Basic Information

Type: gene

Biological Identity

Species

rosa_laevigata

Chromosome

Chr4

Orthogroup ID

OG0005013

Physical Location & Seq

Genomic Coordinates Reverse (-)

1995341 .. 1997894

Gene Structure 2554 bp

Loading structure...

UTR

Exon/CDS

Intron

Transcript / Protein ID

RLM00000015869

Sequence Viewer

CDS Protein Promoter (2k)

Length: 1524 bp

ATGGAAATTGTGCATATTATTCCGTTAAGAATGGCAATAACTATGTTAAGTATGTTGATGTTGGTACAATTGCCAGCAATCCCTGCCTCTGCGTTTGTCATACAGTCAAAGCTTTGTAAGGACCAGTTTGCCTATGCGACTTCTAATGGTAAGGACTTGTTTTACATAAATGGGAATTCAGTGGAGAAAGCTTTATTCTGTAATGCTCTTCAAACATACTATGCAAATGGTTGCATTTTGGAGGGGTGCCTCGGAAGCAGCCATTGTGGATCAGTCATTTCACTAGGAAGAAAACTTATGCAAAAGGAGTTGAAGGATGAATCCACTTCCCATCATGGAGAAAACAAGAACGCTTCTGAATCTATGTCCATTACCAAAACAGTTGGAATAGCAGCAAGTGGAGCGTTATTGGTGTGTTGTGTTTTCCTTTGTCCCTGCTTTTTTAAGAAAAGGACAGAAGCTGCTCACAATGTTCTTGCTAAGGACGCAAATTCAATGGATAATTCAGCTTCTTCTTTTGAAGTGAATCATGCCGCTGAAAAGATACCGGCCAGTCCACATCGTGTTCCACCTAGTCCTAGATATTCAAACTCTCCAAAACTCAGTAGACTTGGGTCTGTACATCTCAATTTGAATCAGATTGCCAGGGCAACCCGCAACTTCTCAAAATCACAACTCATAGGTGAAGGTGGTTTTGGAACTGTCTACAAAGCTCGGCTAGCTGATGGCCAAGTGGTTTCTATAAAACGTGCAAAGAAGGAACTCTTTGAGAATTTGCAAACCGAATTCAGTACAGAAGTTGAACTTCTGGCCAAAATTGATCATCGGAATCTTGTAAAGCTACTAGGTTATGTGGATGAAGGAAATGAACGACTTATCATTACAGAGTATGTGCCGAATGGTAATCTCAGAGAACATCTGGATGGTCAACATGGGAAAACCCTGGAGTTTAATCAGCGACTTGAAATAGCTATTGATGTTGCTCATGCCCTAACTTATCTTCATCTGTACACAGAAAAGCAAATTATCCATCGAGACGTGAAGTCATCCAACATTCTTCTGACAGAAAGCATGCGAGCCAAAGTTGCTGATTTTGGATTTGCCAGGCTTGGTCCGATGGATGCAGATCGAACACACATTTCTACTAAAGTGAAAGGAACTGTTGGCTACCTAGACCCGGAGTACATGAAAACATATCAACTCACTGCCAAAAGTGATGTCTACTCATTTGGGATATTACTTATAGAAATTGTGACGGGTCGTCGTCCAGTGGAGATGAAGAGACCAGTTGAAGAGCGTGTCACTCTCAGATGGGCATTCAAAAAGTTGAACGAAGGAAGAACGTGGGAATTGATTGATCCTGCAATAGAGGAAGCTGTAAATGCAGACGTACTACAGAAAATCTTTGATTTGGCTATCCAGTGTGCAGCTCCAATTCGAGCTGATCGGCCAGACATGAAAACAGTGGGGGAGCAGCTGTGGACAATAAGGGCAGACTACCTTAAAAGTCTCAAAAGAGGGTAG

Functional Annotation

Gene Ontology

Molecular Function
Biological Process
Cellular Component

No ontology terms assigned.

