Rorug02G0030300

Pectinesterase

Design Primers CRISPR Design

Basic Information

Type: gene
Biological Identity
rosa_rugosa
GWHBQTZ00000002
OG0004453
Physical Location & Seq
Reverse (-)
2438692 .. 2440638
1947 bp
Loading structure...
UTR
Exon/CDS
Intron
Rorug02G0030300.1

Sequence Viewer

Length: 1671 bp
ATGGCTGCCTCTGTATCTCCATTGTCCGAATCAATCATGAAGAAAGCAACACATGCCTCTGCCAAGTTCCGAAACTCTTTTACCCGAAAGGGTAAGAGAAGCAGCAGCAAAGTCATGTCTGTCGAAATTGCAGATGTGCATGATGTTGAGGAAGGTCAGGCTGTTGACAAGCTTCGTCAGGCGCTTATATTGGACGAGCTTCTTCCAAGCAAGCACGATGATTATCACATGATGCTTCGGTTCTTGAAGGCCAGGAAATTCGACATTGAGCAGACCAAGCAAATGTGGTCTGAAATGCTTCAATGGAGGAAGGAATTCGGAGCTGACACGATCAGAGATGATTTTGAATTCGAGGAACTCAATGAAGTCCTGGATTATTATCCCCAAGGACATCATGGGGTGGACAAAGAAGGAAGACCGGTCTATATTGAGAGACTGGGACTAGTTGATTCAACCAAGCTCATGCAAGTCACAACCATGGACCGGTATTTGAAATACCATGTTCAGGAATTCGAAAGAACATTTGATGTTAAATTCCCAGCTTGTTCGATTGCAGCGAAGAAGCACATTGATCAAAGTACAACTATTCTTGATGTTCAAGGAGTTGGACTAAAAAATTTCAACAAAGCTGCTAGGGAGCTTATTATGAGCCTTCAGAAGATTGATGGTGACAACTATCCTGAGACCTTGAGCCGGATGTTTATTATCAATGCTGGTTCTGGTTTTAGAATGTTGTGGAGCACTATCAAGTCATTCCTTGATCCGAGGACAACGGTGAAGATTCATGTCCTTGGGAATAAGTACCAGAGCAATTTGCTTGAGATTATTGATGCCAGTGAGTTGCCAGAGTTTTTAGGAGGAAAATGTACTTGTGCTGATGAAGGAGGATGTATGCGTTCCGATAAGGGACCATGGAAAAATCCAGAGATAATGAGTATGGTCCATAATGGTGATCATAAGTGTTCAAAGGAACCAAGGACTCAGGTGGCTGAAGACAAGAAGTCAGTGTATGTATCCGATGACGAATTGGTGTATTCCAAGGGACATTCTTCCCCTTCTTTACCGAAAAATTTCCACGAATCTTTCAAGCATGAAGATTATGCCCCAATGGTTGATAAAGCTATGGATGTAACAAGTCACATTGTTGTGGGAAGTGTCAAATTTGACACATTTGCCTCTTCTAAGTCGGCAGATTGTTTCGACATGCCTGAGGCTTGCAAACCTCCTGAGGGGTTTAGCGCTCGTTTTTTGGGTGCTATTATGGCATTTATTTTGAGCATTGTGGCTATTTTCCGGCTAACAAGTAACGTGCCAAGGAAGCTTACTAATTCCACCTTCTACTCTAGTGCTGTTTGCCACGACGACACAATGCTGAAGAGCAAGGCGCCTTCCAAGCCTGCCATTGCCATTTCTGATGCAGATTTCCAGTCCATCATCAAACGCATGGCGGAGTTGGAAGAGAGAGCCATTGTTTTGAGCATAAAACAAGCCACCATGCCACCCGAGAAGGAGGAGTTGCTTACTGCTGCTTTGAGCCGGGTTGATGCCTTGGAGCAAGAGCTTATGGCAACTACTAAGGCTCTCCAAGAGTCAATTTCCCGCCAAGATGAGCTCTTGGCATATGTTGAAAAGAAGAAGAGGAAAAGGAGGTTTTTCAAGTTCCACTGGTGA

Protein Analysis

556

Amino Acids

63.14

Weight (kDa)

7.0

Isoelectric Point (pI)

48.84

Instability Index
Protein Domains (Pfam)
Domain Name Pfam ID Position E-value Description
CRAL_TRIO PF00650 126 - 287 3.5e-30 CRAL/TRIO domain
Hydrophobicity Profile (Kyte-Doolittle)
AI Structure Prediction Report
Calculating structure properties...

