Basic Information

Type: gene

Biological Identity

Species

pyrus_communis

Chromosome

Chr4

Orthogroup ID

OG0004490

Physical Location & Seq

Genomic Coordinates Forward (+)

2460577 .. 2465222

Gene Structure 4646 bp

Loading structure...

UTR

Exon/CDS

Intron

Transcript / Protein ID

pycom04g02710.1

Sequence Viewer

CDS Protein Promoter (2k)

Length: 1083 bp

ATGTATCCCACAAACTCATCCTCCTCGCCTCAAAAGCCCATGGCCCCGAGACCAAGCACAACGGGCCTGACCCGCTACGGCTCCGCCCCAGGATCCCTCCTAAACTCAGCCGTGGACTCCATCATGGGAGCCGCCGCCGACCGCGAATTCTCATCCCTCCGACCGCAGCACCTACGAGGCCACTACTTCTCCGGAGACGGTGGCGGCGACTCTCCCTCCTTCACTAGCTCCGAGTCCAGCTTCAAAGTCAACTCATCCAACGGCAGCCACAGGGACACCAGCACCACCAAGCCACTGCTCCGATCGCACGGTCTAAATGAGATTGGATCTTCCAACTCTTCTTCTTGTTCTTCGACATCGTCGGCGGCTCTGGTTCGCCAGCGAAGCTCTCCAGCTGGCTTCTTAAGCCATCTCACCGCCCCTGAGAGCAATGGAGGATTCTCAGTTACAAGAGGAACTGGAAGCTATAGATCACAAGGGGGCAATAATGGTGGTCCCGGAGTAACAAGATTAAAGTCTCAGTTGAGCTTTAGTGGGCAAGATTCTCTTTCTCAAATCTCTGAAGTAAGTCAAAATGCTGGTGAAGGTGTTAGCTCTAAGAACGGCCACCATAGTGCCATGCATTCTTATCCTGCCACTAGCTTCGGGATGGAATCCTGGGATAATACCAATTCGATTGTCTTTTCAGCTCCGCCAAGTAAACGAGCTAAAAATATGGATGTTGACATATTTGACTGCCTCAATACTTTAGAATCTCAGTTTAGCCTTCCCCAGACGTCGTTAGAAATGGCAACAGTAGAGAAGCTACTACACATTCCTGAAGATTCTGTTCCTTGCAAAATCCGTGCCAAGCGTGGCTGTGCAACTCATCCGAGAAGCATTGCAGAGAGGGAGAGAAGAACTAGAATAAGCGGGAAGTTAAAAAAACTACAAGACCTTGTTCCTAACATGGATAAGCAAACGAGCTATGCGGACATGTTGGATTTAGCCGTGCAACATATCAAAGGCCTTCAACATCAAATCCAGGATCTTCATAATGATCTTGAGCATTGCACTTGCGGCTGCAAACCAACCGCGAAATGA

Functional Annotation

Gene Ontology

Molecular Function
Biological Process
Cellular Component

GO:0000976 GO:0000981 GO:0001046 GO:0001047 GO:0001067 GO:0001228 GO:0003674 GO:0003676 GO:0003677 GO:0003690 GO:0003700 GO:0005488 GO:0005575 GO:0005622 GO:0005623 GO:0005634 GO:0005737 GO:0005829 GO:0006139 GO:0006351 GO:0006355 GO:0006357 GO:0006366 GO:0006725 GO:0006807 GO:0008150 GO:0008152 GO:0009058 GO:0009059 GO:0009889 GO:0009891 GO:0009893 GO:0009987 GO:0010119 GO:0010467 GO:0010468 GO:0010556 GO:0010557 GO:0010604 GO:0010628 GO:0016070 GO:0018130 GO:0019219 GO:0019222 GO:0019438 GO:0031323 GO:0031325 GO:0031326 GO:0031328 GO:0032774 GO:0034641 GO:0034645 GO:0034654 GO:0043170 GO:0043226 GO:0043227 GO:0043229 GO:0043231 GO:0043565 GO:0044212 GO:0044237 GO:0044238 GO:0044249 GO:0044260 GO:0044271 GO:0044424 GO:0044444 GO:0044464 GO:0045893 GO:0045935 GO:0045944 GO:0046483 GO:0048518 GO:0048522 GO:0050789 GO:0050794 GO:0051171 GO:0051173 GO:0051252 GO:0051254 GO:0060255 GO:0065007 GO:0071704 GO:0080090 GO:0090304 GO:0097159 GO:0097659 GO:0140110 GO:1901360 GO:1901362 GO:1901363 GO:1901576 GO:1902680 GO:1903506 GO:1903508 GO:1990837 GO:2000112 GO:2001141

KEGG Pathways

Metabolic & Signaling

No pathways identified.