KEGG Pathways

Metabolic & Signaling

ko04010 ko04064 ko04620 ko04621 ko04624 ko04722 ko05133 ko05140 ko05142 ko05145 ko05152 ko05162 ko05164 map04010 map04064 map04620 map04621 map04624 map04722 map05133 map05140 map05142 map05145 map05152 map05162 map05164

Pfam Domains

Protein Families

Pkinase Pkinase_Tyr

Protein Analysis

508

Amino Acids

56.53

Weight (kDa)

8.81

Isoelectric Point (pI)

34.19

Instability Index

Protein Domains (Pfam)

Domain Name	Pfam ID	Position	E-value	Description
Pkinase	PF00069	222 - 491	2.2e-46	Protein kinase domain
PK_Tyr_Ser-Thr	PF07714	224 - 493	1.1e-50	Protein tyrosine and serine/threonine kinase

Hydrophobicity Profile (Kyte-Doolittle)

AI Structure Prediction Report

Calculating structure properties...

Genomic Context

Gene Family Tree

Species Filter

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Align Labels (Cladogram)

Image

Tree File

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Publication-ready

iTOL Interactive Tree Of Life · Most popular EvolView Flexible annotations MEGA Desktop phylogenetics suite FigTree Desktop tree editor

Orthologous Genes (Group: OG0005013)

Species	Orthologous Gene IDs
arabidopsis_thaliana	AT2G11520
fragaria_vesca	FvH4_1g02470 FvH4_1g02470 FvH4_1g02470 FvH4_1g02470 FvH4_1g02470 FvH4_1g02470 FvH4_1g02470
malus_domestica	MD02G1022200.v1.1 MD15G1164700.v1.1
prunus_persica	Prupe.7G250000_v2.0.a1 Prupe.7G250000_v2.0.a1 Prupe.7G250000_v2.0.a1 Prupe.7G250000_v2.0.a1 Prupe.7G250000_v2.0.a1 Prupe.7G250000_v2.0.a1
pyrus_communis	pycom02g01910 pycom15g14770
rosa_chinensis	RchiOBHm_Chr2g0087441
rosa_laevigata	RLG00000015869
rosa_multiflora	Rmu_sc0006064.1_g000012
rosa_roxburghii	Rroxscaffold_2G00153530
rosa_rugosa	Rorug01G0475300 Rorug01G0475400 Rorug01G0475500
rosa_samantha	Rh2AG028800 Rh2BG029100 Rh2CG029100 Rh2DG028900
rosa_wichuraiana	Rw2G002300

