous Genes (Group: OG0003187)

Species	Orthologous Gene IDs
arabidopsis_thaliana	AT5G42690 AT5G42690 AT5G42690 AT5G42690 AT5G42690
fragaria_vesca	FvH4_2g34930 FvH4_2g34930 FvH4_2g34930 FvH4_2g34930 FvH4_2g34930 FvH4_2g34930 FvH4_2g34930 FvH4_2g34930 FvH4_2g34930 FvH4_2g34930 FvH4_2g34930
malus_domestica	MD08G1106800.v1.1 MD15G1087500.v1.1
prunus_persica	Prupe.1G441200_v2.0.a1 Prupe.1G441200_v2.0.a1 Prupe.1G441200_v2.0.a1 Prupe.1G441200_v2.0.a1
pyrus_communis	pycom08g08840 pycom15g08230
rosa_chinensis	RchiOBHm_Chr6g0307071
rosa_laevigata	RLG00000010736
rosa_multiflora	Rmu_sc0005634.1_g000003
rosa_roxburghii	Rroxscaffold_7G00161180
rosa_rugosa	Rorug06G0355700 Rorug06G0355800 Rorug06G0355900 Rorug06G0356000
rosa_samantha	Rh6AG467200 Rh6BG433300 Rh6CG482700 Rh6DG469000
rosa_wichuraiana	Rw6G040730

Restriction Enzyme Sites

1 / 10

Enzyme	Recognition Site	Cut Count	Positions (bp)
Acc65I	GGTACC	1 cut(s)	1431
AccB1I	GGYRCC	1 cut(s)	1431
AccBSI	CCGCTC	1 cut(s)	897
AccI	GTMKAC	1 cut(s)	184
AccII	CGCG	2 cut(s)	416, 1097
AccIII	TCCGGA	1 cut(s)	665
AciI	CCGC	4 cut(s)	414, 416, 895, 1265
AclWI	GGATC	6 cut(s)	496, 545, 815, 828, 1484, 1538
AcsI	RAATTY	5 cut(s)	383, 812, 842, 979, 1389
AcuI	CTGAAG	3 cut(s)	122, 725, 1469
AfaI	GTAC	3 cut(s)	132, 1370, 1433
AgsI	TTSAA	8 cut(s)	157, 263, 363, 777, 890, 949, 985, 1529
AjnI	CCWGG	1 cut(s)	165
Alw21I	GWGCWC	2 cut(s)	175, 626
AlwI	GGATC	6 cut(s)	496, 545, 815, 828, 1484, 1538
Aor13HI	TCCGGA	1 cut(s)	665
AoxI	GGCC	1 cut(s)	1079
ApeKI	GCWGC	2 cut(s)	1174, 1379
ApoI	RAATTY	5 cut(s)	383, 812, 842, 979, 1389
AseI	ATTAAT	2 cut(s)	903, 1023
Asp700I	GAANNNNTTC	4 cut(s)	252, 699, 736, 1052
Asp718I	GGTACC	1 cut(s)	1431
AspLEI	GCGC	1 cut(s)	1099
AspS9I	GGNCC	2 cut(s)	863, 1299
AsuHPI	GGTGA	1 cut(s)	430
AvaII	GGWCC	2 cut(s)	863, 1299
BaeGI	GKGCMC	1 cut(s)	1134
BaeI	ACNNNNGTAYC	2 cut(s)	402, 435
BamHI	GGATCC	1 cut(s)	820
BanI	GGYRCC	1 cut(s)	1431
BanII	GRGCYC	1 cut(s)	175
BarI	GAAGNNNNNNTAC	2 cut(s)	114, 146
Bbv12I	GWGCWC	2 cut(s)	175, 626
BbvCI	CCTCAGC	2 cut(s)	225, 343
BbvI	GCAGC	2 cut(s)	1161, 1366
BccI	CCATC	2 cut(s)	439, 670
BciT130I	CCWGG	1 cut(s)	167
