Basic Information

Type: gene

Biological Identity

Species

rosa_samantha

Chromosome

Chr2D

Orthogroup ID

OG0002219

Physical Location & Seq

Genomic Coordinates Reverse (-)

88066809 .. 88070617

Gene Structure 3809 bp

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UTR

Exon/CDS

Intron

Transcript / Protein ID

Rh2DG655700.1

Sequence Viewer

CDS Protein Promoter (2k)

Length: 1419 bp

ATGGGGGCTTACAGCAACGACGGCGTTTTGTTGTCTCCGATATACAGTTTGAGTTCCGGCCAAGACCTCCGTTCCTCCAAGCACGTCTACGACAATGTCCACGGCAACATTTACCTCGATCCGCTTTCTTTGAAGTTTATTGATACTGAGGAGTTTCAGAGGCTTAGAGACCTGAAACAACTCGGAGTGACAAACTCAGTTTTTCCTGGGGCTGTGCATTCCAGGTTTGAGCATTCGCTTGGTGTCTATTGGCTTGCTGGTGAAGCTGTTCAGAGGCTCAAAAATTATCAGGGTCTCGAACTTGATATTGACCGCTTTGATCTCCAGACAGTAAAAATTGCAGGTCTATTGCATGATGTTGGCCATGGGCCTTTCAGTCACTTGTTTGAACGGGAGTTTCTTCCACGGGTTGAGAGTGGTTCCCATTGGTCTCATGAGGATATGTCAGTTAAGATGATTGACCACATTGTTAATCAGCATCATATTGATATTGACCCTGAGATGATTAAAAGAGTTAAGGAGATGATACTTGCTTCTGAAAATGCCAATGCAAAAAGCTCCAGGGAGAAGCCTTTTCTGTATGAAATTGTGGCAAATGGCCGTAATGGTATTGATGTGGACAAGTTTGATTATATTGTCCGTGACAGCCGAGCTTGTGGGCTGGGATGTAATTTTGAGTTTCGAAGATTAATGGAAACAATGCGGGTTTTGGATGATGAGATCTGCTACCGTGCTAAAGACTATCTCACCGTCCACAAATTATTTGTCACTCGGGCGGATTTGTATCGAACAGTTTATACACATCCAAAAGTAAAGGCCATAGAGCTTATGGTGGTGGATGCCCTGGTGAAAGCAAATGATTATCTACAAATATCATCTCACATTGAGGATCCAGCTGAATATTGGAAGTTAGATGACTCAATAATCAGAACTATTGAGACAGCTCCGAATGAAGAGCTAAAGGAGGCTAGGGACTTAATCCTGCGCATTAGGAGGAGAAATCTCTATCAGTTCTGCAATGAGTATGCTGTACCCAAGGAGCAAGTGGAACATTTCAAAAACGTCACTGCTCAAGATATTGTTTGTTCTCAGACAAATGGAGGTGTTATATTGAAAGAGGAGGATGTTGCTGTTACCAACATCAGGATTGATTTGACTCGCGGAAGTCAAAATCCCCTTCAAAGCATCAACTTTTTCCAGGACTATGAGAGTAAGGAGAAATTCCCAATACGTGATGATCGCATTAGTCACTTGCTGCCTACATCGTATCAAGATTTGATAGTGAGGGTCTATTCAAAAAAGCCTGAGCTGGTTGGGCCAATCTCTGAGGCTTTTGAAAATTTCCAGGTAAAAACATATGGGGTCAAGGCACAAGTACATCCCACACCAGAGAAAAAGAAACGCCCTCGTGGTTTCTAA

Functional Annotation

Gene Ontology

Molecular Function
Biological Process
Cellular Component

No ontology terms assigned.

KEGG Pathways

Metabolic & Signaling

No pathways identified.

Pfam Domains

Protein Families

Protein Analysis

472

Amino Acids

54.52

Weight (kDa)

6.16

Isoelectric Point (pI)

38.2

Instability Index

Protein Domains (Pfam)

Domain Name	Pfam ID	Position	E-value	Description
HD	PF01966	75 - 167	1.8e-11	HD domain
HD_assoc_2	PF19276	227 - 338	4.1e-07	HD associated region

Hydrophobicity Profile (Kyte-Doolittle)

AI Structure Prediction Report

Calculating structure properties...