Restriction Enzyme Sites

1 / 10

Enzyme	Recognition Site	Cut Count	Positions (bp)
AccB1I	GGYRCC	1 cut(s)	246
AccI	GTMKAC	3 cut(s)	607, 705, 1221
AciI	CCGC	2 cut(s)	534, 655
AclWI	GGATC	2 cut(s)	277, 1352
AcoI	YGGCCR	4 cut(s)	549, 727, 810, 1448
AcsI	RAATTY	4 cut(s)	175, 490, 772, 785
AdeI	CACNNNGTG	1 cut(s)	563
AfaI	GTAC	6 cut(s)	66, 621, 793, 1010, 1184, 1392
AfiI	CCNNNNNNNGG	1 cut(s)	1177
AhdI	GACNNNNNGTC	2 cut(s)	1042, 1260
AjiI	CACGTC	1 cut(s)	1039
AjnI	CCWGG	3 cut(s)	644, 942, 1103
AjuI	GAANNNNNNNTTGG	2 cut(s)	678, 710
Alw26I	GTCTC	3 cut(s)	1029, 1276, 1514
AlwI	GGATC	2 cut(s)	277, 1352
AlwNI	CAGNNNCTG	1 cut(s)	461
AoxI	GGCC	4 cut(s)	549, 727, 810, 1448
ApeKI	GCWGC	5 cut(s)	258, 392, 461, 1427, 1474
ApoI	RAATTY	4 cut(s)	175, 490, 772, 785
ArsI	GACNNNNNNTTYG	2 cut(s)	1203, 1235
AspS9I	GGNCC	2 cut(s)	121, 1112
AsuC2I	CCSGG	1 cut(s)	1178
AsuHPI	GGTGA	1 cut(s)	695
AsuNHI	GCTAGC	1 cut(s)	718
AvaII	GGWCC	2 cut(s)	121, 1112
BalI	TGGCCA	2 cut(s)	729, 812
BanI	GGYRCC	1 cut(s)	246
BbvI	GCAGC	5 cut(s)	270, 404, 448, 1439, 1486
BccI	CCATC	6 cut(s)	339, 719, 917, 1038, 1111, 1305
BciT130I	CCWGG	3 cut(s)	646, 944, 1105
BclI	TGATCA	1 cut(s)	820
BcnI	CCSGG	1 cut(s)	1178
BcoDI	GTCTC	3 cut(s)	1029, 1276, 1514
BfaI	CTAG	6 cut(s)	284, 573, 579, 719, 845, 1172
BfmI	CTRYAG	1 cut(s)	1394
BisI	GCNGC	6 cut(s)	259, 393, 462, 534, 1428, 1475
BlsI	GCNGC	6 cut(s)	260, 394, 463, 535, 1429, 1476
Bme1390I	CCNGG	4 cut(s)	646, 944, 1105, 1178
Bme18I	GGWCC	2 cut(s)	121, 1112
BmeRI	GACNNNNNGTC	2 cut(s)	1042, 1260
BmgBI	CACGTC	1 cut(s)	1039
BmgT120I	GGNCC	2 cut(s)	121, 1112
BmiI	GGNNCC	1 cut(s)	248
BmrFI	CCNGG	4 cut(s)	646, 944, 1105, 1178
BmsI	GCATC	1 cut(s)	1111
BmtI	GCTAGC	1 cut(s)	722
BoxI	GACNNNNGTC	1 cut(s)	613
BpmI	CTGGAG	1 cut(s)	965
Bpu10I	CCTNAGC	1 cut(s)	480
BpuMI	CCSGG	1 cut(s)	1178
BsaBI	GATNNNNATC	1 cut(s)	1125
BsaI	GGTCTC	1 cut(s)	1276
BsaJI	CCNNGG	3 cut(s)	250, 645, 942
BsaXI	ACNNNNNCTCC	2 cut(s)	1463, 1493
Bsc4I	CCNNNNNNNGG	1 cut(s)	1177
Bse118I	RCCGGY	1 cut(s)	547
Bse1I	ACTGG	5 cut(s)	124, 552, 1268, 1286, 1420
Bse8I	GATNNNNATC	1 cut(s)	1125
BseBI	CCWGG	3 cut(s)	646, 944, 1105
BseDI	CCNNGG	3 cut(s)	250, 645, 942
BseGI	GGATG	5 cut(s)	322, 862, 928, 1046, 1126
BseJI	GATNNNNATC	1 cut(s)	1125
BseLI	CCNNNNNNNGG	1 cut(s)	1177
BseMII	CTCAG	3 cut(s)	616, 922, 1321
BseNI	ACTGG	5 cut(s)	124, 552, 1268, 1286, 1420
BseXI	GCAGC	5 cut(s)	270, 404, 448, 1439, 1486
BsgI	GTGCAG	1 cut(s)	1446
BshFI	GGCC	4 cut(s)	551, 729, 812, 1450