Genomic Context

Gene Family Tree

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Restriction Enzyme Sites

1 / 10
Enzyme Recognition Site Cut Count Positions (bp)
AccB1I GGYRCC 1 cut(s) 1384
AciI CCGC 2 cut(s) 1448, 1600
AclWI GGATC 1 cut(s) 755
AcsI RAATTY 8 cut(s) 257, 314, 347, 509, 533, 616, 1069, 1160
AcuI CTGAAG 3 cut(s) 638, 1011, 1394
AcyI GRCGYC 1 cut(s) 1385
AfaI GTAC 3 cut(s) 580, 803, 868
AfeI AGCGCT 1 cut(s) 1240
AfiI CCNNNNNNNGG 4 cut(s) 483, 505, 693, 1229
AgeI ACCGGT 2 cut(s) 418, 483
AhdI GACNNNNNGTC 1 cut(s) 1000
AhlI ACTAGT 1 cut(s) 442
AjnI CCWGG 2 cut(s) 251, 369
Alw21I GWGCWC 2 cut(s) 743, 1614
Alw26I GTCTC 2 cut(s) 427, 677
AlwI GGATC 1 cut(s) 755
Ama87I CYCGRG 1 cut(s) 1502
Aor51HI AGCGCT 1 cut(s) 1240
AoxI GGCC 1 cut(s) 249
ApeKI GCWGC 6 cut(s) 5, 102, 105, 554, 629, 1526
ApoI RAATTY 8 cut(s) 257, 314, 347, 509, 533, 616, 1069, 1160
AsiGI ACCGGT 2 cut(s) 418, 483
Asp700I GAANNNNTTC 3 cut(s) 297, 314, 1070
AspLEI GCGC 3 cut(s) 184, 1241, 1387
AspS9I GGNCC 3 cut(s) 481, 908, 940
AsuC2I CCSGG 1 cut(s) 1538
AsuHPI GGTGA 3 cut(s) 680, 787, 962
AsuII TTCGAA 1 cut(s) 513
AvaI CYCGRG 1 cut(s) 1502
AvaII GGWCC 3 cut(s) 481, 908, 940
AxyI CCTNAGG 2 cut(s) 1209, 1227
BanI GGYRCC 1 cut(s) 1384
BanII GRGCYC 1 cut(s) 1614
BbsI GAAGAC 2 cut(s) 421, 999
Bbv12I GWGCWC 2 cut(s) 743, 1614
BbvI GCAGC 5 cut(s) 114, 117, 566, 616, 1513
BccI CCATC 2 cut(s) 659, 1439
BciT130I CCWGG 2 cut(s) 253, 371
BciVI GTATCC 1 cut(s) 1024
BclI TGATCA 2 cut(s) 571, 952
BcnI CCSGG 1 cut(s) 1538
BcoDI GTCTC 2 cut(s) 427, 677
BcuI ACTAGT 1 cut(s) 442
BfaI CTAG 3 cut(s) 443, 633, 1344
BfoI RGCGCY 3 cut(s) 185, 1242, 1388
BfuI GTATCC 1 cut(s) 1024
BisI GCNGC 6 cut(s) 6, 103, 106, 555, 630, 1527
BlsI GCNGC 6 cut(s) 7, 104, 107, 556, 631, 1528
Bme1390I CCNGG 3 cut(s) 253, 371, 1538
Bme18I GGWCC 3 cut(s) 481, 908, 940
BmeRI GACNNNNNGTC 1 cut(s) 1000
BmeT110I CYCGRG 1 cut(s) 1502
BmgT120I GGNCC 3 cut(s) 481, 908, 940
BmiI GGNNCC 3 cut(s) 909, 972, 1386
BmrFI CCNGG 3 cut(s) 253, 371, 1538
BmrI ACTGGG 1 cut(s) 446
BmsI GCATC 4 cut(s) 222, 820, 1405, 1534
BmuI ACTGGG 1 cut(s) 446
BpiI GAAGAC 2 cut(s) 421, 999
Bpu14I TTCGAA 1 cut(s) 513
BpuEI CTTGAG 2 cut(s) 709, 839
BpuMI CCSGG 1 cut(s) 1538