Pfam Domains

Protein Families

HLH

Protein Analysis

361

Amino Acids

38.48

Weight (kDa)

8.93

Isoelectric Point (pI)

54.34

Instability Index

Protein Domains (Pfam)

Domain Name	Pfam ID	Position	E-value	Description
HLH	PF00010	293 - 338	3.9e-09	Helix-loop-helix DNA-binding domain

Hydrophobicity Profile (Kyte-Doolittle)

AI Structure Prediction Report

Calculating structure properties...

Genomic Context

Gene Family Tree

Species Filter

Style Settings

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Align Labels (Cladogram)

Image

Tree File

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Publication-ready

iTOL Interactive Tree Of Life · Most popular EvolView Flexible annotations MEGA Desktop phylogenetics suite FigTree Desktop tree editor

Orthologous Genes (Group: OG0004490)

Species	Orthologous Gene IDs
arabidopsis_thaliana	AT1G05805
fragaria_vesca	FvH4_4g11732 FvH4_4g11732 FvH4_4g11732 FvH4_4g11732 FvH4_4g11732 FvH4_4g11732 FvH4_4g11732 FvH4_4g11732 FvH4_4g11740
malus_domestica	MD04G1033000.v1.1 MD05G1062100.v1.1 MD07G1058200.v1.1
prunus_persica	Prupe.1G181300_v2.0.a1 Prupe.1G181300_v2.0.a1
pyrus_communis	pycom04g02710 pycom16g26360
rosa_chinensis	RchiOBHm_Chr4g0409001
rosa_laevigata	RLG00000008598
rosa_multiflora	Rmu_sc0006766.1_g000012
rosa_roxburghii	Rroxscaffold_2G00121800 Rroxscaffold_3G00239380 Rroxscaffold_5G00353460 Rroxscaffold_7G00197350
rosa_rugosa	Rorug04G0089200 Rorug04G0089300 Rorug04G0089400 Rorug04G0089500
rosa_samantha	Rh4AG153300 Rh4BG151400 Rh4CG161700 Rh4DG146600