BshNI	GGYRCC	1 cut(s)	246
BsiSI	CCGG	2 cut(s)	548, 1178
BslFI	GGGAC	1 cut(s)	417
BslI	CCNNNNNNNGG	1 cut(s)	1177
BsmAI	GTCTC	3 cut(s)	1029, 1276, 1514
BsmBI	CGTCTC	1 cut(s)	1029
BsmFI	GGGAC	1 cut(s)	417
BsmI	GAATGC	1 cut(s)	1316
BsnI	GGCC	4 cut(s)	551, 729, 812, 1450
Bso31I	GGTCTC	1 cut(s)	1276
Bsp1407I	TGTACA	2 cut(s)	619, 1008
Bsp143I	GATC	5 cut(s)	269, 820, 1126, 1357, 1444
BspACI	CCGC	2 cut(s)	534, 655
BspANI	GGCC	4 cut(s)	551, 729, 812, 1450
BspCNI	CTCAG	3 cut(s)	615, 921, 1320
BspLI	GGNNCC	1 cut(s)	248
BspOI	GCTAGC	1 cut(s)	722
BspPI	GGATC	2 cut(s)	277, 1352
BspQI	GCTCTTC	2 cut(s)	213, 1287
BspT107I	GGYRCC	1 cut(s)	246
BspTNI	GGTCTC	1 cut(s)	1276
BsrFI	RCCGGY	1 cut(s)	547
BsrGI	TGTACA	2 cut(s)	619, 1008
BsrI	ACTGG	5 cut(s)	124, 552, 1268, 1286, 1420
BssAI	RCCGGY	1 cut(s)	547
BssECI	CCNNGG	3 cut(s)	250, 645, 942
BssMI	GATC	5 cut(s)	269, 820, 1126, 1357, 1444
Bst2UI	CCWGG	3 cut(s)	646, 944, 1105
Bst4CI	ACNGT	5 cut(s)	105, 382, 703, 1162, 1465
Bst6I	CTCTTC	3 cut(s)	213, 1274, 1287
BstAPI	GCANNNNNTGC	1 cut(s)	83
BstAUI	TGTACA	2 cut(s)	619, 1008
BstC8I	GCNNGC	4 cut(s)	75, 720, 1073, 1077
BstDEI	CTNAG	4 cut(s)	480, 602, 908, 1307
BstF5I	GGATG	5 cut(s)	322, 862, 928, 1046, 1126
BstKTI	GATC	5 cut(s)	272, 823, 1129, 1360, 1447
BstMAI	GTCTC	3 cut(s)	1029, 1276, 1514
BstMBI	GATC	5 cut(s)	269, 820, 1126, 1357, 1444
BstMWI	GCNNNNNNNGC	7 cut(s)	83, 255, 401, 485, 719, 1085, 1382
BstNI	CCWGG	3 cut(s)	646, 944, 1105
BstNSI	RCATGY	1 cut(s)	1075
BstPAI	GACNNNNGTC	1 cut(s)	613
BstSCI	CCNGG	4 cut(s)	644, 942, 1103, 1176
BstSFI	CTRYAG	1 cut(s)	1394
BstV1I	GCAGC	5 cut(s)	270, 404, 448, 1439, 1486
BsuRI	GGCC	4 cut(s)	551, 729, 812, 1450
BtrI	CACGTC	1 cut(s)	1039
BtsCI	GGATG	5 cut(s)	322, 862, 928, 1046, 1126
BtsI	GCAGTG	1 cut(s)	1203
BtsIMutI	CAGTG	5 cut(s)	186, 1203, 1275, 1427, 1470
Cac8I	GCNNGC	4 cut(s)	75, 720, 1073, 1077
CaiI	CAGNNNCTG	1 cut(s)	461
Cfr10I	RCCGGY	1 cut(s)	547
Cfr13I	GGNCC	2 cut(s)	121, 1112
CseI	GACGC	1 cut(s)	494
Csp6I	GTAC	6 cut(s)	65, 620, 792, 1009, 1183, 1391
CviAII	CATG	7 cut(s)	335, 530, 932, 986, 1072, 1186, 1456
CviQI	GTAC	6 cut(s)	65, 620, 792, 1009, 1183, 1391
DdeI	CTNAG	4 cut(s)	480, 602, 908, 1307
DpnI	GATC	5 cut(s)	271, 822, 1128, 1359, 1446
DpnII	GATC	5 cut(s)	269, 820, 1126, 1357, 1444
DraIII	CACNNNGTG	1 cut(s)	563
DriI	GACNNNNNGTC	2 cut(s)	1042, 1260
EaeI	YGGCCR	4 cut(s)	549, 727, 810, 1448
Eam1104I	CTCTTC	3 cut(s)	213, 1274, 1287
Eam1105I	GACNNNNNGTC	2 cut(s)	1042, 1260
EarI	CTCTTC	3 cut(s)	213, 1274, 1287
