Basic Information

Type: gene

Biological Identity

Species

rosa_roxburghii

Chromosome

GWHEROQ00000002

Orthogroup ID

OG0009480

Physical Location & Seq

Genomic Coordinates Forward (+)

89284127 .. 89287698

Gene Structure 3572 bp

Loading structure...

UTR

Exon/CDS

Intron

Transcript / Protein ID

Rroxscaffold_2G00152240.1

Sequence Viewer

CDS Protein Promoter (2k)

Length: 990 bp

ATGGCTGCCTTTTTTCAGAGACTGGCCAAGGCCGCCCCAACCGCCTTCACTGCTGCGTTCGCCGCCGGCGCCGCTGCCGGCCAATCTAAGCCCGGCTTCCGCTTTCCAATCGGAGCACTCGCGGCTGTCTCCGGCGGAATCTCGTATATCTACTACCAGTCGTCGGCCGATTCGGTTTATCTAGATGAAATCAAAGAACAGGCTGGTCCAAAAATTGCACTTAATCCAGATAAATGGACTGAATTTAAGCTCCAAGACACGGCAAGAGTTAGCCACAATACTCAATTGTTCAGATTTACATTTGACCCAAATGCCAAGTTAGGTTTAGATGTGGCTTCATGCCTCATTACAAGGGCTCCAATTGGACAAGACGCTGAAGGGAAACCAAAATATGTCATACGGCCGTATACTCCTATATCAGATCCAGATTCCAAGGGATACTTTGACTTATTGATTAAGGTGTATCCAGAAGGGAAAATGAGCCAGCATTTTGCAAGCTTAAAACCAGGCGATGTAGTTGAAGTGAAGGGGCCCATTGAAAAGCTGAGATATGTTCCCAACATGAAGAAACATATTGGCATGATTGCAGGTGGTACTGGTATTACTCCAATGCTTCAGGTCATTGAGGCTATACTTAAAAATCCTGATGACTCTACCAAGGTGTCACTGCTTTATGCTAATGTCTCACCAGATGACATACTGCTTAAACAGAAACTTGACATACTTGCAGCTAGCCACCCTAATTTGAAGGTGTTTTACACAGTAGATAATCCAACAAAGAGTTGGAAAGGAGGTAAAGGCTACATATCAAAGGACATGGCAATTAAAGGCTTACCTGGTCCAGCTGAAGATACTCTCATCCTTGTATGCGGTCCTCCTGGAATGATGGAACACATATCTGGTGGCAAGGCTAAAGACTATTCACAAGGAGAGCTCAGAGGCATACTAAAAGAACTTGGTTACACAGAAGAAATGGTTTACAAATTTTGA

Functional Annotation

Gene Ontology

Molecular Function
Biological Process
Cellular Component

GO:0003674 GO:0003824 GO:0004128 GO:0005488 GO:0005507 GO:0005575 GO:0005618 GO:0005622 GO:0005623 GO:0005737 GO:0005739 GO:0005740 GO:0005758 GO:0005794 GO:0006950 GO:0006970 GO:0008150 GO:0008152 GO:0009505 GO:0009628 GO:0009651 GO:0012505 GO:0016491 GO:0016651 GO:0016653 GO:0030312 GO:0031967 GO:0031970 GO:0031974 GO:0031975 GO:0043167 GO:0043169 GO:0043226 GO:0043227 GO:0043229 GO:0043231 GO:0043233 GO:0044422 GO:0044424 GO:0044429 GO:0044444 GO:0044446 GO:0044464 GO:0046872 GO:0046914 GO:0050896 GO:0055114 GO:0070013 GO:0071944

KEGG Pathways

Metabolic & Signaling

ko00520 map00520

Pfam Domains

Protein Families

FAD_binding_6 NAD_binding_1

Protein Analysis

329

Amino Acids

35.73

Weight (kDa)

8.87

Isoelectric Point (pI)

32.76

Instability Index

Protein Domains (Pfam)

Domain Name	Pfam ID	Position	E-value	Description
FAD_binding_6	PF00970	82 - 184	8.4e-26	Oxidoreductase FAD-binding domain
NAD_binding_1	PF00175	194 - 299	5.9e-31	Oxidoreductase NAD-binding domain

Hydrophobicity Profile (Kyte-Doolittle)

AI Structure Prediction Report

Calculating structure properties...