BsaBI GATNNNNATC 2 cut(s) 222, 378
BsaHI GRCGYC 1 cut(s) 1385
BsaI GGTCTC 1 cut(s) 677
BsaJI CCNNGG 9 cut(s) 385, 477, 764, 790, 911, 974, 1038, 1313, 1548
BsaWI WCCGGW 2 cut(s) 418, 483
Bsc4I CCNNNNNNNGG 4 cut(s) 483, 505, 693, 1229
Bse118I RCCGGY 2 cut(s) 418, 483
Bse1I ACTGG 4 cut(s) 441, 834, 1426, 1670
Bse21I CCTNAGG 2 cut(s) 1209, 1227
Bse3DI GCAATG 1 cut(s) 1401
Bse8I GATNNNNATC 2 cut(s) 222, 378
BseBI CCWGG 2 cut(s) 253, 371
BseDI CCNNGG 9 cut(s) 385, 477, 764, 790, 911, 974, 1038, 1313, 1548
BseGI GGATG 3 cut(s) 702, 893, 1132
BseJI GATNNNNATC 2 cut(s) 222, 378
BseLI CCNNNNNNNGG 4 cut(s) 483, 505, 693, 1229
BseMI GCAATG 1 cut(s) 1401
BseMII CTCAG 4 cut(s) 672, 995, 1200, 1218
BseNI ACTGG 4 cut(s) 441, 834, 1426, 1670
BseRI GAGGAG 1 cut(s) 1526
BseXI GCAGC 5 cut(s) 114, 117, 566, 616, 1513
BseYI CCCAGC 1 cut(s) 538
BshFI GGCC 1 cut(s) 251
BshNI GGYRCC 1 cut(s) 1384
BshTI ACCGGT 2 cut(s) 418, 483
BsiHKAI GWGCWC 2 cut(s) 743, 1614
BsiHKCI CYCGRG 1 cut(s) 1502
BsiSI CCGG 5 cut(s) 419, 484, 694, 1294, 1537
BslFI GGGAC 3 cut(s) 453, 921, 1056
BslI CCNNNNNNNGG 4 cut(s) 483, 505, 693, 1229
BsmAI GTCTC 2 cut(s) 427, 677
BsmFI GGGAC 3 cut(s) 453, 921, 1056
BsnI GGCC 1 cut(s) 251
Bso31I GGTCTC 1 cut(s) 677
BsoBI CYCGRG 1 cut(s) 1502
Bsp119I TTCGAA 1 cut(s) 513
Bsp1286I GDGCHC 2 cut(s) 743, 1614
Bsp143I GATC 4 cut(s) 330, 571, 760, 952
Bsp19I CCATGG 2 cut(s) 477, 911
BspACI CCGC 2 cut(s) 1448, 1600
BspANI GGCC 1 cut(s) 251
BspCNI CTCAG 4 cut(s) 673, 994, 1201, 1219
BspHI TCATGA 1 cut(s) 36
BspLI GGNNCC 3 cut(s) 909, 972, 1386
BspPI GGATC 1 cut(s) 755
BspQI GCTCTTC 1 cut(s) 1370
BspT104I TTCGAA 1 cut(s) 513
BspT107I GGYRCC 1 cut(s) 1384
BspTNI GGTCTC 1 cut(s) 677
BsrDI GCAATG 1 cut(s) 1401
BsrFI RCCGGY 2 cut(s) 418, 483
BsrI ACTGG 4 cut(s) 441, 834, 1426, 1670
BssAI RCCGGY 2 cut(s) 418, 483
BssECI CCNNGG 9 cut(s) 385, 477, 764, 790, 911, 974, 1038, 1313, 1548
BssMI GATC 4 cut(s) 330, 571, 760, 952
BssNI GRCGYC 1 cut(s) 1385
BssT1I CCWWGG 8 cut(s) 385, 477, 790, 911, 974, 1038, 1313, 1548
Bst2UI CCWGG 2 cut(s) 253, 371
Bst4CI ACNGT 1 cut(s) 775
Bst6I CTCTTC 4 cut(s) 1183, 1370, 1452, 1631
BstACI GRCGYC 1 cut(s) 1385
BstAPI GCANNNNNTGC 1 cut(s) 53
BstBI TTCGAA 1 cut(s) 513