Genomic Context

Gene Family Tree

Species Filter

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Align Labels (Cladogram)

Image

Tree File

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Publication-ready

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Orthologous Genes (Group: OG0002219)

Species	Orthologous Gene IDs
arabidopsis_thaliana	AT5G40270 AT5G40270 AT5G40290
fragaria_vesca	FvH4_4g34540 FvH4_4g34540 FvH4_6g50610 FvH4_6g50610 FvH4_6g50610
malus_domestica	MD09G1034800.v1.1 MD16G1092000.v1.1
prunus_persica	Prupe.1G254300_v2.0.a1 Prupe.1G254300_v2.0.a1 Prupe.1G254300_v2.0.a1 Prupe.3G283700_v2.0.a1 Prupe.3G283700_v2.0.a1 Prupe.3G283700_v2.0.a1
pyrus_communis	pycom111g02770 pycom16g07850
rosa_chinensis	RchiOBHm_Chr2g0171401 RchiOBHm_Chr4g0443921
rosa_laevigata	RLG00000005877 RLG00000022010
rosa_multiflora	Rmu_co8513743.1_g000001 Rmu_sc0007646.1_g000008 Rmu_sc0030144.1_g000004
rosa_roxburghii	Rroxscaffold_2G00080690 Rroxscaffold_5G00384470
rosa_rugosa	Rorug02G0553800 Rorug02G0553800 Rorug04G0348200 Rorug04G0348200 Rorug04G0348200
rosa_samantha	Rh2AG628200 Rh2BG638600 Rh2CG607300 Rh2DG649600 Rh2DG655700 Rh4AG408800 Rh4BG419900 Rh4CG433900 Rh4DG414900
rosa_wichuraiana	Rw2G052030 Rw4G035110

