Basic Information

Type: gene

Biological Identity

Species

rosa_wichuraiana

Chromosome

Chr2

Orthogroup ID

OG0007545

Physical Location & Seq

Genomic Coordinates Forward (+)

13450476 .. 13453264

Gene Structure 2789 bp

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UTR

Exon/CDS

Intron

Transcript / Protein ID

Rw2G012860.1

Sequence Viewer

CDS Protein Promoter (2k)

Length: 1815 bp

ATGAGTTCTCCTTGCTGGAAAATTGATCAGCGACCAAAATTTTCTGCTACGCCTCTCAGCTTTCATAAGTCCATTTCGGAGAAAACTCACAGAGACAATAATCTTAGAGTTCCAAGGAGTTATGTGAAGAAATATGAGAAGAAATATGGAGAAGAGCTATCAAATATCGTACATCTTAAGCTTCCATGTGGTACAGAATGGGAAATAGAAGTCACAAGACATAATGGTGTCTTTTGGTTCGACAAGGGTTGGGAAACCTTTTGCAAGTTTTACTCCATCGAGTCTCGCTACTCGCTAGTTTTTCAATATGAAGGGAATTCTAGATTCAAAGTTAACATATTTGATAGAACAAACTCAGAGATTGAGTATCCTATCAAAACCCCGAAGAAGGAAGAAAATGATGTAGATGATCAATTTCCACCGGCTAGTCCTCTGCTGCCTAAAAAGAACAATATAGCAAGTTCAAATGGTGACAAAAAGGATCTTCCTGCGGAAGTAGATGAGGGAGACAAGTCTAGTAGTACTCAAAGATCACAAAAACCAACAACTGAGGTTCTTGGGAGGATGCACCCTTCGACTACAAGAGGAAGTGGAGCTGTACAGACAGTTAAAGAATTCAAGTACACTGAGCAGCCTTCTTGTTTTTTGATTCCTATGTGCCCCTCTTATATCGAAAAACATATGCTTTGGCTGCCAAGTGACTTTGCCATGCATTTGAAACATCTTATTACGAATTCTGGTAATCTTACCCTTCGTGTTTCGGAAGGGAGCACAACGAGAACTTGGACTGTTGGAATGAAATATGAAAAAGAAAGAGCCAAATTCCAATCTGGATGGAGTGAATTTGTGCAAGGCAATAAACTGAATCATGGTGATGTGTGTGACCTCATGTTAATTGACAAAAATGAACTTTTATTTGAAGTTAAGATTTTCCGAGCAAGAGAAGCTGTGGATTGCACCTTCTCACCGGATATTGATGAAGCTGATGCACATCACGAGCATGATGATGACAATGGTGATAATGATGCTGCTCATGACAATCGTTCATTCACAATTATCATACGACAGTCTTATACTACTCGTTTGCCAGTGCCTGCTAAGTTCCACAAAAATGTTACTAAGAAGCCTGATACCGCTAACCTGAGTCTTCCCGGTGGGAGGAGAACTTGGCCTATTTCATTGAAATATGAGGTTTCAACAGAAAGAGTCAGATTCCAGTCCGGTTGGAGCAAGTTTGTACGAGACAATCATTTGAAAGTTGGAGATGCGTGTGAATTTGTGTTGACTGATGAAACTAGACTTTCATTTGAAGTTCATATAGAAGCTGCTAAATGCACCTTGCCAGCACCACGACTTCACAAAAGAAAGAGATCAAGTTCAAGTACCAAAAGCAACATCGAAGATGTAGATGGTTCCACTAGTAATAAACAATTCTTGAAACGAATTCGAAGGAGCATGAGTCCAATAGGGAACAAGAAAAAAGATAATGCTCTTCAGAGAGCTAATGCTTTCAAATCTGTAAAACCTTCTTTCGTGGTTGCAATGCATCCCACTTATGTCTGTGGAGGTGTTTTGAATTTGCCATGTGGTTTTGCCAAGAAACATCTTACTGAGAGGCCTAATCACGTGAGCCTGCGGGGTTCAAATGGGAAAATTTGGTCTGTCAATTTCCATTTCAGCAACCAAGCTAGCTTCCGCAGTGGTTGGACTGACTTTGTGCGAGGCAATAATTTGGAAGAAGGTGATGCGTGTGCCTTTGAGTTGATTAAGAAGAGCAAATTTTTATTTGACGTTCACTTTTCCCGCACCACATAG