BstC8I GCNNGC 3 cut(s) 212, 1216, 1398
BstDEI CTNAG 6 cut(s) 681, 981, 1182, 1209, 1227, 1575
BstDSI CCRYGG 2 cut(s) 477, 911
BstF5I GGATG 3 cut(s) 702, 893, 1132
BstH2I RGCGCY 3 cut(s) 185, 1242, 1388
BstHHI GCGC 3 cut(s) 184, 1241, 1387
BstKTI GATC 4 cut(s) 333, 574, 763, 955
BstMAI GTCTC 2 cut(s) 427, 677
BstMBI GATC 4 cut(s) 330, 571, 760, 952
BstMWI GCNNNNNNNGC 5 cut(s) 53, 277, 1262, 1318, 1393
BstNI CCWGG 2 cut(s) 253, 371
BstNSI RCATGY 2 cut(s) 56, 1207
BstSCI CCNGG 3 cut(s) 251, 369, 1536
BstV1I GCAGC 5 cut(s) 114, 117, 566, 616, 1513
BstV2I GAAGAC 2 cut(s) 421, 999
Bsu36I CCTNAGG 2 cut(s) 1209, 1227
BsuI GTATCC 1 cut(s) 1024
BsuRI GGCC 1 cut(s) 251
BtgI CCRYGG 2 cut(s) 477, 911
BtsCI GGATG 3 cut(s) 702, 893, 1132
BtsIMutI CAGTG 3 cut(s) 841, 1011, 1663
Cac8I GCNNGC 3 cut(s) 212, 1216, 1398
CciI TCATGA 1 cut(s) 36
CfoI GCGC 3 cut(s) 184, 1241, 1387
Cfr10I RCCGGY 2 cut(s) 418, 483
Cfr13I GGNCC 3 cut(s) 481, 908, 940
Csp6I GTAC 3 cut(s) 579, 802, 867
CspAI ACCGGT 2 cut(s) 418, 483
CviQI GTAC 3 cut(s) 579, 802, 867
DdeI CTNAG 6 cut(s) 681, 981, 1182, 1209, 1227, 1575
DinI GGCGCC 1 cut(s) 1386
DpnI GATC 4 cut(s) 332, 573, 762, 954
DpnII GATC 4 cut(s) 330, 571, 760, 952
DriI GACNNNNNGTC 1 cut(s) 1000
Eam1104I CTCTTC 4 cut(s) 1183, 1370, 1452, 1631
Eam1105I GACNNNNNGTC 1 cut(s) 1000
EarI CTCTTC 4 cut(s) 1183, 1370, 1452, 1631
EciI GGCGGA 1 cut(s) 1463
Ecl136II GAGCTC 1 cut(s) 1612
Eco130I CCWWGG 8 cut(s) 385, 477, 790, 911, 974, 1038, 1313, 1548
Eco24I GRGCYC 1 cut(s) 1614
Eco31I GGTCTC 1 cut(s) 677
Eco47I GGWCC 3 cut(s) 481, 908, 940
Eco47III AGCGCT 1 cut(s) 1240
Eco53kI GAGCTC 1 cut(s) 1612
Eco57I CTGAAG 3 cut(s) 638, 1011, 1394
Eco81I CCTNAGG 2 cut(s) 1209, 1227
Eco88I CYCGRG 1 cut(s) 1502
EcoICRI GAGCTC 1 cut(s) 1612
EcoRI GAATTC 3 cut(s) 314, 347, 509
EcoRII CCWGG 2 cut(s) 251, 369
EcoT14I CCWWGG 8 cut(s) 385, 477, 790, 911, 974, 1038, 1313, 1548
EcoT38I GRGCYC 1 cut(s) 1614
EgeI GGCGCC 1 cut(s) 1386
EheI GGCGCC 1 cut(s) 1386
ErhI CCWWGG 8 cut(s) 385, 477, 790, 911, 974, 1038, 1313, 1548
FaqI GGGAC 3 cut(s) 453, 921, 1056
FauI CCCGC 1 cut(s) 1607
FauNDI CATATG 1 cut(s) 1621
FbaI TGATCA 2 cut(s) 571, 952
Fnu4HI GCNGC 6 cut(s) 6, 103, 106, 555, 630, 1527
FokI GGATG 3 cut(s) 709, 900, 1139