Restriction Enzyme Sites

1 / 10

Enzyme	Recognition Site	Cut Count	Positions (bp)
Acc16I	TGCGCA	1 cut(s)	986
Acc36I	ACCTGC	1 cut(s)	332
AccI	GTMKAC	1 cut(s)	87
AccII	CGCG	1 cut(s)	1161
AciI	CCGC	5 cut(s)	122, 313, 703, 776, 1161
AclWI	GGATC	3 cut(s)	113, 884, 897
AcoI	YGGCCR	3 cut(s)	58, 361, 598
AcsI	RAATTY	2 cut(s)	1220, 1339
AfaI	GTAC	2 cut(s)	1032, 1377
AfiI	CCNNNNNNNGG	1 cut(s)	1143
AgsI	TTSAA	7 cut(s)	133, 389, 1057, 1114, 1181, 1296, 1337
AjiI	CACGTC	1 cut(s)	85
AjnI	CCWGG	6 cut(s)	205, 221, 560, 843, 1197, 1344
AluBI	AGCT	8 cut(s)	266, 558, 653, 826, 896, 944, 958, 1309
AluI	AGCT	8 cut(s)	266, 558, 653, 826, 896, 944, 958, 1309
Alw26I	GTCTC	5 cut(s)	39, 162, 299, 435, 932
AlwI	GGATC	3 cut(s)	113, 884, 897
Ama87I	CYCGRG	1 cut(s)	771
AoxI	GGCC	6 cut(s)	58, 361, 368, 598, 816, 1316
ApeKI	GCWGC	1 cut(s)	1255
ApoI	RAATTY	2 cut(s)	1220, 1339
ArsI	GACNNNNNNTTYG	2 cut(s)	750, 782
AseI	ATTAAT	1 cut(s)	689
Asp700I	GAANNNNTTC	1 cut(s)	267
AspLEI	GCGC	1 cut(s)	987
AspS9I	GGNCC	2 cut(s)	368, 1316
AsuHPI	GGTGA	3 cut(s)	272, 739, 859
AsuII	TTCGAA	1 cut(s)	682
AvaI	CYCGRG	1 cut(s)	771
BalI	TGGCCA	1 cut(s)	363
BamHI	GGATCC	1 cut(s)	889
BauI	CACGAG	1 cut(s)	1407
BbvI	GCAGC	1 cut(s)	1242
BceAI	ACGGC	3 cut(s)	37, 118, 585
BciT130I	CCWGG	6 cut(s)	207, 223, 562, 845, 1199, 1346
BcoDI	GTCTC	5 cut(s)	39, 162, 299, 435, 932
BfaI	CTAG	1 cut(s)	969
BfuAI	ACCTGC	1 cut(s)	332
BglII	AGATCT	1 cut(s)	720
BisI	GCNGC	1 cut(s)	1256
BlsI	GCNGC	1 cut(s)	1257
Bme1390I	CCNGG	6 cut(s)	207, 223, 562, 845, 1199, 1346
BmeT110I	CYCGRG	1 cut(s)	771
BmgBI	CACGTC	1 cut(s)	85
BmgT120I	GGNCC	2 cut(s)	368, 1316
BmiI	GGNNCC	2 cut(s)	421, 891
BmrFI	CCNGG	6 cut(s)	207, 223, 562, 845, 1199, 1346
BmsI	GCATC	3 cut(s)	487, 829, 1194
BpmI	CTGGAG	2 cut(s)	308, 544
Bpu10I	CCTNAGC	1 cut(s)	1305
Bpu14I	TTCGAA	1 cut(s)	682
BpuEI	CTTGAG	1 cut(s)	1056
BsaAI	YACGTR	1 cut(s)	1232
BsaBI	GATNNNNATC	1 cut(s)	783
BsaI	GGTCTC	3 cut(s)	162, 299, 435
BsaJI	CCNNGG	7 cut(s)	100, 206, 364, 404, 561, 843, 1035
Bsc4I	CCNNNNNNNGG	1 cut(s)	1143
Bse3DI	GCAATG	1 cut(s)	1024
Bse8I	GATNNNNATC	1 cut(s)	783
BseBI	CCWGG	6 cut(s)	207, 223, 562, 845, 1199, 1346
BseDI	CCNNGG	7 cut(s)	100, 206, 364, 404, 561, 843, 1035
BseGI	GGATG	6 cut(s)	671, 718, 802, 844, 1129, 1378
BseJI	GATNNNNATC	1 cut(s)	783
BseLI	CCNNNNNNNGG	1 cut(s)	1143
BseMI	GCAATG	1 cut(s)	1024
BseMII	CTCAG	6 cut(s)	138, 210, 489, 1103, 1296, 1317
BseRI	GAGGAG	3 cut(s)	164, 1009, 1133
BseXI	GCAGC	1 cut(s)	1242
BseYI	CCCAGC	1 cut(s)	661
Bsh1236I	CGCG	1 cut(s)	1161