Eco31I	GGTCTC	1 cut(s)	1276
Eco47I	GGWCC	2 cut(s)	121, 1112
EcoRI	GAATTC	2 cut(s)	175, 785
EcoRII	CCWGG	3 cut(s)	644, 942, 1103
Esp3I	CGTCTC	1 cut(s)	1029
FaeI	CATG	7 cut(s)	338, 533, 935, 989, 1075, 1189, 1459
FalI	AAGNNNNNCTT	4 cut(s)	749, 781, 789, 821
FaqI	GGGAC	1 cut(s)	417
FatI	CATG	7 cut(s)	334, 529, 931, 985, 1071, 1185, 1455
FauI	CCCGC	1 cut(s)	662
FbaI	TGATCA	1 cut(s)	820
FblI	GTMKAC	3 cut(s)	607, 705, 1221
Fnu4HI	GCNGC	6 cut(s)	259, 393, 462, 534, 1428, 1475
FokI	GGATG	5 cut(s)	329, 869, 935, 1033, 1133
Fsp4HI	GCNGC	6 cut(s)	259, 393, 462, 534, 1428, 1475
FspBI	CTAG	6 cut(s)	284, 573, 579, 719, 845, 1172
GluI	GCNGC	6 cut(s)	259, 393, 462, 534, 1428, 1475
GsuI	CTGGAG	1 cut(s)	965
HaeIII	GGCC	4 cut(s)	551, 729, 812, 1450
HapII	CCGG	2 cut(s)	548, 1178
HgaI	GACGC	1 cut(s)	494
Hin1II	CATG	7 cut(s)	338, 533, 935, 989, 1075, 1189, 1459
HincII	GTYRAC	1 cut(s)	929
HindII	GTYRAC	1 cut(s)	929
HindIII	AAGCTT	2 cut(s)	110, 189
HinfI	GANTC	5 cut(s)	320, 359, 526, 634, 829
HpaII	CCGG	2 cut(s)	548, 1178
HphI	GGTGA	1 cut(s)	695
Hpy166II	GTNNAC	7 cut(s)	557, 608, 706, 929, 1011, 1222, 1482
Hpy188I	TCNGA	8 cut(s)	254, 358, 639, 828, 911, 1062, 1116, 1310
Hpy188III	TCNNGA	2 cut(s)	920, 1034
Hpy8I	GTNNAC	7 cut(s)	557, 608, 706, 929, 1011, 1222, 1482
Hpy99I	CGWCG	1 cut(s)	1266
HpyAV	CCTTC	5 cut(s)	307, 680, 751, 854, 1328
HpyCH4III	ACNGT	5 cut(s)	105, 382, 703, 1162, 1465
HpyCH4IV	ACGT	4 cut(s)	748, 1038, 1343, 1389
HpyF10VI	GCNNNNNNNGC	7 cut(s)	83, 255, 401, 485, 719, 1085, 1382
HpyF3I	CTNAG	4 cut(s)	480, 602, 908, 1307
HpySE526I	ACGT	4 cut(s)	748, 1038, 1343, 1389
Hsp92II	CATG	7 cut(s)	338, 533, 935, 989, 1075, 1189, 1459
Ksp22I	TGATCA	1 cut(s)	820
Kzo9I	GATC	5 cut(s)	269, 820, 1126, 1357, 1444
LguI	GCTCTTC	2 cut(s)	213, 1287
LmnI	GCTCC	3 cut(s)	401, 1435, 1471
Lsp1109I	GCAGC	5 cut(s)	270, 404, 448, 1439, 1486
LweI	GCATC	1 cut(s)	1111
MaeI	CTAG	6 cut(s)	284, 573, 579, 719, 845, 1172
MaeII	ACGT	4 cut(s)	748, 1038, 1343, 1389
MaeIII	GTNAC	2 cut(s)	1252, 1300
MalI	GATC	5 cut(s)	271, 822, 1128, 1359, 1446
MboI	GATC	5 cut(s)	269, 820, 1126, 1357, 1444
MboII	GAAGA	8 cut(s)	200, 300, 504, 992, 1049, 1291, 1304, 1350
MfeI	CAATTG	1 cut(s)	68
MlsI	TGGCCA	2 cut(s)	729, 812
MluNI	TGGCCA	2 cut(s)	729, 812
MmeI	TCCRAC	2 cut(s)	364, 1074
MnlI	CCTC	5 cut(s)	97, 235, 260, 1363, 1511
Mox20I	TGGCCA	2 cut(s)	729, 812
MscI	TGGCCA	2 cut(s)	729, 812
MseI	TTAA	5 cut(s)	26, 47, 444, 951, 1503
MslI	CAYNNNNRTG	1 cut(s)	921
