Basic Information

Type: gene

Biological Identity

Species

arabidopsis_thaliana

Chromosome

Orthogroup ID

OG0001983

Physical Location & Seq

Genomic Coordinates Forward (+)

7922276 .. 7925097

Gene Structure 2822 bp

Loading structure...

UTR

Exon/CDS

Intron

Transcript / Protein ID

AT1G22440.1

Sequence Viewer

CDS Protein Promoter (2k)

Length: 1161 bp

ATGGACAAGGCTTCAATCACTGAAGGAAAGCCGATTCGATGCAAAGCAGCAATCTTAAGAAAAGCAGGAGAACCATTGGTGATAGAAGAGATCCAAGTTGATCCACCTCAAGCTTACGAAGTCAGAATCAAAATTTTATGCACTTCTCTATGTCACACGGATGTTACTTTCTGGAAACTAGACAGTGGACCGCTTGCAAGATTTCCTAGGATTCTAGGTCATGAAGCAGTCGGTGTGGTCGAGAGTATTGGAGAAAAGGTAGACGGATTTAAACAAGGCGACGTCGTTTTGCCAGTGTTTCATCCTCAATGTGAAGAATGCAAAGAGTGCATATCTCCAAAGAGCAACTGGTGCACAAAATATACAAACGATTATTTATCAAACACAAGACGATATGGAATGACTTCAAGATTCAAGGACTCTCGTGGAGAAGATATTCACCATTTCATCTTTGTCTCAAGTTTTACTGAATATACCGTTGTAGATATCGCACATCTCGTCAAAATATCTCCTGAAATTCCCGTTGATATAGCGGCCTTGCTCAGCTGCAGTGTTGCCACAGGACTTGGAGCTGCGTGGAAAGTGGCTGACGTAGAGGAAGGTTCCACTGTCGTTATCTTTGGCCTTGGTGCTGTTGGACTAGCGGTTGCAGAAGGAGTCAGATTGCGTGGAGCCGCAAAAATCATCGGTGTGGATCTCAATCCCGCGAAATTTGAAATAGGTAAAAGATTTGGAATCACAGATTTTGTCAATCCTGCTTTGTGTGGGGAGAAGACAATTAGCGAGGTGATTAGAGAAATGACAGATGTAGGAGCTGACTACAGTTTTGAATGCATTGGTTTAGCTTCCTTGATGGAAGAGGCATTCAAGAGTACTCGCCCGGGATCGGGTAAAACGATAGTATTGGGGATGGAACAAAAGGCTTTGCCAATAAGCTTGGGTAGTTATGATCTTCTCAGAGGCAGAACTGTATGCGGCACTTTATTTGGCGGTTTGAAACCCAAACTTGATATTCCTATTCTCGTGGATCGTTATTTGAAAAAGGAGCTTAATCTAGAGGATCTGATTACACACGAATTAAGCTTTGAAGAAATTAACAAAGCTTTCCATTTATTGGCTGAGGGAAACTCTATCCGTTGCATTATATGGATGGATAAATAG

Functional Annotation

Gene Ontology

Molecular Function
Biological Process
Cellular Component

GO:0005575 GO:0005622 GO:0005623 GO:0005737 GO:0005829 GO:0044424 GO:0044444 GO:0044464

KEGG Pathways

Metabolic & Signaling

ko00010 ko00071 ko00190 ko00350 ko00625 ko00626 ko00680 ko00830 ko00980 ko00982 ko01100 ko01110 ko01120 ko01130 ko01200 ko01220 ko04260 ko04714 ko04932 ko05010 ko05012 ko05016 ko05204 map00010 map00071 map00190 map00350 map00625 map00626 map00680 map00830 map00980 map00982 map01100 map01110 map01120 map01130 map01200 map01220 map04260 map04714 map04932 map05010 map05012 map05016 map05204

Pfam Domains

Protein Families

ADH_N ADH_zinc_N

Protein Analysis

386

Amino Acids

42.38

Weight (kDa)

5.73

Isoelectric Point (pI)

31.74

Instability Index

Protein Domains (Pfam)

Domain Name	Pfam ID	Position	E-value	Description
ADH_N	PF08240	39 - 170	1.9e-19	Alcohol dehydrogenase GroES-like domain
ADH_zinc_N	PF00107	211 - 335	2.3e-22	Zinc-binding dehydrogenase

Hydrophobicity Profile (Kyte-Doolittle)

AI Structure Prediction Report

Calculating structure properties...