BclI	TGATCA	1 cut(s)	442
BfaI	CTAG	6 cut(s)	642, 959, 1349, 1469, 1494, 1502
BfmI	CTRYAG	3 cut(s)	590, 1152, 1380
BisI	GCNGC	2 cut(s)	1175, 1380
BlsI	GCNGC	2 cut(s)	1176, 1381
BmcAI	AGTACT	1 cut(s)	1370
Bme1390I	CCNGG	1 cut(s)	167
Bme18I	GGWCC	2 cut(s)	863, 1299
BmgT120I	GGNCC	2 cut(s)	863, 1299
BmiI	GGNNCC	3 cut(s)	822, 1433, 1534
BmrFI	CCNGG	1 cut(s)	167
BmsI	GCATC	1 cut(s)	1337
BpmI	CTGGAG	2 cut(s)	513, 528
Bpu10I	CCTNAGC	3 cut(s)	225, 343, 516
BpuEI	CTTGAG	1 cut(s)	1189
BsaBI	GATNNNNATC	1 cut(s)	1025
BsaJI	CCNNGG	6 cut(s)	43, 165, 166, 414, 991, 1203
BsaWI	WCCGGW	1 cut(s)	665
Bse1I	ACTGG	2 cut(s)	545, 1478
Bse3DI	GCAATG	1 cut(s)	805
Bse8I	GATNNNNATC	1 cut(s)	1025
BseAI	TCCGGA	1 cut(s)	665
BseBI	CCWGG	1 cut(s)	167
BseDI	CCNNGG	6 cut(s)	43, 165, 166, 414, 991, 1203
BseGI	GGATG	3 cut(s)	431, 662, 1328
BseJI	GATNNNNATC	1 cut(s)	1025
BseMI	GCAATG	1 cut(s)	805
BseMII	CTCAG	4 cut(s)	216, 357, 530, 1153
BseNI	ACTGG	2 cut(s)	545, 1478
BseRI	GAGGAG	1 cut(s)	343
BseSI	GKGCMC	1 cut(s)	1134
BseXI	GCAGC	2 cut(s)	1161, 1366
Bsh1236I	CGCG	2 cut(s)	416, 1097
BshFI	GGCC	1 cut(s)	1081
BshNI	GGYRCC	1 cut(s)	1431
BsiHKAI	GWGCWC	2 cut(s)	175, 626
BsiSI	CCGG	1 cut(s)	666
BsmI	GAATGC	4 cut(s)	152, 424, 1013, 1054
BsnI	GGCC	1 cut(s)	1081
Bsp1286I	GDGCHC	3 cut(s)	175, 626, 1134
Bsp13I	TCCGGA	1 cut(s)	665
Bsp1407I	TGTACA	1 cut(s)	130
Bsp143I	GATC	6 cut(s)	442, 501, 550, 820, 1476, 1543
BspACI	CCGC	4 cut(s)	414, 416, 895, 1265
BspANI	GGCC	1 cut(s)	1081
BspCNI	CTCAG	4 cut(s)	217, 356, 529, 1152
BspEI	TCCGGA	1 cut(s)	665
BspFNI	CGCG	2 cut(s)	416, 1097
BspHI	TCATGA	1 cut(s)	726
BspLI	GGNNCC	3 cut(s)	822, 1433, 1534
BspMAI	CTGCAG	1 cut(s)	1384
BspPI	GGATC	6 cut(s)	496, 545, 815, 828, 1484, 1538
BspT107I	GGYRCC	1 cut(s)	1431
BsrBI	CCGCTC	1 cut(s)	897
BsrDI	GCAATG	1 cut(s)	805
BsrGI	TGTACA	1 cut(s)	130
BsrI	ACTGG	2 cut(s)	545, 1478
BssECI	CCNNGG	6 cut(s)	43, 165, 166, 414, 991, 1203
BssMI	GATC	6 cut(s)	442, 501, 550, 820, 1476, 1543
BssT1I	CCWWGG	2 cut(s)	43, 1203
Bst2UI	CCWGG	1 cut(s)	167
Bst4CI	ACNGT	2 cut(s)	594, 1557
Bst6I	CTCTTC	3 cut(s)	55, 117, 237
BstAUI	TGTACA	1 cut(s)	130
