Rh5AG334700

Protein LATERAL ROOT PRIMORDIUM

Design Primers CRISPR Design

Basic Information

Type: gene
Biological Identity
rosa_samantha
Chr5A
OG0009808
Physical Location & Seq
Reverse (-)
53665806 .. 53667209
1404 bp
Loading structure...
UTR
Exon/CDS
Intron
Rh5AG334700.1

Sequence Viewer

Length: 1236 bp
ATGTGGTCGTCATCATCAGGCCCTTCCAGGCCCATCAACTACGGCTTCCCACCGGAGATGGGTATGCTCGGCCTTCGAGACGTTTTCCTTGTTGCTCCAGCTGCAGCCTACCACAATCACCCGCACCACCACCACCACCCCCCCCACCATCAACAAGACCTTCTCTCATCTGATCATAATTCCATGAATGGCTCTAATCCCGCCACAGCTCTCGGGGTCGGCGTCATTCCACTCCTCACCACCACCCCTTGCCTGGGCCCTCCCAATACCAACATGAATGATGACAACAATGATGATGATGATGATCACAACATGCTGAGCGGTCGCAACAGAAACAGTGCAGGTGGAGGAGGAGGAGGAGGAGGAGGTATTCACTTGTGGCAGCAGCCGACCCAGCAACAACACCAACTTTTCCTCAAGAGAAACCAGAACCTTCACGGTGGCGCCCATTTGATGGTTCAAGGTGGTGGTGGGAATTTTGGTGCCTCGGCGGCATCGGCATCGGCATCGGCATCGTCCGCTCTCACTTGTCAAGACTGCGGGAACCAGGCCAAGAAAGACTGCAGTCATCGGCGGTGCAGGACATGCTGCAAGAGCCGTGGTTTTGATTGTTCCACGCACGTGAAGAGCACGTGGGTCCCAGCGGCTAGACGCAGGGAGCGGCAGCTCATGTCCGAGGGAGTGGCGGCGGCCGGGTCTTCTGCGTCTACCTCGCACACTAATAATACTAGTTCCAATGCCAAAAAACCAAGACTGGTTACTTCGCAAAACACAACTCCCTCTCATACGAACTCCACTTCAAACACAACGCCACCCAGAAGCTATGACAGTACCTCCAAACTCCAACACCAAGATGGTGGTTTTCCAGAGACAATGCCGGGCCAAGTACGGGCGGCAGCGGTTTTTAAGTGCGTTCGAGTAACGGCCGTGGATGATGGTGCAGAGGGAGAGTATGCCTATCACGCTGTCGTCAAAATCAACGGTCATGTGTTTAAGGGGTTCTTGTATGATCAAGGGGTTGAAGGTCAAGGTAGAGATCATCATGGCTTCCCAAATATTTCTGATTTGCATTTGGGTGGTAGAAGCGGAGGTGCCACTAGTACCCCCCCAACTGGAAGAAATGGAGGTGCTCCAATACTCGACCCCGCGGATGCTTATGCAGCCGCTGCAGCAGCGGCCTCCACTAGTGGAGGACTGCTGCTGGGAGTGTCAAACTTTGGTAACTCAATAAATTAA
Functional Annotation
Gene Ontology
Molecular Function
Biological Process
Cellular Component
No ontology terms assigned.
KEGG Pathways
Metabolic & Signaling
No pathways identified.
Pfam Domains
Protein Families
DUF702

Protein Analysis

411

Amino Acids

42.94

Weight (kDa)

8.11

Isoelectric Point (pI)

40.66

Instability Index
Protein Domains (Pfam)
Domain Name Pfam ID Position E-value Description
DUF702 PF05142 175 - 341 1.9e-59 Domain of unknown function (DUF702)
Hydrophobicity Profile (Kyte-Doolittle)
AI Structure Prediction Report
Calculating structure properties...