Genomic Context

Gene Family Tree

Species Filter

Style Settings

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Align Labels (Cladogram)

Image

Tree File

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Publication-ready

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Orthologous Genes (Group: OG0009480)

Species	Orthologous Gene IDs
arabidopsis_thaliana	AT5G20080
fragaria_vesca	FvH4_1g03580
malus_domestica	MD00G1107200.v1.1 MD02G1035100.v1.1
prunus_persica	Prupe.7G239800_v2.0.a1
pyrus_communis	pycom02g02870 pycom02g02890 pycom02g02910 pycom15g15760 pycom810g00010 pycom810g00030
rosa_chinensis	RchiOBHm_Chr2g0089111
rosa_laevigata	RLG00000015992
rosa_multiflora	Rmu_sc0003668.1_g000001 Rmu_sc0005510.1_g000001
rosa_roxburghii	Rroxscaffold_2G00152240
rosa_rugosa	Rorug01G0487500
rosa_samantha	Rh2AG043000 Rh2BG041500 Rh2CG043100 Rh2DG042100
rosa_wichuraiana	Rw2G003460

Restriction Enzyme Sites

1 / 10

Enzyme	Recognition Site	Cut Count	Positions (bp)
AarI	CACCTGC	1 cut(s)	578
Acc36I	ACCTGC	1 cut(s)	578
AccB1I	GGYRCC	1 cut(s)	68
AccI	GTMKAC	1 cut(s)	407
AccII	CGCG	1 cut(s)	122
AciI	CCGC	8 cut(s)	33, 42, 63, 72, 100, 122, 135, 870
AclWI	GGATC	1 cut(s)	416
AcoI	YGGCCR	4 cut(s)	24, 79, 165, 401
AcsI	RAATTY	2 cut(s)	242, 983
AcuI	CTGAAG	3 cut(s)	396, 599, 867
AcyI	GRCGYC	1 cut(s)	69
AfaI	GTAC	1 cut(s)	595
AfiI	CCNNNNNNNGG	2 cut(s)	163, 259
AgsI	TTSAA	3 cut(s)	521, 539, 748
AjnI	CCWGG	3 cut(s)	505, 835, 877
AluBI	AGCT	6 cut(s)	250, 498, 544, 731, 845, 934
AluI	AGCT	6 cut(s)	250, 498, 544, 731, 845, 934
Alw21I	GWGCWC	2 cut(s)	118, 936
Alw26I	GTCTC	3 cut(s)	13, 133, 688
AlwI	GGATC	1 cut(s)	416
AlwNI	CAGNNNCTG	1 cut(s)	22
AoxI	GGCC	6 cut(s)	24, 30, 79, 165, 401, 530
ApaI	GGGCCC	1 cut(s)	534
ApeKI	GCWGC	4 cut(s)	5, 53, 74, 728
ApoI	RAATTY	2 cut(s)	242, 983
AspLEI	GCGC	1 cut(s)	71
AspS9I	GGNCC	5 cut(s)	206, 530, 531, 839, 872
AsuC2I	CCSGG	1 cut(s)	93
AsuHPI	GGTGA	1 cut(s)	678
AsuNHI	GCTAGC	1 cut(s)	731
AvaII	GGWCC	3 cut(s)	206, 839, 872
BaeGI	GKGCMC	1 cut(s)	534
BalI	TGGCCA	1 cut(s)	26
BanI	GGYRCC	1 cut(s)	68
BanII	GRGCYC	3 cut(s)	358, 534, 936
BarI	GAAGNNNNNNTAC	2 cut(s)	740, 772
Bbv12I	GWGCWC	2 cut(s)	118, 936
BbvI	GCAGC	3 cut(s)	40, 61, 740