Genomic Context

Gene Family Tree

Species Filter

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Align Labels (Cladogram)

Image

Tree File

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Publication-ready

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Orthologous Genes (Group: OG0001983)

Species	Orthologous Gene IDs
arabidopsis_thaliana	AT1G22430 AT1G22430 AT1G22430 AT1G22430 AT1G22430 AT1G22440 AT4G22110 AT4G22110 AT4G22110
fragaria_vesca	FvH4_7g34200 FvH4_7g34200 FvH4_7g34200
malus_domestica	MD01G1238700.v1.1
prunus_persica	Prupe.2G328800_v2.0.a1
pyrus_communis	pycom01g24620
rosa_chinensis	RchiOBHm_Chr1g0315691 RchiOBHm_Chr1g0384561
rosa_laevigata	RLG00000026032 RLG00000030737 RLG00000030739
rosa_multiflora	Rmu_co8484511.1_g000001 Rmu_sc0004180.1_g000010 Rmu_sc0006005.1_g000011
rosa_roxburghii	Rroxscaffold_4G00276590 Rroxscaffold_4G00318470 Rroxscaffold_4G00331540
rosa_rugosa	Rorug01G0004700 Rorug01G0066000 Rorug01G0066000 Rorug01G0448900 Rorug01G0449000 Rorug04G0062000 Rorug04G0062100 Rorug04G0062200 RorugMtG0004000.1
rosa_samantha	Rh1AG015900 Rh1AG474000 Rh1BG008400 Rh1BG432100 Rh1CG013400 Rh1CG445300 Rh1DG009100 Rh1DG464700
rosa_wichuraiana	Rw1G000930 Rw1G039850