Genomic Context

Gene Family Tree

Species Filter
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Publication-ready
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Orthologous Genes (Group: OG0009808)

Species Orthologous Gene IDs
arabidopsis_thaliana AT5G12330 AT5G12330 AT5G12330 AT5G12330
fragaria_vesca FvH4_3g27920
malus_domestica MD03G1166900.v1.1 MD11G1186100.v1.1
prunus_persica Prupe.4G267200_v2.0.a1
pyrus_communis pycom03g12230 pycom12g08680
rosa_chinensis RchiOBHm_Chr5g0050641
rosa_laevigata RLG00000034715
rosa_multiflora Rmu_ssc0000350.1_g000034
rosa_roxburghii Rroxscaffold_1G00030150
rosa_rugosa Rorug05G0262400 Rorug05G0262500
rosa_samantha Rh5AG334700 Rh5BG344000 Rh5CG371800 Rh5DG357000
rosa_wichuraiana Rw5G031570

Restriction Enzyme Sites

1 / 10
Enzyme Recognition Site Cut Count Positions (bp)
AarI CACCTGC 1 cut(s) 332
Acc36I ACCTGC 1 cut(s) 332
AccB1I GGYRCC 3 cut(s) 443, 482, 1091
AccB7I CCANNNNNTGG 1 cut(s) 454
AccBSI CCGCTC 3 cut(s) 321, 521, 661
AccI GTMKAC 1 cut(s) 707
AccII CGCG 1 cut(s) 1148
AcoI YGGCCR 2 cut(s) 690, 924
AcsI RAATTY 1 cut(s) 475
AcvI CACGTG 2 cut(s) 622, 633
AcyI GRCGYC 2 cut(s) 222, 444
AfaI GTAC 3 cut(s) 832, 888, 1102
AfiI CCNNNNNNNGG 6 cut(s) 59, 253, 254, 454, 889, 1112
AgsI TTSAA 3 cut(s) 461, 801, 1022
AhlI ACTAGT 3 cut(s) 728, 1097, 1184
AjnI CCWGG 3 cut(s) 26, 252, 546
AleI CACNNNNGTG 1 cut(s) 620
AluBI AGCT 4 cut(s) 101, 209, 667, 822
AluI AGCT 4 cut(s) 101, 209, 667, 822
Alw21I GWGCWC 2 cut(s) 632, 1132
Alw26I GTCTC 2 cut(s) 72, 863
AlwNI CAGNNNCTG 1 cut(s) 1166
Ama87I CYCGRG 1 cut(s) 212
AoxI GGCC 9 cut(s) 19, 29, 70, 256, 549, 690, 880, 924, 1176
ApaI GGGCCC 1 cut(s) 260
ApoI RAATTY 1 cut(s) 475
AspLEI GCGC 1 cut(s) 446
AspS9I GGNCC 6 cut(s) 20, 30, 256, 257, 637, 880
AsuC2I CCSGG 2 cut(s) 694, 879
AsuHPI GGTGA 2 cut(s) 110, 229
AvaI CYCGRG 1 cut(s) 212
AvaII GGWCC 1 cut(s) 637
BaeGI GKGCMC 1 cut(s) 260
BanI GGYRCC 3 cut(s) 443, 482, 1091
BanII GRGCYC 1 cut(s) 260
BbrPI CACGTG 2 cut(s) 622, 633
BbsI GAAGAC 1 cut(s) 690
Bbv12I GWGCWC 2 cut(s) 632, 1132
BccI CCATC 6 cut(s) 41, 52, 156, 448, 848, 929
BceAI ACGGC 4 cut(s) 58, 582, 911, 939
BcgI CGANNNNNNTGC 6 cut(s) 483, 489, 495, 517, 523, 529
BciT130I CCWGG 3 cut(s) 28, 254, 548
BclI TGATCA 3 cut(s) 172, 304, 1009
BcnI CCSGG 2 cut(s) 694, 879
BcoDI GTCTC 2 cut(s) 72, 863