BccI	CCATC	1 cut(s)	880
BceAI	ACGGC	3 cut(s)	276, 388, 416
BciT130I	CCWGG	3 cut(s)	507, 837, 879
BciVI	GTATCC	2 cut(s)	431, 474
BcnI	CCSGG	1 cut(s)	93
BcoDI	GTCTC	3 cut(s)	13, 133, 688
BfaI	CTAG	2 cut(s)	182, 732
BfoI	RGCGCY	1 cut(s)	72
BfuAI	ACCTGC	1 cut(s)	578
BfuI	GTATCC	2 cut(s)	431, 474
BisI	GCNGC	8 cut(s)	6, 33, 54, 63, 72, 75, 123, 729
BlsI	GCNGC	8 cut(s)	7, 34, 55, 64, 73, 76, 124, 730
Bme1390I	CCNGG	4 cut(s)	93, 507, 837, 879
Bme18I	GGWCC	3 cut(s)	206, 839, 872
BmgT120I	GGNCC	5 cut(s)	206, 530, 531, 839, 872
BmiI	GGNNCC	4 cut(s)	70, 357, 531, 532
BmrFI	CCNGG	4 cut(s)	93, 507, 837, 879
BmtI	GCTAGC	1 cut(s)	735
BpuMI	CCSGG	1 cut(s)	93
BsaHI	GRCGYC	1 cut(s)	69
BsaJI	CCNNGG	3 cut(s)	27, 432, 657
BsaXI	ACNNNNNCTCC	2 cut(s)	340, 370
Bsc4I	CCNNNNNNNGG	2 cut(s)	163, 259
Bse118I	RCCGGY	2 cut(s)	65, 77
Bse1I	ACTGG	3 cut(s)	27, 157, 601
BseBI	CCWGG	3 cut(s)	507, 837, 879
BseDI	CCNNGG	3 cut(s)	27, 432, 657
BseGI	GGATG	1 cut(s)	858
BseLI	CCNNNNNNNGG	2 cut(s)	163, 259
BseMII	CTCAG	2 cut(s)	536, 949
BseNI	ACTGG	3 cut(s)	27, 157, 601
BseSI	GKGCMC	1 cut(s)	534
BseX3I	CGGCCG	2 cut(s)	165, 401
BseXI	GCAGC	3 cut(s)	40, 61, 740
Bsh1236I	CGCG	1 cut(s)	122
Bsh1285I	CGRYCG	2 cut(s)	168, 404
BshFI	GGCC	6 cut(s)	26, 32, 81, 167, 403, 532
BshNI	GGYRCC	1 cut(s)	68
BsiEI	CGRYCG	2 cut(s)	168, 404
BsiHKAI	GWGCWC	2 cut(s)	118, 936
BsiSI	CCGG	4 cut(s)	66, 78, 93, 132
BslI	CCNNNNNNNGG	2 cut(s)	163, 259
BsmAI	GTCTC	3 cut(s)	13, 133, 688
BsnI	GGCC	6 cut(s)	26, 32, 81, 167, 403, 532
Bsp120I	GGGCCC	1 cut(s)	530
Bsp1286I	GDGCHC	4 cut(s)	118, 358, 534, 936
Bsp143I	GATC	1 cut(s)	421
BspACI	CCGC	8 cut(s)	33, 42, 63, 72, 100, 122, 135, 870
BspANI	GGCC	6 cut(s)	26, 32, 81, 167, 403, 532
BspCNI	CTCAG	2 cut(s)	537, 948
BspFNI	CGCG	1 cut(s)	122
BspLI	GGNNCC	4 cut(s)	70, 357, 531, 532
BspMI	ACCTGC	1 cut(s)	578
BspOI	GCTAGC	1 cut(s)	735
BspPI	GGATC	1 cut(s)	416
BspT107I	GGYRCC	1 cut(s)	68
BsrFI	RCCGGY	2 cut(s)	65, 77
BsrI	ACTGG	3 cut(s)	27, 157, 601
BssAI	RCCGGY	2 cut(s)	65, 77
BssECI	CCNNGG	3 cut(s)	27, 432, 657
BssMI	GATC	1 cut(s)	421
BssNAI	GTATAC	1 cut(s)	408
BssNI	GRCGYC	1 cut(s)	69
BssT1I	CCWWGG	3 cut(s)	27, 432, 657
Bst1107I	GTATAC	1 cut(s)	408
Bst2UI	CCWGG	3 cut(s)	507, 837, 879