Restriction Enzyme Sites

1 / 10

Enzyme	Recognition Site	Cut Count	Positions (bp)
AatII	GACGTC	1 cut(s)	779
AccII	CGCG	2 cut(s)	144, 1076
AccIII	TCCGGA	1 cut(s)	191
AclWI	GGATC	4 cut(s)	87, 100, 334, 1035
AcoI	YGGCCR	1 cut(s)	604
AcsI	RAATTY	1 cut(s)	146
AcuI	CTGAAG	2 cut(s)	582, 840
AcyI	GRCGYC	1 cut(s)	776
AflII	CTTAAG	1 cut(s)	403
AflIII	ACRYGT	1 cut(s)	975
AgsI	TTSAA	2 cut(s)	244, 1013
AjnI	CCWGG	3 cut(s)	88, 656, 1023
AleI	CACNNNNGTG	1 cut(s)	612
Alw26I	GTCTC	3 cut(s)	43, 189, 522
AlwI	GGATC	4 cut(s)	87, 100, 334, 1035
Ama87I	CYCGRG	1 cut(s)	46
Aor13HI	TCCGGA	1 cut(s)	191
AoxI	GGCC	5 cut(s)	42, 64, 178, 604, 1006
ApeKI	GCWGC	3 cut(s)	166, 264, 1062
ApoI	RAATTY	1 cut(s)	146
AspS9I	GGNCC	3 cut(s)	43, 64, 494
AsuC2I	CCSGG	1 cut(s)	498
AsuHPI	GGTGA	2 cut(s)	406, 593
AvaI	CYCGRG	1 cut(s)	46
AvaII	GGWCC	1 cut(s)	494
BamHI	GGATCC	1 cut(s)	92
BbvI	GCAGC	3 cut(s)	178, 276, 1049
BccI	CCATC	3 cut(s)	128, 417, 643
BceAI	ACGGC	5 cut(s)	94, 95, 277, 619, 974
BciT130I	CCWGG	3 cut(s)	90, 658, 1025
BciVI	GTATCC	1 cut(s)	15
BcnI	CCSGG	1 cut(s)	498
BcoDI	GTCTC	3 cut(s)	43, 189, 522
BfaI	CTAG	3 cut(s)	225, 639, 903
BfmI	CTRYAG	1 cut(s)	466
BfrI	CTTAAG	1 cut(s)	403
BfuI	GTATCC	1 cut(s)	15
BisI	GCNGC	8 cut(s)	132, 135, 167, 205, 265, 366, 1060, 1063
BlsI	GCNGC	8 cut(s)	133, 136, 168, 206, 266, 367, 1061, 1064
Bme1390I	CCNGG	4 cut(s)	90, 498, 658, 1025
Bme18I	GGWCC	1 cut(s)	494
BmeT110I	CYCGRG	1 cut(s)	46
BmgT120I	GGNCC	3 cut(s)	43, 64, 494
BmiI	GGNNCC	5 cut(s)	45, 82, 94, 130, 496
BmrFI	CCNGG	4 cut(s)	90, 498, 658, 1025
BpmI	CTGGAG	1 cut(s)	375
BpuEI	CTTGAG	1 cut(s)	1064
BpuMI	CCSGG	1 cut(s)	498
BsaHI	GRCGYC	1 cut(s)	776
BsaI	GGTCTC	1 cut(s)	43
BsaJI	CCNNGG	4 cut(s)	39, 88, 111, 657
BsaWI	WCCGGW	1 cut(s)	191
BsaXI	ACNNNNNCTCC	6 cut(s)	5, 35, 173, 200, 203, 230
Bse1I	ACTGG	1 cut(s)	463
Bse3DI	GCAATG	3 cut(s)	436, 879, 1048
BseAI	TCCGGA	1 cut(s)	191
BseBI	CCWGG	3 cut(s)	90, 658, 1025
BseDI	CCNNGG	4 cut(s)	39, 88, 111, 657
BseGI	GGATG	6 cut(s)	17, 152, 254, 654, 724, 868
BseMI	GCAATG	3 cut(s)	436, 879, 1048
BseMII	CTCAG	5 cut(s)	120, 414, 456, 533, 770
BseNI	ACTGG	1 cut(s)	463
BseRI	GAGGAG	1 cut(s)	13
BseXI	GCAGC	3 cut(s)	178, 276, 1049
Bsh1236I	CGCG	2 cut(s)	144, 1076
Bsh1285I	CGRYCG	3 cut(s)	142, 164, 305
BshFI	GGCC	5 cut(s)	44, 66, 180, 606, 1008
BsiEI	CGRYCG	3 cut(s)	142, 164, 305
BsiHKCI	CYCGRG	1 cut(s)	46
BsiSI	CCGG	2 cut(s)	192, 498
BslFI	GGGAC	2 cut(s)	287, 480