BshFI	GGCC	6 cut(s)	60, 363, 370, 600, 818, 1318
BsiHKCI	CYCGRG	1 cut(s)	771
BsiSI	CCGG	1 cut(s)	57
BslFI	GGGAC	1 cut(s)	986
BslI	CCNNNNNNNGG	1 cut(s)	1143
BsmAI	GTCTC	5 cut(s)	39, 162, 299, 435, 932
BsmFI	GGGAC	1 cut(s)	986
BsmI	GAATGC	2 cut(s)	217, 232
BsnI	GGCC	6 cut(s)	60, 363, 370, 600, 818, 1318
Bso31I	GGTCTC	3 cut(s)	162, 299, 435
BsoBI	CYCGRG	1 cut(s)	771
Bsp119I	TTCGAA	1 cut(s)	682
Bsp143I	GATC	5 cut(s)	118, 319, 720, 889, 1237
Bsp19I	CCATGG	1 cut(s)	364
BspACI	CCGC	5 cut(s)	122, 313, 703, 776, 1161
BspANI	GGCC	6 cut(s)	60, 363, 370, 600, 818, 1318
BspCNI	CTCAG	6 cut(s)	139, 209, 490, 1102, 1297, 1318
BspFNI	CGCG	1 cut(s)	1161
BspHI	TCATGA	1 cut(s)	433
BspLI	GGNNCC	2 cut(s)	421, 891
BspMI	ACCTGC	1 cut(s)	332
BspPI	GGATC	3 cut(s)	113, 884, 897
BspQI	GCTCTTC	1 cut(s)	948
BspT104I	TTCGAA	1 cut(s)	682
BspTNI	GGTCTC	3 cut(s)	162, 299, 435
BsrDI	GCAATG	1 cut(s)	1024
BssECI	CCNNGG	7 cut(s)	100, 206, 364, 404, 561, 843, 1035
BssMI	GATC	5 cut(s)	118, 319, 720, 889, 1237
BssSI	CACGAG	1 cut(s)	1407
BssT1I	CCWWGG	2 cut(s)	364, 1035
Bst2BI	CACGAG	1 cut(s)	1407
Bst2UI	CCWGG	6 cut(s)	207, 223, 562, 845, 1199, 1346
Bst4CI	ACNGT	5 cut(s)	47, 331, 731, 751, 793
Bst6I	CTCTTC	1 cut(s)	948
BstBAI	YACGTR	1 cut(s)	1232
BstBI	TTCGAA	1 cut(s)	682
BstC8I	GCNNGC	1 cut(s)	255
BstDEI	CTNAG	7 cut(s)	147, 164, 196, 498, 1089, 1305, 1326
BstDSI	CCRYGG	3 cut(s)	100, 364, 404
BstF5I	GGATG	6 cut(s)	671, 718, 802, 844, 1129, 1378
BstFNI	CGCG	1 cut(s)	1161
BstHHI	GCGC	1 cut(s)	987
BstKTI	GATC	5 cut(s)	121, 322, 723, 892, 1240
BstMAI	GTCTC	5 cut(s)	39, 162, 299, 435, 932
BstMBI	GATC	5 cut(s)	118, 319, 720, 889, 1237
BstMWI	GCNNNNNNNGC	3 cut(s)	21, 263, 1315
BstNI	CCWGG	6 cut(s)	207, 223, 562, 845, 1199, 1346
BstSCI	CCNGG	6 cut(s)	205, 221, 560, 843, 1197, 1344
BstUI	CGCG	1 cut(s)	1161
BstV1I	GCAGC	1 cut(s)	1242
BstX2I	RGATCY	2 cut(s)	720, 889
BstYI	RGATCY	2 cut(s)	720, 889
BsuRI	GGCC	6 cut(s)	60, 363, 370, 600, 818, 1318
BtgI	CCRYGG	3 cut(s)	100, 364, 404
BtrI	CACGTC	1 cut(s)	85
BtsCI	GGATG	6 cut(s)	671, 718, 802, 844, 1129, 1378
BtsI	GCAGTG	1 cut(s)	1065
BtsIMutI	CAGTG	1 cut(s)	1065
BveI	ACCTGC	1 cut(s)	332
Cac8I	GCNNGC	1 cut(s)	255
CciI	TCATGA	1 cut(s)	433
CfoI	GCGC	1 cut(s)	987
Cfr13I	GGNCC	2 cut(s)	368, 1316
Csp6I	GTAC	2 cut(s)	1031, 1376
CviAII	CATG	3 cut(s)	353, 365, 434
CviQI	GTAC	2 cut(s)	1031, 1376
DdeI	CTNAG	7 cut(s)	147, 164, 196, 498, 1089, 1305, 1326
DpnI	GATC	5 cut(s)	120, 321, 722, 891, 1239
DpnII	GATC	5 cut(s)	118, 319, 720, 889, 1237
EaeI	YGGCCR	3 cut(s)	58, 361, 598