Functional Annotation

Gene Ontology

Molecular Function
Biological Process
Cellular Component

GO:0003674 GO:0003676 GO:0003677 GO:0005488 GO:0005575 GO:0005622 GO:0005623 GO:0005634 GO:0005654 GO:0006950 GO:0008150 GO:0009266 GO:0009409 GO:0009628 GO:0009909 GO:0010048 GO:0031974 GO:0031981 GO:0043226 GO:0043227 GO:0043229 GO:0043231 GO:0043233 GO:0043565 GO:0044422 GO:0044424 GO:0044428 GO:0044446 GO:0044464 GO:0048580 GO:0048831 GO:0050789 GO:0050793 GO:0050896 GO:0051239 GO:0065007 GO:0070013 GO:0097159 GO:1901363 GO:2000026 GO:2000241

KEGG Pathways

Metabolic & Signaling

No pathways identified.

Pfam Domains

Protein Families

Protein Analysis

604

Amino Acids

69.13

Weight (kDa)

9.17

Isoelectric Point (pI)

51.27

Instability Index

Protein Domains (Pfam)

Domain Name	Pfam ID	Position	E-value	Description
B3	PF02362	26 - 116	1.6e-13	B3 DNA binding domain
B3	PF02362	216 - 313	1.6e-14	B3 DNA binding domain
B3	PF02362	359 - 440	4e-13	B3 DNA binding domain
B3	PF02362	522 - 602	9.3e-14	B3 DNA binding domain

Hydrophobicity Profile (Kyte-Doolittle)

AI Structure Prediction Report

Calculating structure properties...

Genomic Context

Gene Family Tree

Species Filter

Style Settings

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Align Labels (Cladogram)

Image

Tree File

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Publication-ready

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Orthologous Genes (Group: OG0007545)

Species	Orthologous Gene IDs
rosa_chinensis	RchiOBHm_Chr2g0103451 RchiOBHm_Chr2g0103461 RchiOBHm_Chr2g0103471 RchiOBHm_Chr2g0103481
rosa_laevigata	RLG00000017237 RLG00000017238 RLG00000017239
rosa_multiflora	Rmu_co8300361.1_g000001 Rmu_co8395735.1_g000001 Rmu_sc0001852.1_g000027 Rmu_sc0011039.1_g000006
rosa_roxburghii	Rroxscaffold_2G00139600 Rroxscaffold_2G00139610 Rroxscaffold_2G00139620
rosa_rugosa	Rorug02G0115000 Rorug02G0115100
rosa_samantha	Rh2AG163400 Rh2AG163500 Rh2AG163600 Rh2BG170500 Rh2BG170600 Rh2BG170700 Rh2DG169000
rosa_wichuraiana	Rw2G012850 Rw2G012860

