Basic Information

Type: gene

Biological Identity

Species

rosa_samantha

Chromosome

Chr4A

Orthogroup ID

OG0000642

Physical Location & Seq

Genomic Coordinates Forward (+)

63906145 .. 63909229

Gene Structure 3085 bp

Loading structure...

UTR

Exon/CDS

Intron

Transcript / Protein ID

Rh4AG327700.1

Sequence Viewer

CDS Protein Promoter (2k)

Length: 1473 bp

ATGGAAGCGAGCTTGATAACATTGCCAAATGATACTGAAGACGAAAGAAAAAGTAGAAGTGATAGTTCAAGTACTCATCATCTAACATGGGTTTGTTTCTTTGAAGAAGTGAAGAGACAGTGTTACATAGCAGGACCTATGGTTGCTGTGATCCTGTCCCAGAACTTGTTGCGTATTGTTTCGATGATGATGGTTGGTCACTTGGGTGAGCTTGCTCTCTCTAGCAGCTCTATTGCCATCTCCCTCACTGGAGTCACCGGCTTTAGTCTTTTTATAGGAATGGCCAGTGGACTGGAAACATTATGTGGCCAAGCATATGGAGCTAAGCAATATCAGAAACTTGGACTTCAAACTTACACTGCCATATTTTCTATCAACTTAGTTGGTCTACCATTATCTTTGATATGGATATACATGGAGGACTTGCTGATTTTGATTGGTCAAGATCCTTCAATTTCTCGTGAAGCTGGTAAATTCACAATTTGGCTTATTCCAGCTCTATTTGCTTATGCAACTCTTCAAGCATTGATTAGATACTTCCAGACACAAAGTTTAATAGTTCCTATGCTTCTAAGCTCTTGCGCGACCCTTTTGTTCCATATTCCTCTTTGTTGGGTTCTAGTATTTAAGTCTGGACTGGATAATGTTGGAGGGGCATTAGCAATCAGTATTTCATATTGGTTGAATGTTATTTTACTCATATTCTACATGAAGTTTTCTTCTGCCTGTGCAAAAACTAGAGTTCCAATTTCTATGGAGGTATTCTATGGAATCAAAGAGTTCTTCAGCTTTGCTATCCCTTCTGCAATTATGATATGCCTTGAGTGGTGGTCATTTGAGATGCTTATCTTGCTGTCCGGAATTCTACAAAATCCGGCTCTTGAAACTTCAGTTCTGTCTGTATGTCTCCAGACCATTGTGACAGTCTATTACATACCATATGGGTTTGCTGCTGCAGCAAGTATTAGAGTTTCCAACGAACTAGGAGCTGGTAACCCAGAAGGTGCTCGTATAGCAACATGTGCTGCGATGTTTCTTGCAATCACAGAGACAGCTATAGTAACCACAACTCTCTTTGCTTGCCGAAATGTATTTGGTTATGTTTTTAGCAATGAGCAAGAAGTTGTTGATTATGTCACAACCATGGCTCCTCTAGTTTGCTTGTCTGTTTTACTAGACAGCTTGCAAGCCGTCCTTTCAGGTATTGCTAGAGGGTGTGGGTGGCAGCACATAGTGGCTTACATAAACCTTGCAGCATTTTATATTTTTGGAATTCCAATTGCTGCCTTACTGGCTTTCTGGATACAACTAGGAGGAAGGGGCCTTTGGATTGGAATCCAAGTTGGTGCTTTTGTGCAAACAATTCTGCTTTCTTTTGTCACGACTTGTACAAATTGGGATAAACAGGCACATAAGGCGAGGGCTAGGATATATGAGGGAAGCTCTCTCGAAGCTAATGGATTGCTGCACTAA

Functional Annotation

Gene Ontology

Molecular Function
Biological Process
Cellular Component

GO:0003674 GO:0005215 GO:0005575 GO:0005622 GO:0005623 GO:0005737 GO:0006810 GO:0006855 GO:0008150 GO:0009507 GO:0009536 GO:0015238 GO:0015893 GO:0022857 GO:0042221 GO:0042493 GO:0043226 GO:0043227 GO:0043229 GO:0043231 GO:0044424 GO:0044444 GO:0044464 GO:0050896 GO:0051179 GO:0051234 GO:0055085

KEGG Pathways

Metabolic & Signaling

No pathways identified.