Bst4CI	ACNGT	1 cut(s)	763
BstACI	GRCGYC	1 cut(s)	69
BstC8I	GCNNGC	5 cut(s)	67, 79, 485, 496, 733
BstDEI	CTNAG	3 cut(s)	87, 545, 935
BstF5I	GGATG	1 cut(s)	858
BstFNI	CGCG	1 cut(s)	122
BstH2I	RGCGCY	1 cut(s)	72
BstHHI	GCGC	1 cut(s)	71
BstKTI	GATC	1 cut(s)	424
BstMAI	GTCTC	3 cut(s)	13, 133, 688
BstMBI	GATC	1 cut(s)	421
BstMCI	CGRYCG	2 cut(s)	168, 404
BstMWI	GCNNNNNNNGC	8 cut(s)	32, 41, 50, 59, 62, 68, 71, 122
BstNI	CCWGG	3 cut(s)	507, 837, 879
BstSCI	CCNGG	4 cut(s)	91, 505, 835, 877
BstSLI	GKGCMC	1 cut(s)	534
BstUI	CGCG	1 cut(s)	122
BstV1I	GCAGC	3 cut(s)	40, 61, 740
BstX2I	RGATCY	1 cut(s)	421
BstXI	CCANNNNNNTGG	1 cut(s)	234
BstYI	RGATCY	1 cut(s)	421
BstZ17I	GTATAC	1 cut(s)	408
BstZI	CGGCCG	2 cut(s)	165, 401
BsuI	GTATCC	2 cut(s)	431, 474
BsuRI	GGCC	6 cut(s)	26, 32, 81, 167, 403, 532
BtgZI	GCGATG	1 cut(s)	525
BtsCI	GGATG	1 cut(s)	858
BtsI	GCAGTG	2 cut(s)	48, 665
BtsIMutI	CAGTG	2 cut(s)	48, 665
BveI	ACCTGC	1 cut(s)	578
Cac8I	GCNNGC	5 cut(s)	67, 79, 485, 496, 733
CaiI	CAGNNNCTG	1 cut(s)	22
CfoI	GCGC	1 cut(s)	71
Cfr10I	RCCGGY	2 cut(s)	65, 77
Cfr13I	GGNCC	5 cut(s)	206, 530, 531, 839, 872
CseI	GACGC	1 cut(s)	380
CsiI	ACCWGGT	1 cut(s)	835
Csp6I	GTAC	1 cut(s)	594
CviAII	CATG	4 cut(s)	339, 562, 580, 817
CviQI	GTAC	1 cut(s)	594
DdeI	CTNAG	3 cut(s)	87, 545, 935
DinI	GGCGCC	1 cut(s)	70
DpnI	GATC	1 cut(s)	423
DpnII	GATC	1 cut(s)	421
EaeI	YGGCCR	4 cut(s)	24, 79, 165, 401
EagI	CGGCCG	2 cut(s)	165, 401
EciI	GGCGGA	1 cut(s)	150
Ecl136II	GAGCTC	1 cut(s)	934
EclXI	CGGCCG	2 cut(s)	165, 401
Eco130I	CCWWGG	3 cut(s)	27, 432, 657
Eco24I	GRGCYC	3 cut(s)	358, 534, 936
Eco47I	GGWCC	3 cut(s)	206, 839, 872
Eco52I	CGGCCG	2 cut(s)	165, 401
Eco53kI	GAGCTC	1 cut(s)	934
Eco57I	CTGAAG	3 cut(s)	396, 599, 867
EcoICRI	GAGCTC	1 cut(s)	934
EcoO109I	RGGNCCY	1 cut(s)	530
EcoRII	CCWGG	3 cut(s)	505, 835, 877
EcoT14I	CCWWGG	3 cut(s)	27, 432, 657
EcoT38I	GRGCYC	3 cut(s)	358, 534, 936
EgeI	GGCGCC	1 cut(s)	70
EheI	GGCGCC	1 cut(s)	70
ErhI	CCWWGG	3 cut(s)	27, 432, 657
FaeI	CATG	4 cut(s)	342, 565, 583, 820
FalI	AAGNNNNNCTT	4 cut(s)	80, 112, 425, 457
FatI	CATG	4 cut(s)	338, 561, 579, 816
FblI	GTMKAC	1 cut(s)	407
Fnu4HI	GCNGC	8 cut(s)	6, 33, 54, 63, 72, 75, 123, 729
FokI	GGATG	1 cut(s)	845