BstC8I	GCNNGC	4 cut(s)	895, 922, 1011, 1377
BstDEI	CTNAG	7 cut(s)	113, 225, 334, 343, 516, 1057, 1139
BstDSI	CCRYGG	1 cut(s)	414
BstF5I	GGATG	3 cut(s)	431, 662, 1328
BstFNI	CGCG	2 cut(s)	416, 1097
BstHHI	GCGC	1 cut(s)	1099
BstKTI	GATC	6 cut(s)	445, 504, 553, 823, 1479, 1546
BstMBI	GATC	6 cut(s)	442, 501, 550, 820, 1476, 1543
BstMWI	GCNNNNNNNGC	3 cut(s)	230, 717, 890
BstNI	CCWGG	1 cut(s)	167
BstNSI	RCATGY	1 cut(s)	884
BstSCI	CCNGG	1 cut(s)	165
BstSFI	CTRYAG	3 cut(s)	590, 1152, 1380
BstSLI	GKGCMC	1 cut(s)	1134
BstUI	CGCG	2 cut(s)	416, 1097
BstV1I	GCAGC	2 cut(s)	1161, 1366
BstX2I	RGATCY	2 cut(s)	820, 1476
BstXI	CCANNNNNNTGG	1 cut(s)	816
BstYI	RGATCY	2 cut(s)	820, 1476
BsuRI	GGCC	1 cut(s)	1081
BtgI	CCRYGG	1 cut(s)	414
BtsCI	GGATG	3 cut(s)	431, 662, 1328
BtsI	GCAGTG	1 cut(s)	1323
BtsIMutI	CAGTG	3 cut(s)	405, 1147, 1323
Cac8I	GCNNGC	4 cut(s)	895, 922, 1011, 1377
CciI	TCATGA	1 cut(s)	726
CfoI	GCGC	1 cut(s)	1099
Cfr13I	GGNCC	2 cut(s)	863, 1299
Cfr42I	CCGCGG	1 cut(s)	417
CseI	GACGC	1 cut(s)	42
Csp6I	GTAC	3 cut(s)	131, 1369, 1432
CviQI	GTAC	3 cut(s)	131, 1369, 1432
DdeI	CTNAG	7 cut(s)	113, 225, 334, 343, 516, 1057, 1139
DpnI	GATC	6 cut(s)	444, 503, 552, 822, 1478, 1545
DpnII	GATC	6 cut(s)	442, 501, 550, 820, 1476, 1543
Eam1104I	CTCTTC	3 cut(s)	55, 117, 237
EarI	CTCTTC	3 cut(s)	55, 117, 237
Ecl136II	GAGCTC	1 cut(s)	173
Eco130I	CCWWGG	2 cut(s)	43, 1203
Eco147I	AGGCCT	1 cut(s)	1081
Eco24I	GRGCYC	1 cut(s)	175
Eco47I	GGWCC	2 cut(s)	863, 1299
Eco53kI	GAGCTC	1 cut(s)	173
Eco57I	CTGAAG	3 cut(s)	122, 725, 1469
EcoICRI	GAGCTC	1 cut(s)	173
EcoRII	CCWGG	1 cut(s)	165
EcoT14I	CCWWGG	2 cut(s)	43, 1203
EcoT38I	GRGCYC	1 cut(s)	175
ErhI	CCWWGG	2 cut(s)	43, 1203
FalI	AAGNNNNNCTT	8 cut(s)	317, 349, 482, 514, 621, 653, 993, 1025
FauI	CCCGC	1 cut(s)	902
FbaI	TGATCA	1 cut(s)	442
FblI	GTMKAC	1 cut(s)	184
Fnu4HI	GCNGC	2 cut(s)	1175, 1380
FokI	GGATG	3 cut(s)	418, 649, 1315
FriOI	GRGCYC	1 cut(s)	175
Fsp4HI	GCNGC	2 cut(s)	1175, 1380
FspBI	CTAG	6 cut(s)	642, 959, 1349, 1469, 1494, 1502
GlaI	GCGC	1 cut(s)	1098
GluI	GCNGC	2 cut(s)	1175, 1380
GsuI	CTGGAG	2 cut(s)	513, 528
HaeIII	GGCC	1 cut(s)	1081
HapII	CCGG	1 cut(s)	666