BsmAI	GTCTC	3 cut(s)	43, 189, 522
BsmBI	CGTCTC	1 cut(s)	189
BsmFI	GGGAC	2 cut(s)	287, 480
BsmI	GAATGC	1 cut(s)	622
BsnI	GGCC	5 cut(s)	44, 66, 180, 606, 1008
Bso31I	GGTCTC	1 cut(s)	43
BsoBI	CYCGRG	1 cut(s)	46
Bsp13I	TCCGGA	1 cut(s)	191
Bsp143I	GATC	6 cut(s)	92, 302, 326, 470, 1027, 1039
Bsp19I	CCATGG	1 cut(s)	39
BspANI	GGCC	5 cut(s)	44, 66, 180, 606, 1008
BspCNI	CTCAG	5 cut(s)	119, 415, 455, 532, 769
BspEI	TCCGGA	1 cut(s)	191
BspFNI	CGCG	2 cut(s)	144, 1076
BspLI	GGNNCC	5 cut(s)	45, 82, 94, 130, 496
BspPI	GGATC	4 cut(s)	87, 100, 334, 1035
BspTI	CTTAAG	1 cut(s)	403
BspTNI	GGTCTC	1 cut(s)	43
BsrDI	GCAATG	3 cut(s)	436, 879, 1048
BsrI	ACTGG	1 cut(s)	463
BssECI	CCNNGG	4 cut(s)	39, 88, 111, 657
BssMI	GATC	6 cut(s)	92, 302, 326, 470, 1027, 1039
BssNI	GRCGYC	1 cut(s)	776
BssT1I	CCWWGG	1 cut(s)	39
Bst2UI	CCWGG	3 cut(s)	90, 658, 1025
Bst4CI	ACNGT	3 cut(s)	200, 311, 796
Bst6I	CTCTTC	1 cut(s)	343
BstACI	GRCGYC	1 cut(s)	776
BstAFI	CTTAAG	1 cut(s)	403
BstC8I	GCNNGC	2 cut(s)	380, 397
BstDEI	CTNAG	6 cut(s)	106, 423, 442, 519, 597, 756
BstDSI	CCRYGG	2 cut(s)	39, 111
BstF5I	GGATG	6 cut(s)	17, 152, 254, 654, 724, 868
BstFNI	CGCG	2 cut(s)	144, 1076
BstKTI	GATC	6 cut(s)	95, 305, 329, 473, 1030, 1042
BstMAI	GTCTC	3 cut(s)	43, 189, 522
BstMBI	GATC	6 cut(s)	92, 302, 326, 470, 1027, 1039
BstMCI	CGRYCG	3 cut(s)	142, 164, 305
BstMWI	GCNNNNNNNGC	7 cut(s)	34, 63, 72, 304, 384, 405, 1059
BstNI	CCWGG	3 cut(s)	90, 658, 1025
BstNSI	RCATGY	1 cut(s)	979
BstSCI	CCNGG	4 cut(s)	88, 496, 656, 1023
BstSFI	CTRYAG	1 cut(s)	466
BstUI	CGCG	2 cut(s)	144, 1076
BstV1I	GCAGC	3 cut(s)	178, 276, 1049
BstX2I	RGATCY	3 cut(s)	92, 326, 1027
BstYI	RGATCY	3 cut(s)	92, 326, 1027
BsuI	GTATCC	1 cut(s)	15
BsuRI	GGCC	5 cut(s)	44, 66, 180, 606, 1008
BtgI	CCRYGG	2 cut(s)	39, 111
BtsCI	GGATG	6 cut(s)	17, 152, 254, 654, 724, 868
BtsI	GCAGTG	1 cut(s)	293
BtsIMutI	CAGTG	1 cut(s)	293
Cac8I	GCNNGC	2 cut(s)	380, 397
Cfr13I	GGNCC	3 cut(s)	43, 64, 494
CspCI	CAANNNNNGTGG	2 cut(s)	472, 507
CviAII	CATG	5 cut(s)	40, 124, 619, 949, 976
DdeI	CTNAG	6 cut(s)	106, 423, 442, 519, 597, 756
DpnI	GATC	6 cut(s)	94, 304, 328, 472, 1029, 1041
DpnII	GATC	6 cut(s)	92, 302, 326, 470, 1027, 1039
EaeI	YGGCCR	1 cut(s)	604
Eam1104I	CTCTTC	1 cut(s)	343
EarI	CTCTTC	1 cut(s)	343
EciI	GGCGGA	2 cut(s)	73, 681
Eco130I	CCWWGG	1 cut(s)	39
Eco147I	AGGCCT	1 cut(s)	1008
Eco31I	GGTCTC	1 cut(s)	43
Eco47I	GGWCC	1 cut(s)	494