BcuI ACTAGT 3 cut(s) 728, 1097, 1184
BfaI CTAG 4 cut(s) 648, 729, 1098, 1185
BfmI CTRYAG 3 cut(s) 102, 562, 1167
BfoI RGCGCY 1 cut(s) 447
BfuAI ACCTGC 1 cut(s) 332
BglI GCCNNNNNGGC 1 cut(s) 491
BlpI GCTNAGC 1 cut(s) 317
Bme1390I CCNGG 5 cut(s) 28, 254, 548, 694, 879
Bme18I GGWCC 1 cut(s) 637
BmeT110I CYCGRG 1 cut(s) 212
BmgT120I GGNCC 6 cut(s) 20, 30, 256, 257, 637, 880
BmiI GGNNCC 7 cut(s) 258, 445, 484, 545, 638, 639, 1093
BmrFI CCNGG 5 cut(s) 28, 254, 548, 694, 879
BmsI GCATC 5 cut(s) 503, 509, 515, 521, 1141
BoxI GACNNNNGTC 1 cut(s) 564
BpiI GAAGAC 1 cut(s) 690
BpmI CTGGAG 1 cut(s) 81
Bpu1102I GCTNAGC 1 cut(s) 317
BpuEI CTTGAG 1 cut(s) 401
BpuMI CCSGG 2 cut(s) 694, 879
BsaAI YACGTR 2 cut(s) 622, 633
BsaBI GATNNNNATC 1 cut(s) 303
BsaHI GRCGYC 2 cut(s) 222, 444
BsaJI CCNNGG 6 cut(s) 253, 486, 598, 675, 927, 1146
BsaWI WCCGGW 1 cut(s) 52
BsaXI ACNNNNNCTCC 2 cut(s) 818, 848
Bsc4I CCNNNNNNNGG 6 cut(s) 59, 253, 254, 454, 889, 1112
Bse1I ACTGG 2 cut(s) 759, 1117
Bse8I GATNNNNATC 1 cut(s) 303
BseBI CCWGG 3 cut(s) 28, 254, 548
BseDI CCNNGG 6 cut(s) 253, 486, 598, 675, 927, 1146
BseGI GGATG 2 cut(s) 937, 1156
BseJI GATNNNNATC 1 cut(s) 303
BseLI CCNNNNNNNGG 6 cut(s) 59, 253, 254, 454, 889, 1112
BseMII CTCAG 1 cut(s) 308
BseNI ACTGG 2 cut(s) 759, 1117
BseRI GAGGAG 7 cut(s) 224, 363, 366, 369, 372, 375, 378
BseSI GKGCMC 1 cut(s) 260
BseX3I CGGCCG 2 cut(s) 690, 924
BseYI CCCAGC 3 cut(s) 393, 640, 1201
BsgI GTGCAG 3 cut(s) 360, 598, 960
Bsh1236I CGCG 1 cut(s) 1148
Bsh1285I CGRYCG 3 cut(s) 325, 693, 927
BshFI GGCC 9 cut(s) 21, 31, 72, 258, 551, 692, 882, 926, 1178
BshNI GGYRCC 3 cut(s) 443, 482, 1091
BsiEI CGRYCG 3 cut(s) 325, 693, 927
BsiHKAI GWGCWC 2 cut(s) 632, 1132
BsiHKCI CYCGRG 1 cut(s) 212
BsiSI CCGG 3 cut(s) 53, 693, 878
BslFI GGGAC 1 cut(s) 623
BslI CCNNNNNNNGG 6 cut(s) 59, 253, 254, 454, 889, 1112
BsmAI GTCTC 2 cut(s) 72, 863
BsmBI CGTCTC 1 cut(s) 72
BsmFI GGGAC 1 cut(s) 623
BsnI GGCC 9 cut(s) 21, 31, 72, 258, 551, 692, 882, 926, 1178
BsoBI CYCGRG 1 cut(s) 212
Bsp120I GGGCCC 1 cut(s) 256
Bsp1286I GDGCHC 3 cut(s) 260, 632, 1132
Bsp143I GATC 4 cut(s) 172, 304, 1009, 1036
Bsp1720I GCTNAGC 1 cut(s) 317
BspANI GGCC 9 cut(s) 21, 31, 72, 258, 551, 692, 882, 926, 1178
BspCNI CTCAG 1 cut(s) 309
BspFNI CGCG 1 cut(s) 1148
BspLI GGNNCC 7 cut(s) 258, 445, 484, 545, 638, 639, 1093
BspMAI CTGCAG 3 cut(s) 106, 566, 1171