HgaI	GACGC	1 cut(s)	42
HhaI	GCGC	1 cut(s)	1099
Hin6I	GCGC	1 cut(s)	1097
HinP1I	GCGC	1 cut(s)	1097
HincII	GTYRAC	2 cut(s)	310, 1339
HindII	GTYRAC	2 cut(s)	310, 1339
HindIII	AAGCTT	3 cut(s)	110, 1007, 1418
HinfI	GANTC	7 cut(s)	359, 376, 457, 526, 578, 786, 1460
HpaII	CCGG	1 cut(s)	666
HphI	GGTGA	1 cut(s)	430
Hpy166II	GTNNAC	7 cut(s)	133, 185, 310, 403, 1321, 1339, 1499
Hpy188I	TCNGA	7 cut(s)	442, 555, 694, 705, 841, 941, 1261
Hpy188III	TCNNGA	9 cut(s)	57, 299, 650, 666, 727, 1001, 1018, 1168, 1526
Hpy8I	GTNNAC	7 cut(s)	133, 185, 310, 403, 1321, 1339, 1499
HpyAV	CCTTC	7 cut(s)	280, 447, 777, 1102, 1207, 1493, 1523
HpyCH4III	ACNGT	2 cut(s)	594, 1557
HpyF10VI	GCNNNNNNNGC	3 cut(s)	230, 717, 890
HpyF3I	CTNAG	7 cut(s)	113, 225, 334, 343, 516, 1057, 1139
HspAI	GCGC	1 cut(s)	1097
Kpn2I	TCCGGA	1 cut(s)	665
KpnI	GGTACC	1 cut(s)	1435
Ksp22I	TGATCA	1 cut(s)	442
KspI	CCGCGG	1 cut(s)	417
Kzo9I	GATC	6 cut(s)	442, 501, 550, 820, 1476, 1543
LmnI	GCTCC	4 cut(s)	170, 330, 942, 1532
Lsp1109I	GCAGC	2 cut(s)	1161, 1366
LweI	GCATC	1 cut(s)	1337
MaeI	CTAG	6 cut(s)	642, 959, 1349, 1469, 1494, 1502
MaeIII	GTNAC	2 cut(s)	607, 1270
MalI	GATC	6 cut(s)	444, 503, 552, 822, 1478, 1545
MbiI	CCGCTC	1 cut(s)	897
MboI	GATC	6 cut(s)	442, 501, 550, 820, 1476, 1543
MboII	GAAGA	9 cut(s)	72, 115, 118, 134, 254, 275, 493, 954, 1589
MflI	RGATCY	2 cut(s)	820, 1476
MhlI	GDGCHC	3 cut(s)	175, 626, 1134
MlyI	GAGTC	2 cut(s)	520, 1469
MmeI	TCCRAC	2 cut(s)	814, 1366
MroI	TCCGGA	1 cut(s)	665
MroXI	GAANNNNTTC	4 cut(s)	252, 699, 736, 1052
MseI	TTAA	2 cut(s)	903, 1023
MslI	CAYNNNNRTG	3 cut(s)	510, 725, 731
MspA1I	CMGCKG	3 cut(s)	347, 416, 1267
MspI	CCGG	1 cut(s)	666
MspR9I	CCNGG	1 cut(s)	167
Mva1269I	GAATGC	4 cut(s)	152, 424, 1013, 1054
MvaI	CCWGG	1 cut(s)	167
MvnI	CGCG	2 cut(s)	416, 1097
MwoI	GCNNNNNNNGC	3 cut(s)	230, 717, 890
NdeII	GATC	6 cut(s)	442, 501, 550, 820, 1476, 1543
NlaIV	GGNNCC	3 cut(s)	822, 1433, 1534
NspI	RCATGY	1 cut(s)	884
PagI	TCATGA	1 cut(s)	726
PasI	CCCWGGG	1 cut(s)	166
PceI	AGGCCT	1 cut(s)	1081
PctI	GAATGC	4 cut(s)	152, 424, 1013, 1054
PdmI	GAANNNNTTC	4 cut(s)	252, 699, 736, 1052
PfeI	GAWTC	5 cut(s)	359, 376, 457, 578, 786