Restriction Enzyme Sites

1 / 10

Enzyme	Recognition Site	Cut Count	Positions (bp)
AciI	CCGC	5 cut(s)	491, 1134, 1636, 1696, 1804
AclWI	GGATC	1 cut(s)	489
AcuI	CTGAAG	1 cut(s)	1478
AcvI	CACGTG	1 cut(s)	1627
AfaI	GTAC	7 cut(s)	171, 193, 523, 600, 623, 1239, 1384
AfiI	CCNNNNNNNGG	2 cut(s)	388, 1158
AflII	CTTAAG	1 cut(s)	176
AhlI	ACTAGT	1 cut(s)	1418
Alw21I	GWGCWC	1 cut(s)	773
Alw26I	GTCTC	4 cut(s)	87, 288, 501, 1236
AlwI	GGATC	1 cut(s)	489
AlwNI	CAGNNNCTG	1 cut(s)	1094
AoxI	GGCC	2 cut(s)	1169, 1616
ApeKI	GCWGC	5 cut(s)	436, 631, 691, 1028, 1325
AsuC2I	CCSGG	1 cut(s)	1152
AsuHPI	GGTGA	5 cut(s)	482, 884, 957, 1028, 1754
AsuII	TTCGAA	1 cut(s)	1447
AsuNHI	GCTAGC	1 cut(s)	1688
BaeGI	GKGCMC	1 cut(s)	662
BauI	CACGAG	1 cut(s)	995
BbrPI	CACGTG	1 cut(s)	1627
BbsI	GAAGAC	1 cut(s)	1139
Bbv12I	GWGCWC	1 cut(s)	773
BbvI	GCAGC	5 cut(s)	423, 643, 678, 1015, 1312
BccI	CCATC	3 cut(s)	284, 828, 1403
BciVI	GTATCC	1 cut(s)	378
BclI	TGATCA	2 cut(s)	25, 409
BcnI	CCSGG	1 cut(s)	1152
BcoDI	GTCTC	4 cut(s)	87, 288, 501, 1236
BcuI	ACTAGT	1 cut(s)	1418
BfaI	CTAG	7 cut(s)	296, 321, 426, 516, 1296, 1419, 1689
BfrI	CTTAAG	1 cut(s)	176
BfuI	GTATCC	1 cut(s)	378
BisI	GCNGC	5 cut(s)	437, 632, 692, 1029, 1326
BlsI	GCNGC	5 cut(s)	438, 633, 693, 1030, 1327
BmcAI	AGTACT	1 cut(s)	523
Bme1390I	CCNGG	1 cut(s)	1152
BmiI	GGNNCC	1 cut(s)	1414
BmrFI	CCNGG	1 cut(s)	1152
BmsI	GCATC	6 cut(s)	555, 976, 1015, 1255, 1555, 1735
BmtI	GCTAGC	1 cut(s)	1692
BpiI	GAAGAC	1 cut(s)	1139
Bpu14I	TTCGAA	1 cut(s)	1447
BpuMI	CCSGG	1 cut(s)	1152
BsaAI	YACGTR	1 cut(s)	1627
BsaBI	GATNNNNATC	1 cut(s)	990
BsaJI	CCNNGG	1 cut(s)	113
BsaWI	WCCGGW	2 cut(s)	967, 1220
BsaXI	ACNNNNNCTCC	2 cut(s)	1444, 1474
Bsc4I	CCNNNNNNNGG	2 cut(s)	388, 1158
Bse118I	RCCGGY	1 cut(s)	421
Bse1I	ACTGG	2 cut(s)	1088, 1216
Bse3DI	GCAATG	1 cut(s)	1548
Bse8I	GATNNNNATC	1 cut(s)	990
BseDI	CCNNGG	1 cut(s)	113
BseGI	GGATG	3 cut(s)	570, 839, 1546
BseJI	GATNNNNATC	1 cut(s)	990
BseLI	CCNNNNNNNGG	2 cut(s)	388, 1158
BseMI	GCAATG	1 cut(s)	1548
BseMII	CTCAG	6 cut(s)	70, 369, 540, 618, 1133, 1602
BseNI	ACTGG	2 cut(s)	1088, 1216
BseRI	GAGGAG	1 cut(s)	1174
BseSI	GKGCMC	1 cut(s)	662
BseXI	GCAGC	5 cut(s)	423, 643, 678, 1015, 1312
BshFI	GGCC	2 cut(s)	1171, 1618
BsiHKAI	GWGCWC	1 cut(s)	773
BsiSI	CCGG	4 cut(s)	422, 968, 1152, 1221
BslI	CCNNNNNNNGG	2 cut(s)	388, 1158
BsmAI	GTCTC	4 cut(s)	87, 288, 501, 1236
BsnI	GGCC	2 cut(s)	1171, 1618