FriOI	GRGCYC	3 cut(s)	358, 534, 936
Fsp4HI	GCNGC	8 cut(s)	6, 33, 54, 63, 72, 75, 123, 729
FspBI	CTAG	2 cut(s)	182, 732
GlaI	GCGC	1 cut(s)	70
GluI	GCNGC	8 cut(s)	6, 33, 54, 63, 72, 75, 123, 729
HaeII	RGCGCY	1 cut(s)	72
HaeIII	GGCC	6 cut(s)	26, 32, 81, 167, 403, 532
HapII	CCGG	4 cut(s)	66, 78, 93, 132
HgaI	GACGC	1 cut(s)	380
HhaI	GCGC	1 cut(s)	71
Hin1I	GRCGYC	1 cut(s)	69
Hin1II	CATG	4 cut(s)	342, 565, 583, 820
Hin6I	GCGC	1 cut(s)	69
HinP1I	GCGC	1 cut(s)	69
HindIII	AAGCTT	1 cut(s)	496
HinfI	GANTC	4 cut(s)	138, 170, 428, 650
HpaII	CCGG	4 cut(s)	66, 78, 93, 132
HphI	GGTGA	1 cut(s)	678
Hpy166II	GTNNAC	2 cut(s)	408, 979
Hpy188I	TCNGA	5 cut(s)	18, 113, 293, 421, 938
Hpy188III	TCNNGA	5 cut(s)	182, 227, 425, 467, 644
Hpy8I	GTNNAC	2 cut(s)	408, 979
Hpy99I	CGWCG	1 cut(s)	166
HpyAV	CCTTC	5 cut(s)	55, 371, 464, 520, 742
HpyCH4III	ACNGT	1 cut(s)	763
HpyCH4V	TGCA	4 cut(s)	218, 494, 587, 728
HpyF10VI	GCNNNNNNNGC	8 cut(s)	32, 41, 50, 59, 62, 68, 71, 122
HpyF3I	CTNAG	3 cut(s)	87, 545, 935
Hsp92I	GRCGYC	1 cut(s)	69
Hsp92II	CATG	4 cut(s)	342, 565, 583, 820
HspAI	GCGC	1 cut(s)	69
KasI	GGCGCC	1 cut(s)	68
KroI	GCCGGC	2 cut(s)	65, 77
KroNI	GCCGGC	2 cut(s)	67, 79
Kzo9I	GATC	1 cut(s)	421
LmnI	GCTCC	3 cut(s)	113, 255, 361
Lsp1109I	GCAGC	3 cut(s)	40, 61, 740
MabI	ACCWGGT	1 cut(s)	835
MaeI	CTAG	2 cut(s)	182, 732
MaeIII	GTNAC	2 cut(s)	663, 959
MalI	GATC	1 cut(s)	423
MboI	GATC	1 cut(s)	421
MboII	GAAGA	3 cut(s)	577, 860, 980
MfeI	CAATTG	2 cut(s)	284, 360
MflI	RGATCY	1 cut(s)	421
MhlI	GDGCHC	4 cut(s)	118, 358, 534, 936
MlsI	TGGCCA	1 cut(s)	26
MluCI	AATT	7 cut(s)	213, 242, 284, 360, 742, 822, 983
MluNI	TGGCCA	1 cut(s)	26
Mly113I	GGCGCC	1 cut(s)	69
MlyI	GAGTC	1 cut(s)	644
MmeI	TCCRAC	2 cut(s)	764, 797
MnlI	CCTC	5 cut(s)	353, 619, 785, 885, 932
Mox20I	TGGCCA	1 cut(s)	26
MreI	CGCCGGCG	1 cut(s)	65
MroNI	GCCGGC	2 cut(s)	65, 77
MscI	TGGCCA	1 cut(s)	26
MseI	TTAA	7 cut(s)	222, 246, 456, 500, 636, 705, 825
Msp20I	TGGCCA	1 cut(s)	26
MspA1I	CMGCKG	2 cut(s)	74, 845
MspI	CCGG	4 cut(s)	66, 78, 93, 132
MspR9I	CCNGG	4 cut(s)	93, 507, 837, 879
MunI	CAATTG	2 cut(s)	284, 360
MvaI	CCWGG	3 cut(s)	507, 837, 879
MvnI	CGCG	1 cut(s)	122
MwoI	GCNNNNNNNGC	8 cut(s)	32, 41, 50, 59, 62, 68, 71, 122
NaeI	GCCGGC	2 cut(s)	67, 79