Pfam Domains

Protein Families

MatE

Protein Analysis

490

Amino Acids

53.71

Weight (kDa)

5.83

Isoelectric Point (pI)

34.58

Instability Index

Protein Domains (Pfam)

Domain Name	Pfam ID	Position	E-value	Description
MatE	PF01554	46 - 206	1.8e-30	MatE
MatE	PF01554	267 - 429	2.1e-26	MatE

Hydrophobicity Profile (Kyte-Doolittle)

AI Structure Prediction Report

Calculating structure properties...

Genomic Context

Gene Family Tree

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Align Labels (Cladogram)

Image

Tree File

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Publication-ready

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Orthologous Genes (Group: OG0000642)

Species	Orthologous Gene IDs
arabidopsis_thaliana	AT1G15150 AT1G15150 AT1G15160 AT1G15170 AT1G15170 AT1G15180 AT1G15180
fragaria_vesca	FvH4_4g26360 FvH4_4g26361 FvH4_4g26362 FvH4_4g26362 FvH4_4g26362 FvH4_4g26362 FvH4_4g26362
malus_domestica	MD13G1031800.v1.1 MD13G1031900.v1.1 MD13G1032000.v1.1 MD16G1033900.v1.1 MD16G1034100.v1.1 MD16G1034200.v1.1 MD16G1034300.v1.1
prunus_persica	Prupe.1G320300_v2.0.a1 Prupe.1G320400_v2.0.a1 Prupe.1G320400_v2.0.a1
pyrus_communis	pycom13g02770 pycom16g02870 pycom16g02880 pycom16g02900
rosa_chinensis	RchiOBHm_Chr4g0433941 RchiOBHm_Chr4g0433951 RchiOBHm_Chr4g0433961 RchiOBHm_Chr4g0433971 RchiOBHm_Chr4g0433981 RchiOBHm_Chr4g0433991
rosa_laevigata	RLG00000006710 RLG00000006711 RLG00000006712 RLG00000006713 RLG00000006714 RLG00000006715
rosa_multiflora	Rmu_co8148826.1_g000001 Rmu_co8171710.1_g000001 Rmu_sc0000352.1_g000016 Rmu_sc0000352.1_g000017 Rmu_sc0000352.1_g000020 Rmu_sc0004983.1_g000006 Rmu_sc0004983.1_g000007 Rmu_sc0004983.1_g000008 Rmu_sc0023229.1_g000002
rosa_roxburghii	Rroxscaffold_5G00375360 Rroxscaffold_5G00375370 Rroxscaffold_5G00375380 Rroxscaffold_5G00375390 Rroxscaffold_5G00375400
rosa_rugosa	Rorug04G0272400
rosa_samantha	Rh4AG327500 Rh4AG327600 Rh4AG327700 Rh4AG327800 Rh4AG327900 Rh4AG328000 Rh4BG334800 Rh4BG334900 Rh4BG335300 Rh4BG335400 Rh4CG350600 Rh4CG350800 Rh4CG350900 Rh4DG331700 Rh4DG331800 Rh4DG331900
rosa_wichuraiana	Rw4G028390 Rw4G028400 Rw4G028410 Rw4G028420 Rw4G028430 Rw4G028440