Restriction Enzyme Sites

1 / 10

Enzyme	Recognition Site	Cut Count	Positions (bp)
AatII	GACGTC	1 cut(s)	285
AccB7I	CCANNNNNTGG	1 cut(s)	1114
AccI	GTMKAC	1 cut(s)	261
AccII	CGCG	1 cut(s)	707
AciI	CCGC	7 cut(s)	191, 533, 644, 675, 705, 975, 990
AclWI	GGATC	6 cut(s)	85, 95, 702, 892, 1035, 1068
AcsI	RAATTY	3 cut(s)	132, 516, 710
AcuI	CTGAAG	1 cut(s)	42
AcyI	GRCGYC	1 cut(s)	282
AfaI	GTAC	1 cut(s)	874
AfiI	CCNNNNNNNGG	3 cut(s)	886, 887, 1114
AflII	CTTAAG	1 cut(s)	55
AgsI	TTSAA	9 cut(s)	15, 408, 415, 716, 830, 868, 997, 1039, 1088
AluBI	AGCT	9 cut(s)	113, 546, 572, 815, 845, 936, 1048, 1083, 1103
AluI	AGCT	9 cut(s)	113, 546, 572, 815, 845, 936, 1048, 1083, 1103
Alw21I	GWGCWC	1 cut(s)	356
Alw26I	GTCTC	1 cut(s)	460
Alw44I	GTGCAC	1 cut(s)	352
AlwI	GGATC	6 cut(s)	85, 95, 702, 892, 1035, 1068
Ama87I	CYCGRG	1 cut(s)	880
AoxI	GGCC	2 cut(s)	534, 622
ApaLI	GTGCAC	1 cut(s)	352
ApeKI	GCWGC	3 cut(s)	47, 546, 572
ApoI	RAATTY	3 cut(s)	132, 516, 710
ArsI	GACNNNNNNTTYG	2 cut(s)	809, 841
Asp700I	GAANNNNTTC	2 cut(s)	403, 435
AspA2I	CCTAGG	1 cut(s)	206
AspS9I	GGNCC	1 cut(s)	188
AsuC2I	CCSGG	2 cut(s)	881, 882
AsuHPI	GGTGA	3 cut(s)	91, 431, 799
AvaI	CYCGRG	1 cut(s)	880
AvaII	GGWCC	1 cut(s)	188
AvrII	CCTAGG	1 cut(s)	206
BaeGI	GKGCMC	1 cut(s)	356
BauI	CACGAG	2 cut(s)	423, 1022
BbsI	GAAGAC	1 cut(s)	779
Bbv12I	GWGCWC	1 cut(s)	356
BbvCI	CCTCAGC	1 cut(s)	1119
BbvI	GCAGC	3 cut(s)	59, 533, 559
BccI	CCATC	3 cut(s)	847, 904, 1144
BcnI	CCSGG	2 cut(s)	881, 882
BcoDI	GTCTC	1 cut(s)	460
BfaI	CTAG	5 cut(s)	179, 207, 215, 641, 1055
BfmI	CTRYAG	2 cut(s)	547, 820
BfrI	CTTAAG	1 cut(s)	55
BisI	GCNGC	6 cut(s)	48, 534, 547, 573, 675, 976
BlnI	CCTAGG	1 cut(s)	206
BlpI	GCTNAGC	1 cut(s)	542
BlsI	GCNGC	6 cut(s)	49, 535, 548, 574, 676, 977
BmcAI	AGTACT	1 cut(s)	874
Bme1390I	CCNGG	2 cut(s)	881, 882
Bme18I	GGWCC	1 cut(s)	188
BmeT110I	CYCGRG	1 cut(s)	880
BmgT120I	GGNCC	1 cut(s)	188
BmiI	GGNNCC	2 cut(s)	604, 673
BmrFI	CCNGG	2 cut(s)	881, 882
BmsI	GCATC	1 cut(s)	29
BpiI	GAAGAC	1 cut(s)	779
BplI	GAGNNNNNCTC	2 cut(s)	1112, 1144
Bpu10I	CCTNAGC	1 cut(s)	1119
Bpu1102I	GCTNAGC	1 cut(s)	542
BpuEI	CTTGAG	2 cut(s)	93, 442
BpuMI	CCSGG	2 cut(s)	881, 882
BsaBI	GATNNNNATC	1 cut(s)	699
BsaHI	GRCGYC	1 cut(s)	282
BsaJI	CCNNGG	3 cut(s)	206, 625, 880
BsaXI	ACNNNNNCTCC	2 cut(s)	243, 273
Bsc4I	CCNNNNNNNGG	3 cut(s)	886, 887, 1114
Bse1I	ACTGG	2 cut(s)	293, 353
Bse8I	GATNNNNATC	1 cut(s)	699
BseDI	CCNNGG	3 cut(s)	206, 625, 880
BseGI	GGATG	4 cut(s)	166, 301, 915, 1155
BseJI	GATNNNNATC	1 cut(s)	699