Eco57I	CTGAAG	2 cut(s)	582, 840
Eco88I	CYCGRG	1 cut(s)	46
EcoRI	GAATTC	1 cut(s)	146
EcoRII	CCWGG	3 cut(s)	88, 656, 1023
EcoT14I	CCWWGG	1 cut(s)	39
EcoT22I	ATGCAT	1 cut(s)	624
ErhI	CCWWGG	1 cut(s)	39
Esp3I	CGTCTC	1 cut(s)	189
FaeI	CATG	5 cut(s)	43, 127, 622, 952, 979
FalI	AAGNNNNNCTT	2 cut(s)	531, 563
FaqI	GGGAC	2 cut(s)	287, 480
FatI	CATG	5 cut(s)	39, 123, 618, 948, 975
FauI	CCCGC	2 cut(s)	80, 905
Fnu4HI	GCNGC	8 cut(s)	132, 135, 167, 205, 265, 366, 1060, 1063
FokI	GGATG	6 cut(s)	4, 139, 241, 661, 731, 855
Fsp4HI	GCNGC	8 cut(s)	132, 135, 167, 205, 265, 366, 1060, 1063
FspBI	CTAG	3 cut(s)	225, 639, 903
GluI	GCNGC	8 cut(s)	132, 135, 167, 205, 265, 366, 1060, 1063
GsuI	CTGGAG	1 cut(s)	375
HaeIII	GGCC	5 cut(s)	44, 66, 180, 606, 1008
HapII	CCGG	2 cut(s)	192, 498
Hin1I	GRCGYC	1 cut(s)	776
Hin1II	CATG	5 cut(s)	43, 127, 622, 952, 979
HincII	GTYRAC	2 cut(s)	250, 724
HindII	GTYRAC	2 cut(s)	250, 724
HinfI	GANTC	8 cut(s)	116, 209, 233, 438, 542, 653, 752, 824
HpaII	CCGG	2 cut(s)	192, 498
HphI	GGTGA	2 cut(s)	406, 593
Hpy166II	GTNNAC	4 cut(s)	115, 250, 701, 724
Hpy188I	TCNGA	5 cut(s)	161, 232, 302, 562, 873
Hpy188III	TCNNGA	4 cut(s)	192, 646, 818, 1043
Hpy8I	GTNNAC	4 cut(s)	115, 250, 701, 724
Hpy99I	CGWCG	2 cut(s)	364, 781
HpyAV	CCTTC	4 cut(s)	229, 578, 776, 1019
HpyCH4III	ACNGT	3 cut(s)	200, 311, 796
HpyCH4IV	ACGT	1 cut(s)	776
HpyCH4V	TGCA	7 cut(s)	622, 837, 863, 884, 994, 1053, 1065
HpyF10VI	GCNNNNNNNGC	7 cut(s)	34, 63, 72, 304, 384, 405, 1059
HpyF3I	CTNAG	6 cut(s)	106, 423, 442, 519, 597, 756
HpySE526I	ACGT	1 cut(s)	776
Hsp92I	GRCGYC	1 cut(s)	776
Hsp92II	CATG	5 cut(s)	43, 127, 622, 952, 979
Kpn2I	TCCGGA	1 cut(s)	191
Kzo9I	GATC	6 cut(s)	92, 302, 326, 470, 1027, 1039
LmnI	GCTCC	5 cut(s)	86, 128, 233, 303, 694
Lsp1109I	GCAGC	3 cut(s)	178, 276, 1049
MaeI	CTAG	3 cut(s)	225, 639, 903
MaeII	ACGT	1 cut(s)	776
MaeIII	GTNAC	2 cut(s)	445, 502
MalI	GATC	6 cut(s)	94, 304, 328, 472, 1029, 1041
MboI	GATC	6 cut(s)	92, 302, 326, 470, 1027, 1039
MboII	GAAGA	7 cut(s)	321, 330, 333, 342, 833, 909, 1022
MflI	RGATCY	3 cut(s)	92, 326, 1027
MluCI	AATT	2 cut(s)	146, 670
MlyI	GAGTC	3 cut(s)	110, 203, 242
MmeI	TCCRAC	4 cut(s)	184, 282, 357, 960
Mph1103I	ATGCAT	1 cut(s)	624
MroI	TCCGGA	1 cut(s)	191
MseI	TTAA	3 cut(s)	404, 512, 920
MslI	CAYNNNNRTG	1 cut(s)	612
MspA1I	CMGCKG	1 cut(s)	395
MspCI	CTTAAG	1 cut(s)	403
MspI	CCGG	2 cut(s)	192, 498
MspR9I	CCNGG	4 cut(s)	90, 498, 658, 1025