Restriction Enzyme Sites

1 / 10

Enzyme	Recognition Site	Cut Count	Positions (bp)
AccI	GTMKAC	1 cut(s)	388
AccII	CGCG	1 cut(s)	584
AccIII	TCCGGA	1 cut(s)	857
AclWI	GGATC	2 cut(s)	145, 440
AcoI	YGGCCR	2 cut(s)	282, 307
AcsI	RAATTY	3 cut(s)	473, 861, 1272
AcuI	CTGAAG	3 cut(s)	57, 769, 873
AdeI	CACNNNGTG	1 cut(s)	1234
AfaI	GTAC	2 cut(s)	73, 1390
AflIII	ACRYGT	1 cut(s)	1019
AgsI	TTSAA	7 cut(s)	69, 104, 350, 453, 521, 685, 884
AjuI	GAANNNNNNNTTGG	2 cut(s)	1309, 1341
AleI	CACNNNNGTG	1 cut(s)	204
Alw21I	GWGCWC	1 cut(s)	1009
Alw26I	GTCTC	3 cut(s)	109, 911, 1043
AlwI	GGATC	2 cut(s)	145, 440
Aor13HI	TCCGGA	1 cut(s)	857
AoxI	GGCC	3 cut(s)	282, 307, 1321
ApeKI	GCWGC	9 cut(s)	225, 950, 953, 956, 1025, 1225, 1253, 1283, 1465
ApoI	RAATTY	3 cut(s)	473, 861, 1272
AspLEI	GCGC	1 cut(s)	584
AspS9I	GGNCC	2 cut(s)	134, 1321
AsuHPI	GGTGA	2 cut(s)	218, 247
AvaII	GGWCC	1 cut(s)	134
BalI	TGGCCA	2 cut(s)	284, 309
BauI	CACGAG	1 cut(s)	459
BbsI	GAAGAC	1 cut(s)	45
Bbv12I	GWGCWC	1 cut(s)	1009
BbvI	GCAGC	9 cut(s)	237, 937, 940, 968, 1012, 1237, 1265, 1270, 1452
BccI	CCATC	2 cut(s)	184, 245
BceAI	ACGGC	1 cut(s)	1175
BciVI	GTATCC	1 cut(s)	1296
BcoDI	GTCTC	3 cut(s)	109, 911, 1043
BfaI	CTAG	9 cut(s)	222, 620, 738, 983, 1154, 1175, 1209, 1310, 1425
BfmI	CTRYAG	2 cut(s)	954, 1056
BfuI	GTATCC	1 cut(s)	1296
BglI	GCCNNNNNGGC	1 cut(s)	1292
BisI	GCNGC	9 cut(s)	226, 951, 954, 957, 1026, 1226, 1254, 1284, 1466
BlpI	GCTNAGC	1 cut(s)	324
BlsI	GCNGC	9 cut(s)	227, 952, 955, 958, 1027, 1227, 1255, 1285, 1467
BmcAI	AGTACT	1 cut(s)	73
Bme18I	GGWCC	1 cut(s)	134
BmgT120I	GGNCC	2 cut(s)	134, 1321
BmiI	GGNNCC	2 cut(s)	1149, 1322
BmsI	GCATC	1 cut(s)	831
BpiI	GAAGAC	1 cut(s)	45
BplI	GAGNNNNNCTC	2 cut(s)	1427, 1459
BpmI	CTGGAG	2 cut(s)	270, 893
Bpu1102I	GCTNAGC	1 cut(s)	324
BpuEI	CTTGAG	1 cut(s)	842
BsaBI	GATNNNNATC	2 cut(s)	845, 1334
BsaJI	CCNNGG	1 cut(s)	1143
BsaWI	WCCGGW	1 cut(s)	857
BsaXI	ACNNNNNCTCC	2 cut(s)	1132, 1162
Bse118I	RCCGGY	1 cut(s)	257
Bse1I	ACTGG	5 cut(s)	253, 285, 297, 642, 1296
Bse3DI	GCAATG	2 cut(s)	20, 1117
Bse8I	GATNNNNATC	2 cut(s)	845, 1334
BseAI	TCCGGA	1 cut(s)	857
BseDI	CCNNGG	1 cut(s)	1143
BseJI	GATNNNNATC	2 cut(s)	845, 1334
BseMI	GCAATG	2 cut(s)	20, 1117
BseNI	ACTGG	5 cut(s)	253, 285, 297, 642, 1296
BseRI	GAGGAG	1 cut(s)	1140
BseXI	GCAGC	9 cut(s)	237, 937, 940, 968, 1012, 1237, 1265, 1270, 1452
BsgI	GTGCAG	1 cut(s)	1451
Bsh1236I	CGCG	1 cut(s)	584
BshFI	GGCC	3 cut(s)	284, 309, 1323
BsiHKAI	GWGCWC	1 cut(s)	1009
BsiSI	CCGG	3 cut(s)	258, 858, 875
BslFI	GGGAC	1 cut(s)	142