NarI	GGCGCC	1 cut(s)	69
NciI	CCSGG	1 cut(s)	93
NdeII	GATC	1 cut(s)	421
NgoMIV	GCCGGC	2 cut(s)	65, 77
NheI	GCTAGC	1 cut(s)	731
NlaIII	CATG	4 cut(s)	342, 565, 583, 820
NlaIV	GGNNCC	4 cut(s)	70, 357, 531, 532
NmuCI	GTSAC	1 cut(s)	663
PaqCI	CACCTGC	1 cut(s)	578
PdiI	GCCGGC	2 cut(s)	67, 79
PfeI	GAWTC	3 cut(s)	138, 170, 428
PfoI	TCCNGGA	1 cut(s)	877
PkrI	GCNGC	8 cut(s)	7, 34, 55, 64, 73, 76, 124, 730
PleI	GAGTC	1 cut(s)	644
PluTI	GGCGCC	1 cut(s)	72
PpsI	GAGTC	1 cut(s)	644
Psp124BI	GAGCTC	1 cut(s)	936
Psp6I	CCWGG	3 cut(s)	505, 835, 877
PspGI	CCWGG	3 cut(s)	505, 835, 877
PspN4I	GGNNCC	4 cut(s)	70, 357, 531, 532
PspOMI	GGGCCC	1 cut(s)	530
PspPI	GGNCC	5 cut(s)	206, 530, 531, 839, 872
PstNI	CAGNNNCTG	1 cut(s)	22
PsuI	RGATCY	1 cut(s)	421
PvuII	CAGCTG	1 cut(s)	845
RsaI	GTAC	1 cut(s)	595
RsaNI	GTAC	1 cut(s)	594
SacI	GAGCTC	1 cut(s)	936
SaqAI	TTAA	7 cut(s)	222, 246, 456, 500, 636, 705, 825
SatI	GCNGC	8 cut(s)	6, 33, 54, 63, 72, 75, 123, 729
Sau3AI	GATC	1 cut(s)	421
Sau96I	GGNCC	5 cut(s)	206, 530, 531, 839, 872
SchI	GAGTC	1 cut(s)	644
ScrFI	CCNGG	4 cut(s)	93, 507, 837, 879
SduI	GDGCHC	4 cut(s)	118, 358, 534, 936
SexAI	ACCWGGT	1 cut(s)	835
SfoI	GGCGCC	1 cut(s)	70
SgrAI	CRCCGGYG	1 cut(s)	65
SinI	GGWCC	3 cut(s)	206, 839, 872
Sse9I	AATT	7 cut(s)	213, 242, 284, 360, 742, 822, 983
SsiI	CCGC	8 cut(s)	33, 42, 63, 72, 100, 122, 135, 870
SspDI	GGCGCC	1 cut(s)	68
SspMI	CTAG	2 cut(s)	182, 732
SstI	GAGCTC	1 cut(s)	936
StyD4I	CCNGG	4 cut(s)	91, 505, 835, 877
StyI	CCWWGG	3 cut(s)	27, 432, 657
TaaI	ACNGT	1 cut(s)	763
TasI	AATT	7 cut(s)	213, 242, 284, 360, 742, 822, 983
TauI	GCSGC	4 cut(s)	35, 65, 74, 125
TfiI	GAWTC	3 cut(s)	138, 170, 428
Tru1I	TTAA	7 cut(s)	222, 246, 456, 500, 636, 705, 825
Tru9I	TTAA	7 cut(s)	222, 246, 456, 500, 636, 705, 825
TscAI	CASTG	2 cut(s)	55, 672
TseFI	GTSAC	1 cut(s)	663
TseI	GCWGC	4 cut(s)	5, 53, 74, 728
Tsp45I	GTSAC	1 cut(s)	663
TspDTI	ATGAA	3 cut(s)	201, 327, 578
TspRI	CASTG	2 cut(s)	55, 672
VpaK11BI	GGWCC	3 cut(s)	206, 839, 872
XapI	RAATTY	2 cut(s)	242, 983
XbaI	TCTAGA	1 cut(s)	181
XcmI	CCANNNNNNNNNTGG	1 cut(s)	780
XmiI	GTMKAC	1 cut(s)	407
XspI	CTAG	2 cut(s)	182, 732

Using CommOnly database (standard laboratory enzymes). Scanned on CDS sequence.