Msp20I	TGGCCA	2 cut(s)	729, 812
MspA1I	CMGCKG	2 cut(s)	536, 1477
MspI	CCGG	2 cut(s)	548, 1178
MspR9I	CCNGG	4 cut(s)	646, 944, 1105, 1178
MunI	CAATTG	1 cut(s)	68
Mva1269I	GAATGC	1 cut(s)	1316
MvaI	CCWGG	3 cut(s)	646, 944, 1105
MwoI	GCNNNNNNNGC	7 cut(s)	83, 255, 401, 485, 719, 1085, 1382
NciI	CCSGG	1 cut(s)	1178
NdeII	GATC	5 cut(s)	269, 820, 1126, 1357, 1444
NheI	GCTAGC	1 cut(s)	718
NlaIII	CATG	7 cut(s)	338, 533, 935, 989, 1075, 1189, 1459
NlaIV	GGNNCC	1 cut(s)	248
NmeAIII	GCCGAG	1 cut(s)	694
NmuCI	GTSAC	2 cut(s)	1252, 1300
NspI	RCATGY	1 cut(s)	1075
PaeI	GCATGC	1 cut(s)	1075
PciSI	GCTCTTC	2 cut(s)	213, 1287
PctI	GAATGC	1 cut(s)	1316
PfeI	GAWTC	5 cut(s)	320, 359, 526, 634, 829
PkrI	GCNGC	6 cut(s)	260, 394, 463, 535, 1429, 1476
PshAI	GACNNNNGTC	1 cut(s)	613
Psp6I	CCWGG	3 cut(s)	644, 942, 1103
PspGI	CCWGG	3 cut(s)	644, 942, 1103
PspN4I	GGNNCC	1 cut(s)	248
PspPI	GGNCC	2 cut(s)	121, 1112
PstNI	CAGNNNCTG	1 cut(s)	461
PvuII	CAGCTG	1 cut(s)	1477
RsaI	GTAC	6 cut(s)	66, 621, 793, 1010, 1184, 1392
RsaNI	GTAC	6 cut(s)	65, 620, 792, 1009, 1183, 1391
RseI	CAYNNNNRTG	1 cut(s)	921
SapI	GCTCTTC	2 cut(s)	213, 1287
SaqAI	TTAA	5 cut(s)	26, 47, 444, 951, 1503
SatI	GCNGC	6 cut(s)	259, 393, 462, 534, 1428, 1475
Sau3AI	GATC	5 cut(s)	269, 820, 1126, 1357, 1444
Sau96I	GGNCC	2 cut(s)	121, 1112
ScrFI	CCNGG	4 cut(s)	646, 944, 1105, 1178
SfaNI	GCATC	1 cut(s)	1111
SfcI	CTRYAG	1 cut(s)	1394
SinI	GGWCC	2 cut(s)	121, 1112
SmiMI	CAYNNNNRTG	1 cut(s)	921
SphI	GCATGC	1 cut(s)	1075
SsiI	CCGC	2 cut(s)	534, 655
SspMI	CTAG	6 cut(s)	284, 573, 579, 719, 845, 1172
StyD4I	CCNGG	4 cut(s)	644, 942, 1103, 1176
TaaI	ACNGT	5 cut(s)	105, 382, 703, 1162, 1465
TaiI	ACGT	4 cut(s)	751, 1041, 1346, 1392
TaqI	TCGA	3 cut(s)	1033, 1129, 1438
TatI	WGTACW	4 cut(s)	619, 791, 1008, 1182
TauI	GCSGC	1 cut(s)	536
TfiI	GAWTC	5 cut(s)	320, 359, 526, 634, 829
Tru1I	TTAA	5 cut(s)	26, 47, 444, 951, 1503
Tru9I	TTAA	5 cut(s)	26, 47, 444, 951, 1503
TscAI	CASTG	5 cut(s)	186, 1210, 1275, 1427, 1470
TseFI	GTSAC	2 cut(s)	1252, 1300
TseI	GCWGC	5 cut(s)	258, 392, 461, 1427, 1474
Tsp45I	GTSAC	2 cut(s)	1252, 1300
TspDTI	ATGAA	7 cut(s)	333, 873, 882, 992, 1202, 1292, 1472
TspGWI	ACGGA	1 cut(s)	12
TspRI	CASTG	5 cut(s)	186, 1210, 1275, 1427, 1470
VpaK11BI	GGWCC	2 cut(s)	121, 1112
XapI	RAATTY	4 cut(s)	175, 490, 772, 785
XceI	RCATGY	1 cut(s)	1075
XmiI	GTMKAC	3 cut(s)	607, 705, 1221
XspI	CTAG	6 cut(s)	284, 573, 579, 719, 845, 1172

Using CommOnly database (standard laboratory enzymes). Scanned on CDS sequence.