BseLI	CCNNNNNNNGG	3 cut(s)	886, 887, 1114
BseMII	CTCAG	3 cut(s)	556, 970, 1110
BseNI	ACTGG	2 cut(s)	293, 353
BseSI	GKGCMC	1 cut(s)	356
BseXI	GCAGC	3 cut(s)	59, 533, 559
Bsh1236I	CGCG	1 cut(s)	707
BshFI	GGCC	2 cut(s)	536, 624
BsiHKAI	GWGCWC	1 cut(s)	356
BsiHKCI	CYCGRG	1 cut(s)	880
BsiSI	CCGG	1 cut(s)	881
BslI	CCNNNNNNNGG	3 cut(s)	886, 887, 1114
BsmAI	GTCTC	1 cut(s)	460
BsmI	GAATGC	3 cut(s)	323, 836, 863
BsnI	GGCC	2 cut(s)	536, 624
BsoBI	CYCGRG	1 cut(s)	880
Bsp1286I	GDGCHC	1 cut(s)	356
Bsp143I	GATC	7 cut(s)	90, 100, 694, 884, 949, 1027, 1060
Bsp1720I	GCTNAGC	1 cut(s)	542
BspACI	CCGC	7 cut(s)	191, 533, 644, 675, 705, 975, 990
BspANI	GGCC	2 cut(s)	536, 624
BspCNI	CTCAG	3 cut(s)	555, 969, 1111
BspFNI	CGCG	1 cut(s)	707
BspHI	TCATGA	1 cut(s)	220
BspLI	GGNNCC	2 cut(s)	604, 673
BspMAI	CTGCAG	1 cut(s)	551
BspPI	GGATC	6 cut(s)	85, 95, 702, 892, 1035, 1068
BspTI	CTTAAG	1 cut(s)	55
BsrI	ACTGG	2 cut(s)	293, 353
BssECI	CCNNGG	3 cut(s)	206, 625, 880
BssMI	GATC	7 cut(s)	90, 100, 694, 884, 949, 1027, 1060
BssNI	GRCGYC	1 cut(s)	282
BssSI	CACGAG	2 cut(s)	423, 1022
BssT1I	CCWWGG	2 cut(s)	206, 625
Bst2BI	CACGAG	2 cut(s)	423, 1022
Bst4CI	ACNGT	5 cut(s)	185, 478, 610, 824, 970
Bst6I	CTCTTC	2 cut(s)	81, 852
BstACI	GRCGYC	1 cut(s)	282
BstAFI	CTTAAG	1 cut(s)	55
BstAPI	GCANNNNNTGC	2 cut(s)	327, 351
BstC8I	GCNNGC	1 cut(s)	195
BstDEI	CTNAG	3 cut(s)	542, 956, 1119
BstF5I	GGATG	4 cut(s)	166, 301, 915, 1155
BstFNI	CGCG	1 cut(s)	707
BstKTI	GATC	7 cut(s)	93, 103, 697, 887, 952, 1030, 1063
BstMAI	GTCTC	1 cut(s)	460
BstMBI	GATC	7 cut(s)	90, 100, 694, 884, 949, 1027, 1060
BstMWI	GCNNNNNNNGC	2 cut(s)	327, 351
BstSCI	CCNGG	2 cut(s)	879, 880
BstSFI	CTRYAG	2 cut(s)	547, 820
BstSLI	GKGCMC	1 cut(s)	356
BstUI	CGCG	1 cut(s)	707
BstV1I	GCAGC	3 cut(s)	59, 533, 559
BstV2I	GAAGAC	1 cut(s)	779
BstX2I	RGATCY	3 cut(s)	90, 694, 1060
BstYI	RGATCY	3 cut(s)	90, 694, 1060
BsuRI	GGCC	2 cut(s)	536, 624
BtsCI	GGATG	4 cut(s)	166, 301, 915, 1155
BtsI	GCAGTG	1 cut(s)	556
BtsIMutI	CAGTG	5 cut(s)	18, 190, 300, 556, 606
Cac8I	GCNNGC	1 cut(s)	195
CciI	TCATGA	1 cut(s)	220
Cfr13I	GGNCC	1 cut(s)	188
Cfr9I	CCCGGG	1 cut(s)	880
Csp6I	GTAC	1 cut(s)	873
CspCI	CAANNNNNGTGG	2 cut(s)	547, 582
CviAII	CATG	1 cut(s)	221
CviQI	GTAC	1 cut(s)	873
DdeI	CTNAG	3 cut(s)	542, 956, 1119
DpnI	GATC	7 cut(s)	92, 102, 696, 886, 951, 1029, 1062
DpnII	GATC	7 cut(s)	90, 100, 694, 884, 949, 1027, 1060
DraI	TTTAAA	1 cut(s)	271
Eam1104I	CTCTTC	2 cut(s)	81, 852
EarI	CTCTTC	2 cut(s)	81, 852
Eco130I	CCWWGG	2 cut(s)	206, 625