Mva1269I	GAATGC	1 cut(s)	622
MvaI	CCWGG	3 cut(s)	90, 658, 1025
MvnI	CGCG	2 cut(s)	144, 1076
MwoI	GCNNNNNNNGC	7 cut(s)	34, 63, 72, 304, 384, 405, 1059
NciI	CCSGG	1 cut(s)	498
NcoI	CCATGG	1 cut(s)	39
NdeII	GATC	6 cut(s)	92, 302, 326, 470, 1027, 1039
NlaIII	CATG	5 cut(s)	43, 127, 622, 952, 979
NlaIV	GGNNCC	5 cut(s)	45, 82, 94, 130, 496
NsiI	ATGCAT	1 cut(s)	624
NspI	RCATGY	1 cut(s)	979
OliI	CACNNNNGTG	1 cut(s)	612
PceI	AGGCCT	1 cut(s)	1008
PciI	ACATGT	1 cut(s)	975
PcsI	WCGNNNNNNNCGW	1 cut(s)	204
PctI	GAATGC	1 cut(s)	622
PfeI	GAWTC	5 cut(s)	438, 542, 653, 752, 824
PflFI	GACNNNGTC	1 cut(s)	358
PfoI	TCCNGGA	2 cut(s)	496, 1023
PkrI	GCNGC	8 cut(s)	133, 136, 168, 206, 266, 367, 1061, 1064
Ple19I	CGATCG	1 cut(s)	305
PleI	GAGTC	3 cut(s)	110, 203, 241
PpsI	GAGTC	3 cut(s)	110, 203, 241
PscI	ACATGT	1 cut(s)	975
Psp6I	CCWGG	3 cut(s)	88, 656, 1023
PspGI	CCWGG	3 cut(s)	88, 656, 1023
PspN4I	GGNNCC	5 cut(s)	45, 82, 94, 130, 496
PspPI	GGNCC	3 cut(s)	43, 64, 494
PsuI	RGATCY	3 cut(s)	92, 326, 1027
PsyI	GACNNNGTC	1 cut(s)	358
PvuI	CGATCG	1 cut(s)	305
PvuII	CAGCTG	1 cut(s)	395
RseI	CAYNNNNRTG	1 cut(s)	612
SaqAI	TTAA	3 cut(s)	404, 512, 920
SatI	GCNGC	8 cut(s)	132, 135, 167, 205, 265, 366, 1060, 1063
Sau3AI	GATC	6 cut(s)	92, 302, 326, 470, 1027, 1039
Sau96I	GGNCC	3 cut(s)	43, 64, 494
SchI	GAGTC	3 cut(s)	110, 203, 242
ScrFI	CCNGG	4 cut(s)	90, 498, 658, 1025
SfcI	CTRYAG	1 cut(s)	466
SinI	GGWCC	1 cut(s)	494
SmiMI	CAYNNNNRTG	1 cut(s)	612
SmlI	CTYRAG	2 cut(s)	403, 1043
SmoI	CTYRAG	2 cut(s)	403, 1043
Sse9I	AATT	2 cut(s)	146, 670
SseBI	AGGCCT	1 cut(s)	1008
SspMI	CTAG	3 cut(s)	225, 639, 903
StuI	AGGCCT	1 cut(s)	1008
StyD4I	CCNGG	4 cut(s)	88, 496, 656, 1023
StyI	CCWWGG	1 cut(s)	39
TaaI	ACNGT	3 cut(s)	200, 311, 796
TaiI	ACGT	1 cut(s)	779
TaqI	TCGA	2 cut(s)	353, 674
TasI	AATT	2 cut(s)	146, 670
TauI	GCSGC	5 cut(s)	134, 137, 207, 368, 1062
TfiI	GAWTC	5 cut(s)	438, 542, 653, 752, 824
Tru1I	TTAA	3 cut(s)	404, 512, 920
Tru9I	TTAA	3 cut(s)	404, 512, 920
TscAI	CASTG	1 cut(s)	300
TseI	GCWGC	3 cut(s)	166, 264, 1062
TspDTI	ATGAA	1 cut(s)	1022
TspGWI	ACGGA	1 cut(s)	833
TspRI	CASTG	1 cut(s)	300
Tth111I	GACNNNGTC	1 cut(s)	358
Vha464I	CTTAAG	1 cut(s)	403
VpaK11BI	GGWCC	1 cut(s)	494
XapI	RAATTY	1 cut(s)	146
XceI	RCATGY	1 cut(s)	979
XspI	CTAG	3 cut(s)	225, 639, 903
ZraI	GACGTC	1 cut(s)	777
Zsp2I	ATGCAT	1 cut(s)	624

Using CommOnly database (standard laboratory enzymes). Scanned on CDS sequence.