Eam1104I	CTCTTC	1 cut(s)	948
EarI	CTCTTC	1 cut(s)	948
EciI	GGCGGA	1 cut(s)	791
Eco130I	CCWWGG	2 cut(s)	364, 1035
Eco31I	GGTCTC	3 cut(s)	162, 299, 435
Eco88I	CYCGRG	1 cut(s)	771
EcoRII	CCWGG	6 cut(s)	205, 221, 560, 843, 1197, 1344
EcoT14I	CCWWGG	2 cut(s)	364, 1035
ErhI	CCWWGG	2 cut(s)	364, 1035
FaeI	CATG	3 cut(s)	356, 368, 437
FaqI	GGGAC	1 cut(s)	986
FatI	CATG	3 cut(s)	352, 364, 433
FauI	CCCGC	1 cut(s)	696
FauNDI	CATATG	1 cut(s)	1357
FblI	GTMKAC	1 cut(s)	87
Fnu4HI	GCNGC	1 cut(s)	1256
FokI	GGATG	6 cut(s)	678, 725, 789, 851, 1136, 1365
Fsp4HI	GCNGC	1 cut(s)	1256
FspBI	CTAG	1 cut(s)	969
FspI	TGCGCA	1 cut(s)	986
GlaI	GCGC	1 cut(s)	986
GluI	GCNGC	1 cut(s)	1256
GsaI	CCCAGC	1 cut(s)	665
GsuI	CTGGAG	2 cut(s)	308, 544
HaeIII	GGCC	6 cut(s)	60, 363, 370, 600, 818, 1318
HapII	CCGG	1 cut(s)	57
HhaI	GCGC	1 cut(s)	987
Hin1II	CATG	3 cut(s)	356, 368, 437
Hin6I	GCGC	1 cut(s)	985
HinP1I	GCGC	1 cut(s)	985
HinfI	GANTC	2 cut(s)	917, 1156
HpaII	CCGG	1 cut(s)	57
HphI	GGTGA	3 cut(s)	272, 739, 859
Hpy166II	GTNNAC	4 cut(s)	88, 100, 619, 754
Hpy188I	TCNGA	9 cut(s)	39, 159, 185, 273, 538, 929, 948, 1092, 1327
Hpy188III	TCNNGA	6 cut(s)	296, 325, 434, 1073, 1144, 1271
Hpy8I	GTNNAC	4 cut(s)	88, 100, 619, 754
Hpy99I	CGWCG	1 cut(s)	23
HpyAV	CCTTC	1 cut(s)	1187
HpyCH4III	ACNGT	5 cut(s)	47, 331, 731, 751, 793
HpyCH4IV	ACGT	3 cut(s)	84, 1062, 1231
HpyCH4V	TGCA	5 cut(s)	217, 341, 352, 551, 1017
HpyF10VI	GCNNNNNNNGC	3 cut(s)	21, 263, 1315
HpyF3I	CTNAG	7 cut(s)	147, 164, 196, 498, 1089, 1305, 1326
HpySE526I	ACGT	3 cut(s)	84, 1062, 1231
Hsp92II	CATG	3 cut(s)	356, 368, 437
HspAI	GCGC	1 cut(s)	985
Kzo9I	GATC	5 cut(s)	118, 319, 720, 889, 1237
LguI	GCTCTTC	1 cut(s)	948
LmnI	GCTCC	3 cut(s)	563, 949, 1039
Lsp1109I	GCAGC	1 cut(s)	1242
LweI	GCATC	3 cut(s)	487, 829, 1194
MaeI	CTAG	1 cut(s)	969
MaeII	ACGT	3 cut(s)	84, 1062, 1231
MaeIII	GTNAC	7 cut(s)	187, 377, 641, 766, 1063, 1132, 1247
MalI	GATC	5 cut(s)	120, 321, 722, 891, 1239
MboI	GATC	5 cut(s)	118, 319, 720, 889, 1237
MboII	GAAGA	3 cut(s)	392, 696, 965
MflI	RGATCY	2 cut(s)	720, 889
MlsI	TGGCCA	1 cut(s)	363
MluCI	AATT	7 cut(s)	283, 336, 585, 670, 758, 1220, 1339
MluNI	TGGCCA	1 cut(s)	363
MlyI	GAGTC	2 cut(s)	911, 1150
Mox20I	TGGCCA	1 cut(s)	363
MroXI	GAANNNNTTC	1 cut(s)	267
MscI	TGGCCA	1 cut(s)	363
MseI	TTAA	6 cut(s)	450, 471, 507, 516, 689, 977
Msp20I	TGGCCA	1 cut(s)	363
MspA1I	CMGCKG	1 cut(s)	896
MspI	CCGG	1 cut(s)	57
MspR9I	CCNGG	6 cut(s)	207, 223, 562, 845, 1199, 1346
Mva1269I	GAATGC	2 cut(s)	217, 232
MvaI	CCWGG	6 cut(s)	207, 223, 562, 845, 1199, 1346