FriOI GRGCYC 1 cut(s) 1614
Fsp4HI GCNGC 6 cut(s) 6, 103, 106, 555, 630, 1527
FspBI CTAG 3 cut(s) 443, 633, 1344
GlaI GCGC 3 cut(s) 183, 1240, 1386
GluI GCNGC 6 cut(s) 6, 103, 106, 555, 630, 1527
GsaI CCCAGC 1 cut(s) 542
HaeII RGCGCY 3 cut(s) 185, 1242, 1388
HaeIII GGCC 1 cut(s) 251
HapII CCGG 5 cut(s) 419, 484, 694, 1294, 1537
HhaI GCGC 3 cut(s) 184, 1241, 1387
Hin1I GRCGYC 1 cut(s) 1385
Hin6I GCGC 3 cut(s) 182, 1239, 1385
HinP1I GCGC 3 cut(s) 182, 1239, 1385
HincII GTYRAC 1 cut(s) 166
HindII GTYRAC 1 cut(s) 166
HindIII AAGCTT 2 cut(s) 170, 1319
HinfI GANTC 6 cut(s) 29, 449, 781, 979, 1079, 1589
HpaII CCGG 5 cut(s) 419, 484, 694, 1294, 1537
HphI GGTGA 3 cut(s) 680, 787, 962
Hpy166II GTNNAC 2 cut(s) 166, 403
Hpy188III TCNNGA 7 cut(s) 37, 244, 506, 590, 680, 923, 1226
Hpy8I GTNNAC 2 cut(s) 166, 403
Hpy99I CGWCG 1 cut(s) 1364
HpyCH4III ACNGT 1 cut(s) 775
HpyCH4IV ACGT 1 cut(s) 1308
HpyCH4V TGCA 6 cut(s) 131, 139, 466, 554, 1218, 1418
HpyF10VI GCNNNNNNNGC 5 cut(s) 53, 277, 1262, 1318, 1393
HpyF3I CTNAG 6 cut(s) 681, 981, 1182, 1209, 1227, 1575
HpySE526I ACGT 1 cut(s) 1308
Hsp92I GRCGYC 1 cut(s) 1385
HspAI GCGC 3 cut(s) 182, 1239, 1385
KasI GGCGCC 1 cut(s) 1384
Ksp22I TGATCA 2 cut(s) 571, 952
Kzo9I GATC 4 cut(s) 330, 571, 760, 952
LguI GCTCTTC 1 cut(s) 1370
LmnI GCTCC 4 cut(s) 320, 637, 738, 1552
Lsp1109I GCAGC 5 cut(s) 114, 117, 566, 616, 1513
LweI GCATC 4 cut(s) 222, 820, 1405, 1534
MaeI CTAG 3 cut(s) 443, 633, 1344
MaeII ACGT 1 cut(s) 1308
MaeIII GTNAC 5 cut(s) 469, 668, 1129, 1136, 1304
MalI GATC 4 cut(s) 332, 573, 762, 954
MboI GATC 4 cut(s) 330, 571, 760, 952
MhlI GDGCHC 2 cut(s) 743, 1614
Mly113I GGCGCC 1 cut(s) 1385
MlyI GAGTC 2 cut(s) 973, 1598
MmeI TCCRAC 2 cut(s) 586, 1434
MroXI GAANNNNTTC 3 cut(s) 297, 314, 1070
MseI TTAA 1 cut(s) 531
MslI CAYNNNNRTG 3 cut(s) 476, 948, 1145
MspI CCGG 5 cut(s) 419, 484, 694, 1294, 1537
MspR9I CCNGG 3 cut(s) 253, 371, 1538
MvaI CCWGG 2 cut(s) 253, 371
MwoI GCNNNNNNNGC 5 cut(s) 53, 277, 1262, 1318, 1393
NarI GGCGCC 1 cut(s) 1385
NciI CCSGG 1 cut(s) 1538
NcoI CCATGG 2 cut(s) 477, 911
NdeI CATATG 1 cut(s) 1621
NdeII GATC 4 cut(s) 330, 571, 760, 952
NlaIV GGNNCC 3 cut(s) 909, 972, 1386
NmuCI GTSAC 3 cut(s) 469, 668, 1136
NspI RCATGY 2 cut(s) 56, 1207