Eco32I	GATATC	1 cut(s)	487
Eco47I	GGWCC	1 cut(s)	188
Eco57I	CTGAAG	1 cut(s)	42
Eco88I	CYCGRG	1 cut(s)	880
EcoRV	GATATC	1 cut(s)	487
EcoT14I	CCWWGG	2 cut(s)	206, 625
EcoT22I	ATGCAT	1 cut(s)	836
ErhI	CCWWGG	2 cut(s)	206, 625
FaeI	CATG	1 cut(s)	224
FatI	CATG	1 cut(s)	220
FauI	CCCGC	1 cut(s)	712
FblI	GTMKAC	1 cut(s)	261
Fnu4HI	GCNGC	6 cut(s)	48, 534, 547, 573, 675, 976
FokI	GGATG	3 cut(s)	173, 288, 922
Fsp4HI	GCNGC	6 cut(s)	48, 534, 547, 573, 675, 976
FspBI	CTAG	5 cut(s)	179, 207, 215, 641, 1055
GluI	GCNGC	6 cut(s)	48, 534, 547, 573, 675, 976
HaeIII	GGCC	2 cut(s)	536, 624
HapII	CCGG	1 cut(s)	881
Hin1I	GRCGYC	1 cut(s)	282
Hin1II	CATG	1 cut(s)	224
HindIII	AAGCTT	4 cut(s)	111, 934, 1081, 1101
HinfI	GANTC	7 cut(s)	34, 126, 211, 411, 419, 657, 735
HpaII	CCGG	1 cut(s)	881
HphI	GGTGA	3 cut(s)	91, 431, 799
Hpy166II	GTNNAC	3 cut(s)	188, 262, 354
Hpy188I	TCNGA	4 cut(s)	125, 662, 959, 1065
Hpy188III	TCNNGA	7 cut(s)	172, 221, 241, 408, 512, 868, 1055
Hpy8I	GTNNAC	3 cut(s)	188, 262, 354
Hpy99I	CGWCG	2 cut(s)	284, 287
HpyAV	CCTTC	3 cut(s)	17, 593, 647
HpyCH4III	ACNGT	5 cut(s)	185, 478, 610, 824, 970
HpyCH4IV	ACGT	2 cut(s)	282, 591
HpyF10VI	GCNNNNNNNGC	2 cut(s)	327, 351
HpyF3I	CTNAG	3 cut(s)	542, 956, 1119
HpySE526I	ACGT	2 cut(s)	282, 591
Hsp92I	GRCGYC	1 cut(s)	282
Hsp92II	CATG	1 cut(s)	224
Kzo9I	GATC	7 cut(s)	90, 100, 694, 884, 949, 1027, 1060
LmnI	GCTCC	4 cut(s)	569, 671, 812, 1045
LpnPI	CCDG	8 cut(s)	51, 157, 306, 334, 525, 546, 768, 894
Lsp1109I	GCAGC	3 cut(s)	59, 533, 559
LweI	GCATC	1 cut(s)	29
MaeI	CTAG	5 cut(s)	179, 207, 215, 641, 1055
MaeII	ACGT	2 cut(s)	282, 591
MaeIII	GTNAC	2 cut(s)	152, 163
MalI	GATC	7 cut(s)	92, 102, 696, 886, 951, 1029, 1062
MboI	GATC	7 cut(s)	90, 100, 694, 884, 949, 1027, 1060
MboII	GAAGA	7 cut(s)	98, 326, 443, 784, 869, 944, 1100
MflI	RGATCY	3 cut(s)	90, 694, 1060
MhlI	GDGCHC	1 cut(s)	356
MluCI	AATT	6 cut(s)	132, 516, 710, 777, 1076, 1092
MlyI	GAGTC	2 cut(s)	413, 666
MmeI	TCCRAC	1 cut(s)	616
MnlI	CCTC	8 cut(s)	117, 315, 589, 778, 853, 953, 1051, 1114
Mph1103I	ATGCAT	1 cut(s)	836
MroXI	GAANNNNTTC	2 cut(s)	403, 435
MseI	TTAA	5 cut(s)	56, 270, 1050, 1079, 1095
MslI	CAYNNNNRTG	2 cut(s)	159, 689
MspA1I	CMGCKG	1 cut(s)	546
MspCI	CTTAAG	1 cut(s)	55
MspI	CCGG	1 cut(s)	881
MspR9I	CCNGG	2 cut(s)	881, 882
Mva1269I	GAATGC	3 cut(s)	323, 836, 863
MvnI	CGCG	1 cut(s)	707
MwoI	GCNNNNNNNGC	2 cut(s)	327, 351
NciI	CCSGG	2 cut(s)	881, 882
NdeII	GATC	7 cut(s)	90, 100, 694, 884, 949, 1027, 1060
NlaIII	CATG	1 cut(s)	224