BspMI ACCTGC 1 cut(s) 332
BspQI GCTCTTC 1 cut(s) 620
BspT107I GGYRCC 3 cut(s) 443, 482, 1091
BsrBI CCGCTC 3 cut(s) 321, 521, 661
BsrI ACTGG 2 cut(s) 759, 1117
BssECI CCNNGG 6 cut(s) 253, 486, 598, 675, 927, 1146
BssMI GATC 4 cut(s) 172, 304, 1009, 1036
BssNI GRCGYC 2 cut(s) 222, 444
Bst2UI CCWGG 3 cut(s) 28, 254, 548
Bst4CI ACNGT 4 cut(s) 338, 440, 830, 983
Bst6I CTCTTC 1 cut(s) 620
BstACI GRCGYC 2 cut(s) 222, 444
BstAPI GCANNNNNTGC 2 cut(s) 585, 1166
BstBAI YACGTR 2 cut(s) 622, 633
BstDEI CTNAG 1 cut(s) 317
BstDSI CCRYGG 3 cut(s) 598, 927, 1146
BstF5I GGATG 2 cut(s) 937, 1156
BstFNI CGCG 1 cut(s) 1148
BstH2I RGCGCY 1 cut(s) 447
BstHHI GCGC 1 cut(s) 446
BstKTI GATC 4 cut(s) 175, 307, 1012, 1039
BstMAI GTCTC 2 cut(s) 72, 863
BstMBI GATC 4 cut(s) 172, 304, 1009, 1036
BstMCI CGRYCG 3 cut(s) 325, 693, 927
BstNI CCWGG 3 cut(s) 28, 254, 548
BstNSI RCATGY 2 cut(s) 316, 588
BstPAI GACNNNNGTC 1 cut(s) 564
BstSCI CCNGG 5 cut(s) 26, 252, 546, 692, 877
BstSFI CTRYAG 3 cut(s) 102, 562, 1167
BstSLI GKGCMC 1 cut(s) 260
BstUI CGCG 1 cut(s) 1148
BstV2I GAAGAC 1 cut(s) 690
BstXI CCANNNNNNTGG 1 cut(s) 857
BstZI CGGCCG 2 cut(s) 690, 924
BsuRI GGCC 9 cut(s) 21, 31, 72, 258, 551, 692, 882, 926, 1178
BtgI CCRYGG 3 cut(s) 598, 927, 1146
BtsCI GGATG 2 cut(s) 937, 1156
BtsIMutI CAGTG 1 cut(s) 343
BveI ACCTGC 1 cut(s) 332
CaiI CAGNNNCTG 1 cut(s) 1166
CfoI GCGC 1 cut(s) 446
Cfr13I GGNCC 6 cut(s) 20, 30, 256, 257, 637, 880
Cfr42I CCGCGG 1 cut(s) 1149
CseI GACGC 3 cut(s) 211, 660, 693
Csp6I GTAC 3 cut(s) 831, 887, 1101
CspCI CAANNNNNGTGG 2 cut(s) 580, 615
CviAII CATG 7 cut(s) 184, 274, 313, 585, 670, 986, 1043
CviQI GTAC 3 cut(s) 831, 887, 1101
DdeI CTNAG 1 cut(s) 317
DinI GGCGCC 1 cut(s) 445
DpnI GATC 4 cut(s) 174, 306, 1011, 1038
DpnII GATC 4 cut(s) 172, 304, 1009, 1036
EaeI YGGCCR 2 cut(s) 690, 924
EagI CGGCCG 2 cut(s) 690, 924
Eam1104I CTCTTC 1 cut(s) 620
EarI CTCTTC 1 cut(s) 620
EclXI CGGCCG 2 cut(s) 690, 924
Eco24I GRGCYC 1 cut(s) 260
Eco47I GGWCC 1 cut(s) 637
Eco52I CGGCCG 2 cut(s) 690, 924
Eco72I CACGTG 2 cut(s) 622, 633
Eco88I CYCGRG 1 cut(s) 212
EcoO109I RGGNCCY 3 cut(s) 20, 257, 637
EcoRII CCWGG 3 cut(s) 26, 252, 546
EcoT38I GRGCYC 1 cut(s) 260
EgeI GGCGCC 1 cut(s) 445
EheI GGCGCC 1 cut(s) 445
Esp3I CGTCTC 1 cut(s) 72
FaeI CATG 7 cut(s) 187, 277, 316, 588, 673, 989, 1046
FalI AAGNNNNNCTT 2 cut(s) 986, 1018