Bsp119I	TTCGAA	1 cut(s)	1447
Bsp1286I	GDGCHC	2 cut(s)	662, 773
Bsp1407I	TGTACA	1 cut(s)	598
Bsp143I	GATC	5 cut(s)	25, 409, 481, 530, 1370
BspACI	CCGC	5 cut(s)	491, 1134, 1636, 1696, 1804
BspANI	GGCC	2 cut(s)	1171, 1618
BspCNI	CTCAG	6 cut(s)	69, 368, 541, 619, 1134, 1603
BspHI	TCATGA	1 cut(s)	1033
BspLI	GGNNCC	1 cut(s)	1414
BspOI	GCTAGC	1 cut(s)	1692
BspPI	GGATC	1 cut(s)	489
BspQI	GCTCTTC	3 cut(s)	147, 1497, 1766
BspT104I	TTCGAA	1 cut(s)	1447
BspTI	CTTAAG	1 cut(s)	176
BsrDI	GCAATG	1 cut(s)	1548
BsrFI	RCCGGY	1 cut(s)	421
BsrGI	TGTACA	1 cut(s)	598
BsrI	ACTGG	2 cut(s)	1088, 1216
BssAI	RCCGGY	1 cut(s)	421
BssECI	CCNNGG	1 cut(s)	113
BssMI	GATC	5 cut(s)	25, 409, 481, 530, 1370
BssSI	CACGAG	1 cut(s)	995
BssT1I	CCWWGG	1 cut(s)	113
Bst2BI	CACGAG	1 cut(s)	995
Bst4CI	ACNGT	3 cut(s)	607, 790, 1068
Bst6I	CTCTTC	3 cut(s)	147, 1497, 1766
BstAFI	CTTAAG	1 cut(s)	176
BstAUI	TGTACA	1 cut(s)	598
BstBAI	YACGTR	1 cut(s)	1627
BstBI	TTCGAA	1 cut(s)	1447
BstC8I	GCNNGC	4 cut(s)	1095, 1344, 1634, 1690
BstDEI	CTNAG	9 cut(s)	56, 104, 355, 549, 627, 1098, 1119, 1142, 1611
BstF5I	GGATG	3 cut(s)	570, 839, 1546
BstKTI	GATC	5 cut(s)	28, 412, 484, 533, 1373
BstMAI	GTCTC	4 cut(s)	87, 288, 501, 1236
BstMBI	GATC	5 cut(s)	25, 409, 481, 530, 1370
BstMWI	GCNNNNNNNGC	2 cut(s)	691, 944
BstSCI	CCNGG	1 cut(s)	1150
BstSLI	GKGCMC	1 cut(s)	662
BstV1I	GCAGC	5 cut(s)	423, 643, 678, 1015, 1312
BstV2I	GAAGAC	1 cut(s)	1139
BstX2I	RGATCY	1 cut(s)	481
BstYI	RGATCY	1 cut(s)	481
BsuI	GTATCC	1 cut(s)	378
BsuRI	GGCC	2 cut(s)	1171, 1618
BtsCI	GGATG	3 cut(s)	570, 839, 1546
BtsI	GCAGTG	1 cut(s)	1705
BtsIMutI	CAGTG	3 cut(s)	624, 1095, 1705
Cac8I	GCNNGC	4 cut(s)	1095, 1344, 1634, 1690
CaiI	CAGNNNCTG	1 cut(s)	1094
CciI	TCATGA	1 cut(s)	1033
Cfr10I	RCCGGY	1 cut(s)	421
Csp6I	GTAC	7 cut(s)	170, 192, 522, 599, 622, 1238, 1383
CviAII	CATG	8 cut(s)	186, 709, 869, 889, 1001, 1034, 1456, 1584
CviQI	GTAC	7 cut(s)	170, 192, 522, 599, 622, 1238, 1383
DdeI	CTNAG	9 cut(s)	56, 104, 355, 549, 627, 1098, 1119, 1142, 1611
DpnI	GATC	5 cut(s)	27, 411, 483, 532, 1372
DpnII	GATC	5 cut(s)	25, 409, 481, 530, 1370
Eam1104I	CTCTTC	3 cut(s)	147, 1497, 1766
EarI	CTCTTC	3 cut(s)	147, 1497, 1766
Eco130I	CCWWGG	1 cut(s)	113
Eco147I	AGGCCT	1 cut(s)	1618
Eco57I	CTGAAG	1 cut(s)	1478
Eco72I	CACGTG	1 cut(s)	1627
EcoRI	GAATTC	4 cut(s)	316, 614, 733, 1443
EcoT14I	CCWWGG	1 cut(s)	113
EcoT22I	ATGCAT	2 cut(s)	714, 1548