NlaIV	GGNNCC	2 cut(s)	604, 673
NmuCI	GTSAC	1 cut(s)	152
NsiI	ATGCAT	1 cut(s)	836
PagI	TCATGA	1 cut(s)	220
PcsI	WCGNNNNNNNCGW	3 cut(s)	237, 495, 893
PctI	GAATGC	3 cut(s)	323, 836, 863
PdmI	GAANNNNTTC	2 cut(s)	403, 435
PfeI	GAWTC	5 cut(s)	34, 126, 211, 411, 735
PflMI	CCANNNNNTGG	1 cut(s)	1114
PkrI	GCNGC	6 cut(s)	49, 535, 548, 574, 676, 977
PleI	GAGTC	2 cut(s)	413, 665
PpsI	GAGTC	2 cut(s)	413, 665
PspN4I	GGNNCC	2 cut(s)	604, 673
PspPI	GGNCC	1 cut(s)	188
PstI	CTGCAG	1 cut(s)	551
PsuI	RGATCY	3 cut(s)	90, 694, 1060
PvuII	CAGCTG	1 cut(s)	546
RsaI	GTAC	1 cut(s)	874
RsaNI	GTAC	1 cut(s)	873
RseI	CAYNNNNRTG	2 cut(s)	159, 689
SaqAI	TTAA	5 cut(s)	56, 270, 1050, 1079, 1095
SatI	GCNGC	6 cut(s)	48, 534, 547, 573, 675, 976
Sau3AI	GATC	7 cut(s)	90, 100, 694, 884, 949, 1027, 1060
Sau96I	GGNCC	1 cut(s)	188
ScaI	AGTACT	1 cut(s)	874
SchI	GAGTC	2 cut(s)	413, 666
ScrFI	CCNGG	2 cut(s)	881, 882
SduI	GDGCHC	1 cut(s)	356
SfaNI	GCATC	1 cut(s)	29
SfcI	CTRYAG	2 cut(s)	547, 820
SinI	GGWCC	1 cut(s)	188
SmaI	CCCGGG	1 cut(s)	882
SmiMI	CAYNNNNRTG	2 cut(s)	159, 689
SmlI	CTYRAG	3 cut(s)	55, 108, 457
SmoI	CTYRAG	3 cut(s)	55, 108, 457
Sse9I	AATT	6 cut(s)	132, 516, 710, 777, 1076, 1092
SsiI	CCGC	7 cut(s)	191, 533, 644, 675, 705, 975, 990
SspMI	CTAG	5 cut(s)	179, 207, 215, 641, 1055
StyD4I	CCNGG	2 cut(s)	879, 880
StyI	CCWWGG	2 cut(s)	206, 625
TaaI	ACNGT	5 cut(s)	185, 478, 610, 824, 970
TaiI	ACGT	2 cut(s)	285, 594
TaqI	TCGA	2 cut(s)	37, 240
TasI	AATT	6 cut(s)	132, 516, 710, 777, 1076, 1092
TatI	WGTACW	1 cut(s)	872
TauI	GCSGC	3 cut(s)	536, 677, 978
TfiI	GAWTC	5 cut(s)	34, 126, 211, 411, 735
Tru1I	TTAA	5 cut(s)	56, 270, 1050, 1079, 1095
Tru9I	TTAA	5 cut(s)	56, 270, 1050, 1079, 1095
TscAI	CASTG	5 cut(s)	25, 190, 300, 556, 613
TseFI	GTSAC	1 cut(s)	152
TseI	GCWGC	3 cut(s)	47, 546, 572
Tsp45I	GTSAC	1 cut(s)	152
TspDTI	ATGAA	3 cut(s)	237, 290, 436
TspGWI	ACGGA	3 cut(s)	173, 279, 1124
TspMI	CCCGGG	1 cut(s)	880
TspRI	CASTG	5 cut(s)	25, 190, 300, 556, 613
Van91I	CCANNNNNTGG	1 cut(s)	1114
Vha464I	CTTAAG	1 cut(s)	55
VneI	GTGCAC	1 cut(s)	352
VpaK11BI	GGWCC	1 cut(s)	188
XapI	RAATTY	3 cut(s)	132, 516, 710
XbaI	TCTAGA	1 cut(s)	1054
XcmI	CCANNNNNNNNNTGG	1 cut(s)	345
XmaI	CCCGGG	1 cut(s)	880
XmaJI	CCTAGG	1 cut(s)	206
XmiI	GTMKAC	1 cut(s)	261
XmnI	GAANNNNTTC	2 cut(s)	403, 435
XspI	CTAG	5 cut(s)	179, 207, 215, 641, 1055
ZraI	GACGTC	1 cut(s)	283
ZrmI	AGTACT	1 cut(s)	874
Zsp2I	ATGCAT	1 cut(s)	836

Using CommOnly database (standard laboratory enzymes). Scanned on CDS sequence.