FaqI GGGAC 1 cut(s) 623
FatI CATG 7 cut(s) 183, 273, 312, 584, 669, 985, 1042
FauI CCCGC 4 cut(s) 129, 208, 533, 1153
FbaI TGATCA 3 cut(s) 172, 304, 1009
FblI GTMKAC 1 cut(s) 707
FokI GGATG 2 cut(s) 944, 1163
FriOI GRGCYC 1 cut(s) 260
FspBI CTAG 4 cut(s) 648, 729, 1098, 1185
GlaI GCGC 1 cut(s) 445
GsaI CCCAGC 3 cut(s) 397, 644, 1205
GsuI CTGGAG 1 cut(s) 81
HaeII RGCGCY 1 cut(s) 447
HaeIII GGCC 9 cut(s) 21, 31, 72, 258, 551, 692, 882, 926, 1178
HapII CCGG 3 cut(s) 53, 693, 878
HgaI GACGC 3 cut(s) 211, 660, 693
HhaI GCGC 1 cut(s) 446
Hin1I GRCGYC 2 cut(s) 222, 444
Hin1II CATG 7 cut(s) 187, 277, 316, 588, 673, 989, 1046
Hin6I GCGC 1 cut(s) 444
HinP1I GCGC 1 cut(s) 444
HpaII CCGG 3 cut(s) 53, 693, 878
HphI GGTGA 2 cut(s) 110, 229
Hpy166II GTNNAC 1 cut(s) 708
Hpy188I TCNGA 3 cut(s) 172, 676, 1063
Hpy188III TCNNGA 4 cut(s) 77, 418, 533, 866
Hpy8I GTNNAC 1 cut(s) 708
HpyAV CCTTC 5 cut(s) 33, 83, 170, 443, 1016
HpyCH4III ACNGT 4 cut(s) 338, 440, 830, 983
HpyCH4IV ACGT 3 cut(s) 81, 621, 632
HpyCH4V TGCA 9 cut(s) 104, 341, 564, 579, 591, 941, 1069, 1160, 1169
HpyF3I CTNAG 1 cut(s) 317
HpySE526I ACGT 3 cut(s) 81, 621, 632
Hsp92I GRCGYC 2 cut(s) 222, 444
Hsp92II CATG 7 cut(s) 187, 277, 316, 588, 673, 989, 1046
HspAI GCGC 1 cut(s) 444
KasI GGCGCC 1 cut(s) 443
KflI GGGWCCC 1 cut(s) 637
Ksp22I TGATCA 3 cut(s) 172, 304, 1009
KspI CCGCGG 1 cut(s) 1149
Kzo9I GATC 4 cut(s) 172, 304, 1009, 1036
LguI GCTCTTC 1 cut(s) 620
LmnI GCTCC 3 cut(s) 100, 658, 1135
LweI GCATC 5 cut(s) 503, 509, 515, 521, 1141
MaeI CTAG 4 cut(s) 648, 729, 1098, 1185
MaeII ACGT 3 cut(s) 81, 621, 632
MaeIII GTNAC 3 cut(s) 757, 919, 1220
MalI GATC 4 cut(s) 174, 306, 1011, 1038
MbiI CCGCTC 3 cut(s) 321, 521, 661
MboI GATC 4 cut(s) 172, 304, 1009, 1036
MboII GAAGA 3 cut(s) 637, 690, 1128
MhlI GDGCHC 3 cut(s) 260, 632, 1132
MluCI AATT 3 cut(s) 178, 475, 1231
Mly113I GGCGCC 1 cut(s) 444
MmeI TCCRAC 1 cut(s) 868
MseI TTAA 3 cut(s) 906, 993, 1234
MslI CAYNNNNRTG 3 cut(s) 620, 852, 1074
MspA1I CMGCKG 6 cut(s) 101, 644, 899, 1148, 1166, 1175
MspI CCGG 3 cut(s) 53, 693, 878
MspR9I CCNGG 5 cut(s) 28, 254, 548, 694, 879
MvaI CCWGG 3 cut(s) 28, 254, 548
MvnI CGCG 1 cut(s) 1148
NarI GGCGCC 1 cut(s) 444
NciI CCSGG 2 cut(s) 694, 879
NdeII GATC 4 cut(s) 172, 304, 1009, 1036
NlaIII CATG 7 cut(s) 187, 277, 316, 588, 673, 989, 1046
NlaIV GGNNCC 7 cut(s) 258, 445, 484, 545, 638, 639, 1093