ErhI	CCWWGG	1 cut(s)	113
FaeI	CATG	8 cut(s)	189, 712, 872, 892, 1004, 1037, 1459, 1587
FatI	CATG	8 cut(s)	185, 708, 868, 888, 1000, 1033, 1455, 1583
FauI	CCCGC	2 cut(s)	1629, 1811
FauNDI	CATATG	1 cut(s)	681
FbaI	TGATCA	2 cut(s)	25, 409
Fnu4HI	GCNGC	5 cut(s)	437, 632, 692, 1029, 1326
FokI	GGATG	3 cut(s)	577, 846, 1533
Fsp4HI	GCNGC	5 cut(s)	437, 632, 692, 1029, 1326
FspBI	CTAG	7 cut(s)	296, 321, 426, 516, 1296, 1419, 1689
GluI	GCNGC	5 cut(s)	437, 632, 692, 1029, 1326
HaeIII	GGCC	2 cut(s)	1171, 1618
HapII	CCGG	4 cut(s)	422, 968, 1152, 1221
Hin1II	CATG	8 cut(s)	189, 712, 872, 892, 1004, 1037, 1459, 1587
HincII	GTYRAC	2 cut(s)	334, 1284
HindII	GTYRAC	2 cut(s)	334, 1284
HindIII	AAGCTT	1 cut(s)	179
HinfI	GANTC	8 cut(s)	281, 324, 649, 865, 1144, 1206, 1212, 1459
HpaI	GTTAAC	1 cut(s)	334
HpaII	CCGG	4 cut(s)	422, 968, 1152, 1221
HphI	GGTGA	5 cut(s)	482, 884, 957, 1028, 1754
Hpy166II	GTNNAC	4 cut(s)	334, 624, 1284, 1795
Hpy188I	TCNGA	6 cut(s)	79, 358, 763, 935, 1211, 1497
Hpy188III	TCNNGA	5 cut(s)	321, 831, 995, 1034, 1435
Hpy8I	GTNNAC	4 cut(s)	334, 624, 1284, 1795
HpyCH4III	ACNGT	3 cut(s)	607, 790, 1068
HpyCH4IV	ACGT	2 cut(s)	1626, 1791
HpyCH4V	TGCA	9 cut(s)	264, 568, 712, 850, 957, 989, 1335, 1541, 1546
HpyF10VI	GCNNNNNNNGC	2 cut(s)	691, 944
HpyF3I	CTNAG	9 cut(s)	56, 104, 355, 549, 627, 1098, 1119, 1142, 1611
HpySE526I	ACGT	2 cut(s)	1626, 1791
Hsp92II	CATG	8 cut(s)	189, 712, 872, 892, 1004, 1037, 1459, 1587
Ksp22I	TGATCA	2 cut(s)	25, 409
KspAI	GTTAAC	1 cut(s)	334
Kzo9I	GATC	5 cut(s)	25, 409, 481, 530, 1370
LguI	GCTCTTC	3 cut(s)	147, 1497, 1766
LmnI	GCTCC	4 cut(s)	593, 768, 1227, 1452
Lsp1109I	GCAGC	5 cut(s)	423, 643, 678, 1015, 1312
LweI	GCATC	6 cut(s)	555, 976, 1015, 1255, 1555, 1735
MaeI	CTAG	7 cut(s)	296, 321, 426, 516, 1296, 1419, 1689
MaeII	ACGT	2 cut(s)	1626, 1791
MaeIII	GTNAC	5 cut(s)	211, 470, 698, 881, 1114
MalI	GATC	5 cut(s)	27, 411, 483, 532, 1372
MboI	GATC	5 cut(s)	25, 409, 481, 530, 1370
MflI	RGATCY	1 cut(s)	481
MhlI	GDGCHC	2 cut(s)	662, 773
MlyI	GAGTC	4 cut(s)	290, 1153, 1215, 1468
MmeI	TCCRAC	4 cut(s)	772, 1205, 1240, 1685
Mph1103I	ATGCAT	2 cut(s)	714, 1548
MseI	TTAA	6 cut(s)	177, 333, 609, 893, 924, 1767
MslI	CAYNNNNRTG	5 cut(s)	225, 873, 999, 1005, 1110
MspCI	CTTAAG	1 cut(s)	176
MspI	CCGG	4 cut(s)	422, 968, 1152, 1221
MspR9I	CCNGG	1 cut(s)	1152
MwoI	GCNNNNNNNGC	2 cut(s)	691, 944
NciI	CCSGG	1 cut(s)	1152