MvnI	CGCG	1 cut(s)	1161
MwoI	GCNNNNNNNGC	3 cut(s)	21, 263, 1315
NcoI	CCATGG	1 cut(s)	364
NdeI	CATATG	1 cut(s)	1357
NdeII	GATC	5 cut(s)	118, 319, 720, 889, 1237
NlaIII	CATG	3 cut(s)	356, 368, 437
NlaIV	GGNNCC	2 cut(s)	421, 891
NmeAIII	GCCGAG	1 cut(s)	674
NmuCI	GTSAC	6 cut(s)	187, 377, 641, 766, 1063, 1247
NsbI	TGCGCA	1 cut(s)	986
NspV	TTCGAA	1 cut(s)	682
PagI	TCATGA	1 cut(s)	433
PciSI	GCTCTTC	1 cut(s)	948
PctI	GAATGC	2 cut(s)	217, 232
PdmI	GAANNNNTTC	1 cut(s)	267
PflFI	GACNNNGTC	1 cut(s)	95
PfoI	TCCNGGA	1 cut(s)	1197
PkrI	GCNGC	1 cut(s)	1257
PleI	GAGTC	2 cut(s)	911, 1150
PpsI	GAGTC	2 cut(s)	911, 1150
Ppu21I	YACGTR	1 cut(s)	1232
PshBI	ATTAAT	1 cut(s)	689
Psp6I	CCWGG	6 cut(s)	205, 221, 560, 843, 1197, 1344
PspFI	CCCAGC	1 cut(s)	661
PspGI	CCWGG	6 cut(s)	205, 221, 560, 843, 1197, 1344
PspN4I	GGNNCC	2 cut(s)	421, 891
PspPI	GGNCC	2 cut(s)	368, 1316
PsrI	GAACNNNNNNTAC	2 cut(s)	781, 813
PsuI	RGATCY	2 cut(s)	720, 889
PsyI	GACNNNGTC	1 cut(s)	95
PvuII	CAGCTG	1 cut(s)	896
RsaI	GTAC	2 cut(s)	1032, 1377
RsaNI	GTAC	2 cut(s)	1031, 1376
SapI	GCTCTTC	1 cut(s)	948
SaqAI	TTAA	6 cut(s)	450, 471, 507, 516, 689, 977
SatI	GCNGC	1 cut(s)	1256
Sau3AI	GATC	5 cut(s)	118, 319, 720, 889, 1237
Sau96I	GGNCC	2 cut(s)	368, 1316
SchI	GAGTC	2 cut(s)	911, 1150
ScrFI	CCNGG	6 cut(s)	207, 223, 562, 845, 1199, 1346
SfaNI	GCATC	3 cut(s)	487, 829, 1194
SfuI	TTCGAA	1 cut(s)	682
SmlI	CTYRAG	1 cut(s)	1071
SmoI	CTYRAG	1 cut(s)	1071
Sse9I	AATT	7 cut(s)	283, 336, 585, 670, 758, 1220, 1339
SsiI	CCGC	5 cut(s)	122, 313, 703, 776, 1161
SspI	AATATT	1 cut(s)	902
SspMI	CTAG	1 cut(s)	969
StyD4I	CCNGG	6 cut(s)	205, 221, 560, 843, 1197, 1344
StyI	CCWWGG	2 cut(s)	364, 1035
TaaI	ACNGT	5 cut(s)	47, 331, 731, 751, 793
TaiI	ACGT	3 cut(s)	87, 1065, 1234
TaqI	TCGA	4 cut(s)	117, 297, 682, 787
TasI	AATT	7 cut(s)	283, 336, 585, 670, 758, 1220, 1339
TatI	WGTACW	1 cut(s)	1375
Tru1I	TTAA	6 cut(s)	450, 471, 507, 516, 689, 977
Tru9I	TTAA	6 cut(s)	450, 471, 507, 516, 689, 977
TscAI	CASTG	1 cut(s)	1072
TseFI	GTSAC	6 cut(s)	187, 377, 641, 766, 1063, 1247
TseI	GCWGC	1 cut(s)	1255
Tsp45I	GTSAC	6 cut(s)	187, 377, 641, 766, 1063, 1247
TspDTI	ATGAA	2 cut(s)	597, 966
TspGWI	ACGGA	2 cut(s)	59, 629
TspRI	CASTG	1 cut(s)	1072
Tth111I	GACNNNGTC	1 cut(s)	95
VspI	ATTAAT	1 cut(s)	689
XapI	RAATTY	2 cut(s)	1220, 1339
XcmI	CCANNNNNNNNNTGG	3 cut(s)	826, 900, 1042
XmiI	GTMKAC	1 cut(s)	87
XmnI	GAANNNNTTC	1 cut(s)	267
XspI	CTAG	1 cut(s)	969

Using CommOnly database (standard laboratory enzymes). Scanned on CDS sequence.