NmeAIII GCCGAG 2 cut(s) 48, 467
NspI RCATGY 2 cut(s) 316, 588
OliI CACNNNNGTG 1 cut(s) 620
PaqCI CACCTGC 1 cut(s) 332
PciSI GCTCTTC 1 cut(s) 620
PcsI WCGNNNNNNNCGW 1 cut(s) 219
PflMI CCANNNNNTGG 1 cut(s) 454
PluTI GGCGCC 1 cut(s) 447
PmaCI CACGTG 2 cut(s) 622, 633
PmlI CACGTG 2 cut(s) 622, 633
Ppu21I YACGTR 2 cut(s) 622, 633
PpuMI RGGWCCY 1 cut(s) 637
PshAI GACNNNNGTC 1 cut(s) 564
Psp5II RGGWCCY 1 cut(s) 637
Psp6I CCWGG 3 cut(s) 26, 252, 546
PspCI CACGTG 2 cut(s) 622, 633
PspFI CCCAGC 3 cut(s) 393, 640, 1201
PspGI CCWGG 3 cut(s) 26, 252, 546
PspN4I GGNNCC 7 cut(s) 258, 445, 484, 545, 638, 639, 1093
PspOMI GGGCCC 1 cut(s) 256
PspPI GGNCC 6 cut(s) 20, 30, 256, 257, 637, 880
PspPPI RGGWCCY 1 cut(s) 637
PstI CTGCAG 3 cut(s) 106, 566, 1171
PstNI CAGNNNCTG 1 cut(s) 1166
PvuII CAGCTG 1 cut(s) 101
RsaI GTAC 3 cut(s) 832, 888, 1102
RsaNI GTAC 3 cut(s) 831, 887, 1101
RseI CAYNNNNRTG 3 cut(s) 620, 852, 1074
SacII CCGCGG 1 cut(s) 1149
SapI GCTCTTC 1 cut(s) 620
SaqAI TTAA 3 cut(s) 906, 993, 1234
Sau3AI GATC 4 cut(s) 172, 304, 1009, 1036
Sau96I GGNCC 6 cut(s) 20, 30, 256, 257, 637, 880
ScrFI CCNGG 5 cut(s) 28, 254, 548, 694, 879
SduI GDGCHC 3 cut(s) 260, 632, 1132
SfaNI GCATC 5 cut(s) 503, 509, 515, 521, 1141
SfcI CTRYAG 3 cut(s) 102, 562, 1167
SfoI GGCGCC 1 cut(s) 445
Sfr303I CCGCGG 1 cut(s) 1149
SgrBI CCGCGG 1 cut(s) 1149
SinI GGWCC 1 cut(s) 637
SmiMI CAYNNNNRTG 3 cut(s) 620, 852, 1074
SmlI CTYRAG 1 cut(s) 416
SmoI CTYRAG 1 cut(s) 416
SpeI ACTAGT 3 cut(s) 728, 1097, 1184
Sse9I AATT 3 cut(s) 178, 475, 1231
SspDI GGCGCC 1 cut(s) 443
SspI AATATT 1 cut(s) 1057
SspMI CTAG 4 cut(s) 648, 729, 1098, 1185
StyD4I CCNGG 5 cut(s) 26, 252, 546, 692, 877
TaaI ACNGT 4 cut(s) 338, 440, 830, 983
TaiI ACGT 3 cut(s) 84, 624, 635
TaqI TCGA 3 cut(s) 76, 916, 1140
TasI AATT 3 cut(s) 178, 475, 1231
TauI GCSGC 8 cut(s) 494, 647, 664, 689, 692, 896, 1166, 1178
Tru1I TTAA 3 cut(s) 906, 993, 1234
Tru9I TTAA 3 cut(s) 906, 993, 1234
TscAI CASTG 1 cut(s) 343
TspDTI ATGAA 2 cut(s) 200, 290
TspRI CASTG 1 cut(s) 343
Van91I CCANNNNNTGG 1 cut(s) 454
VpaK11BI GGWCC 1 cut(s) 637
XapI RAATTY 1 cut(s) 475
XceI RCATGY 2 cut(s) 316, 588
XcmI CCANNNNNNNNNTGG 2 cut(s) 250, 851
XmiI GTMKAC 1 cut(s) 707
XspI CTAG 4 cut(s) 648, 729, 1098, 1185
Using CommOnly database (standard laboratory enzymes). Scanned on CDS sequence.