NdeI	CATATG	1 cut(s)	681
NdeII	GATC	5 cut(s)	25, 409, 481, 530, 1370
NheI	GCTAGC	1 cut(s)	1688
NlaIII	CATG	8 cut(s)	189, 712, 872, 892, 1004, 1037, 1459, 1587
NlaIV	GGNNCC	1 cut(s)	1414
NmuCI	GTSAC	4 cut(s)	211, 470, 698, 881
NsiI	ATGCAT	2 cut(s)	714, 1548
NspV	TTCGAA	1 cut(s)	1447
PagI	TCATGA	1 cut(s)	1033
PceI	AGGCCT	1 cut(s)	1618
PciSI	GCTCTTC	3 cut(s)	147, 1497, 1766
PfeI	GAWTC	4 cut(s)	324, 649, 865, 1212
PkrI	GCNGC	5 cut(s)	438, 633, 693, 1030, 1327
PleI	GAGTC	4 cut(s)	289, 1152, 1214, 1467
PmaCI	CACGTG	1 cut(s)	1627
PmlI	CACGTG	1 cut(s)	1627
PpsI	GAGTC	4 cut(s)	289, 1152, 1214, 1467
Ppu21I	YACGTR	1 cut(s)	1627
PspCI	CACGTG	1 cut(s)	1627
PspN4I	GGNNCC	1 cut(s)	1414
PstNI	CAGNNNCTG	1 cut(s)	1094
PsuI	RGATCY	1 cut(s)	481
RsaI	GTAC	7 cut(s)	171, 193, 523, 600, 623, 1239, 1384
RsaNI	GTAC	7 cut(s)	170, 192, 522, 599, 622, 1238, 1383
RseI	CAYNNNNRTG	5 cut(s)	225, 873, 999, 1005, 1110
SapI	GCTCTTC	3 cut(s)	147, 1497, 1766
SaqAI	TTAA	6 cut(s)	177, 333, 609, 893, 924, 1767
SatI	GCNGC	5 cut(s)	437, 632, 692, 1029, 1326
Sau3AI	GATC	5 cut(s)	25, 409, 481, 530, 1370
ScaI	AGTACT	1 cut(s)	523
SchI	GAGTC	4 cut(s)	290, 1153, 1215, 1468
ScrFI	CCNGG	1 cut(s)	1152
SduI	GDGCHC	2 cut(s)	662, 773
SfaNI	GCATC	6 cut(s)	555, 976, 1015, 1255, 1555, 1735
SfuI	TTCGAA	1 cut(s)	1447
SmiMI	CAYNNNNRTG	5 cut(s)	225, 873, 999, 1005, 1110
SmlI	CTYRAG	1 cut(s)	176
SmoI	CTYRAG	1 cut(s)	176
SpeI	ACTAGT	1 cut(s)	1418
SseBI	AGGCCT	1 cut(s)	1618
SsiI	CCGC	5 cut(s)	491, 1134, 1636, 1696, 1804
SspMI	CTAG	7 cut(s)	296, 321, 426, 516, 1296, 1419, 1689
StuI	AGGCCT	1 cut(s)	1618
StyD4I	CCNGG	1 cut(s)	1150
StyI	CCWWGG	1 cut(s)	113
TaaI	ACNGT	3 cut(s)	607, 790, 1068
TaiI	ACGT	2 cut(s)	1629, 1794
TaqI	TCGA	6 cut(s)	240, 279, 575, 672, 1398, 1447
TatI	WGTACW	3 cut(s)	521, 598, 621
TfiI	GAWTC	4 cut(s)	324, 649, 865, 1212
Tru1I	TTAA	6 cut(s)	177, 333, 609, 893, 924, 1767
Tru9I	TTAA	6 cut(s)	177, 333, 609, 893, 924, 1767
TscAI	CASTG	3 cut(s)	631, 1095, 1705
TseFI	GTSAC	4 cut(s)	211, 470, 698, 881
TseI	GCWGC	5 cut(s)	436, 631, 691, 1028, 1325
Tsp45I	GTSAC	4 cut(s)	211, 470, 698, 881
TspRI	CASTG	3 cut(s)	631, 1095, 1705
Vha464I	CTTAAG	1 cut(s)	176
XbaI	TCTAGA	1 cut(s)	320
XspI	CTAG	7 cut(s)	296, 321, 426, 516, 1296, 1419, 1689
ZrmI	AGTACT	1 cut(s)	523
Zsp2I	ATGCAT	2 cut(s)	714, 1548

Using CommOnly database (standard laboratory enzymes). Scanned on CDS sequence.