PkrI	GCNGC	2 cut(s)	1176, 1381
PleI	GAGTC	2 cut(s)	520, 1468
PpsI	GAGTC	2 cut(s)	520, 1468
PshBI	ATTAAT	2 cut(s)	903, 1023
Psp124BI	GAGCTC	1 cut(s)	175
Psp6I	CCWGG	1 cut(s)	165
PspGI	CCWGG	1 cut(s)	165
PspN4I	GGNNCC	3 cut(s)	822, 1433, 1534
PspPI	GGNCC	2 cut(s)	863, 1299
PsrI	GAACNNNNNNTAC	2 cut(s)	1053, 1085
PstI	CTGCAG	1 cut(s)	1384
PsuI	RGATCY	2 cut(s)	820, 1476
PvuII	CAGCTG	1 cut(s)	347
RsaI	GTAC	3 cut(s)	132, 1370, 1433
RsaNI	GTAC	3 cut(s)	131, 1369, 1432
RseI	CAYNNNNRTG	3 cut(s)	510, 725, 731
SacI	GAGCTC	1 cut(s)	175
SacII	CCGCGG	1 cut(s)	417
SaqAI	TTAA	2 cut(s)	903, 1023
SatI	GCNGC	2 cut(s)	1175, 1380
Sau3AI	GATC	6 cut(s)	442, 501, 550, 820, 1476, 1543
Sau96I	GGNCC	2 cut(s)	863, 1299
ScaI	AGTACT	1 cut(s)	1370
SchI	GAGTC	2 cut(s)	520, 1469
ScrFI	CCNGG	1 cut(s)	167
SduI	GDGCHC	3 cut(s)	175, 626, 1134
SfaNI	GCATC	1 cut(s)	1337
SfcI	CTRYAG	3 cut(s)	590, 1152, 1380
Sfr303I	CCGCGG	1 cut(s)	417
SgrBI	CCGCGG	1 cut(s)	417
SinI	GGWCC	2 cut(s)	863, 1299
SmiMI	CAYNNNNRTG	3 cut(s)	510, 725, 731
SmlI	CTYRAG	1 cut(s)	1168
SmoI	CTYRAG	1 cut(s)	1168
SseBI	AGGCCT	1 cut(s)	1081
SsiI	CCGC	4 cut(s)	414, 416, 895, 1265
SspI	AATATT	1 cut(s)	700
SspMI	CTAG	6 cut(s)	642, 959, 1349, 1469, 1494, 1502
SstI	GAGCTC	1 cut(s)	175
StuI	AGGCCT	1 cut(s)	1081
StyD4I	CCNGG	1 cut(s)	165
StyI	CCWWGG	2 cut(s)	43, 1203
TaaI	ACNGT	2 cut(s)	594, 1557
TaqI	TCGA	3 cut(s)	317, 913, 1397
TaqII	GACCGA	1 cut(s)	983
TatI	WGTACW	2 cut(s)	130, 1368
TfiI	GAWTC	5 cut(s)	359, 376, 457, 578, 786
Tru1I	TTAA	2 cut(s)	903, 1023
Tru9I	TTAA	2 cut(s)	903, 1023
TscAI	CASTG	3 cut(s)	412, 1147, 1330
TseI	GCWGC	2 cut(s)	1174, 1379
TspDTI	ATGAA	8 cut(s)	135, 377, 494, 715, 816, 1061, 1359, 1383
TspGWI	ACGGA	2 cut(s)	770, 865
TspRI	CASTG	3 cut(s)	412, 1147, 1330
VpaK11BI	GGWCC	2 cut(s)	863, 1299
VspI	ATTAAT	2 cut(s)	903, 1023
XapI	RAATTY	5 cut(s)	383, 812, 842, 979, 1389
XceI	RCATGY	1 cut(s)	884
XmiI	GTMKAC	1 cut(s)	184
XmnI	GAANNNNTTC	4 cut(s)	252, 699, 736, 1052
XspI	CTAG	6 cut(s)	642, 959, 1349, 1469, 1494, 1502
ZrmI	AGTACT	1 cut(s)	1370

Using CommOnly database (standard laboratory enzymes). Scanned on CDS sequence.