NspV TTCGAA 1 cut(s) 513
PagI TCATGA 1 cut(s) 36
PciSI GCTCTTC 1 cut(s) 1370
PcsI WCGNNNNNNNCGW 1 cut(s) 554
PdmI GAANNNNTTC 3 cut(s) 297, 314, 1070
PfeI GAWTC 4 cut(s) 29, 449, 781, 1079
PfoI TCCNGGA 1 cut(s) 369
PinAI ACCGGT 2 cut(s) 418, 483
PkrI GCNGC 6 cut(s) 7, 104, 107, 556, 631, 1528
PleI GAGTC 2 cut(s) 973, 1597
PluTI GGCGCC 1 cut(s) 1388
PpsI GAGTC 2 cut(s) 973, 1597
Psp124BI GAGCTC 1 cut(s) 1614
Psp6I CCWGG 2 cut(s) 251, 369
PspFI CCCAGC 1 cut(s) 538
PspGI CCWGG 2 cut(s) 251, 369
PspN4I GGNNCC 3 cut(s) 909, 972, 1386
PspPI GGNCC 3 cut(s) 481, 908, 940
RsaI GTAC 3 cut(s) 580, 803, 868
RsaNI GTAC 3 cut(s) 579, 802, 867
RseI CAYNNNNRTG 3 cut(s) 476, 948, 1145
SacI GAGCTC 1 cut(s) 1614
SapI GCTCTTC 1 cut(s) 1370
SaqAI TTAA 1 cut(s) 531
SatI GCNGC 6 cut(s) 6, 103, 106, 555, 630, 1527
Sau3AI GATC 4 cut(s) 330, 571, 760, 952
Sau96I GGNCC 3 cut(s) 481, 908, 940
SchI GAGTC 2 cut(s) 973, 1598
ScrFI CCNGG 3 cut(s) 253, 371, 1538
SduI GDGCHC 2 cut(s) 743, 1614
SfaNI GCATC 4 cut(s) 222, 820, 1405, 1534
SfoI GGCGCC 1 cut(s) 1386
SfuI TTCGAA 1 cut(s) 513
SinI GGWCC 3 cut(s) 481, 908, 940
SmiMI CAYNNNNRTG 3 cut(s) 476, 948, 1145
SmlI CTYRAG 2 cut(s) 688, 818
SmoI CTYRAG 2 cut(s) 688, 818
SpeI ACTAGT 1 cut(s) 442
SsiI CCGC 2 cut(s) 1448, 1600
SspDI GGCGCC 1 cut(s) 1384
SspMI CTAG 3 cut(s) 443, 633, 1344
SstI GAGCTC 1 cut(s) 1614
StyD4I CCNGG 3 cut(s) 251, 369, 1536
StyI CCWWGG 8 cut(s) 385, 477, 790, 911, 974, 1038, 1313, 1548
TaaI ACNGT 1 cut(s) 775
TaiI ACGT 1 cut(s) 1311
TaqI TCGA 6 cut(s) 123, 261, 351, 513, 548, 1200
TatI WGTACW 2 cut(s) 578, 866
TfiI GAWTC 4 cut(s) 29, 449, 781, 1079
Tru1I TTAA 1 cut(s) 531
Tru9I TTAA 1 cut(s) 531
TscAI CASTG 3 cut(s) 841, 1011, 1670
TseFI GTSAC 3 cut(s) 469, 668, 1136
TseI GCWGC 6 cut(s) 5, 102, 105, 554, 629, 1526
Tsp45I GTSAC 3 cut(s) 469, 668, 1136
TspDTI ATGAA 5 cut(s) 53, 378, 773, 894, 1107
TspRI CASTG 3 cut(s) 841, 1011, 1670
VpaK11BI GGWCC 3 cut(s) 481, 908, 940
XapI RAATTY 8 cut(s) 257, 314, 347, 509, 533, 616, 1069, 1160
XceI RCATGY 2 cut(s) 56, 1207
XcmI CCANNNNNNNNNTGG 1 cut(s) 392
XmnI GAANNNNTTC 3 cut(s) 297, 314, 1070
XspI CTAG 3 cut(s) 443, 633, 1344
Using CommOnly database (standard laboratory enzymes). Scanned on CDS sequence.