BsmAI	GTCTC	3 cut(s)	109, 911, 1043
BsmFI	GGGAC	1 cut(s)	142
BsnI	GGCC	3 cut(s)	284, 309, 1323
Bsp1286I	GDGCHC	1 cut(s)	1009
Bsp13I	TCCGGA	1 cut(s)	857
Bsp1407I	TGTACA	1 cut(s)	1388
Bsp143I	GATC	2 cut(s)	150, 445
Bsp1720I	GCTNAGC	1 cut(s)	324
Bsp19I	CCATGG	1 cut(s)	1143
BspANI	GGCC	3 cut(s)	284, 309, 1323
BspEI	TCCGGA	1 cut(s)	857
BspFNI	CGCG	1 cut(s)	584
BspLI	GGNNCC	2 cut(s)	1149, 1322
BspMAI	CTGCAG	1 cut(s)	958
BspPI	GGATC	2 cut(s)	145, 440
BsrDI	GCAATG	2 cut(s)	20, 1117
BsrFI	RCCGGY	1 cut(s)	257
BsrGI	TGTACA	1 cut(s)	1388
BsrI	ACTGG	5 cut(s)	253, 285, 297, 642, 1296
BssAI	RCCGGY	1 cut(s)	257
BssECI	CCNNGG	1 cut(s)	1143
BssMI	GATC	2 cut(s)	150, 445
BssSI	CACGAG	1 cut(s)	459
BssT1I	CCWWGG	1 cut(s)	1143
Bst2BI	CACGAG	1 cut(s)	459
Bst4CI	ACNGT	2 cut(s)	120, 925
Bst6I	CTCTTC	2 cut(s)	107, 522
BstAPI	GCANNNNNTGC	1 cut(s)	1022
BstAUI	TGTACA	1 cut(s)	1388
BstC8I	GCNNGC	5 cut(s)	10, 213, 1081, 1184, 1188
BstDEI	CTNAG	3 cut(s)	324, 379, 572
BstDSI	CCRYGG	1 cut(s)	1143
BstEII	GGTNACC	1 cut(s)	992
BstFNI	CGCG	1 cut(s)	584
BstHHI	GCGC	1 cut(s)	584
BstKTI	GATC	2 cut(s)	153, 448
BstMAI	GTCTC	3 cut(s)	109, 911, 1043
BstMBI	GATC	2 cut(s)	150, 445
BstMWI	GCNNNNNNNGC	8 cut(s)	320, 503, 850, 956, 1013, 1022, 1292, 1415
BstNSI	RCATGY	1 cut(s)	1023
BstPI	GGTNACC	1 cut(s)	992
BstSFI	CTRYAG	2 cut(s)	954, 1056
BstUI	CGCG	1 cut(s)	584
BstV1I	GCAGC	9 cut(s)	237, 937, 940, 968, 1012, 1237, 1265, 1270, 1452
BstV2I	GAAGAC	1 cut(s)	45
BstX2I	RGATCY	1 cut(s)	445
BstXI	CCANNNNNNTGG	2 cut(s)	292, 317
BstYI	RGATCY	1 cut(s)	445
BsuI	GTATCC	1 cut(s)	1296
BsuRI	GGCC	3 cut(s)	284, 309, 1323
BtgI	CCRYGG	1 cut(s)	1143
BtgZI	GCGATG	1 cut(s)	1043
BtsI	GCAGTG	1 cut(s)	357
BtsIMutI	CAGTG	4 cut(s)	125, 246, 292, 357
Cac8I	GCNNGC	5 cut(s)	10, 213, 1081, 1184, 1188
CfoI	GCGC	1 cut(s)	584
Cfr10I	RCCGGY	1 cut(s)	257
Cfr13I	GGNCC	2 cut(s)	134, 1321
Csp6I	GTAC	2 cut(s)	72, 1389
CviAII	CATG	5 cut(s)	87, 415, 709, 1020, 1144
CviQI	GTAC	2 cut(s)	72, 1389
DdeI	CTNAG	3 cut(s)	324, 379, 572
DpnI	GATC	2 cut(s)	152, 447
DpnII	GATC	2 cut(s)	150, 445
DraIII	CACNNNGTG	1 cut(s)	1234
EaeI	YGGCCR	2 cut(s)	282, 307
Eam1104I	CTCTTC	2 cut(s)	107, 522
EarI	CTCTTC	2 cut(s)	107, 522
Eco130I	CCWWGG	1 cut(s)	1143
Eco47I	GGWCC	1 cut(s)	134
Eco57I	CTGAAG	3 cut(s)	57, 769, 873
Eco91I	GGTNACC	1 cut(s)	992
EcoO109I	RGGNCCY	2 cut(s)	134, 1321
EcoO65I	GGTNACC	1 cut(s)	992
EcoRI	GAATTC	2 cut(s)	861, 1272
EcoT14I	CCWWGG	1 cut(s)	1143
ErhI	CCWWGG	1 cut(s)	1143
FaeI	CATG	5 cut(s)	90, 418, 712, 1023, 1147
FalI	AAGNNNNNCTT	2 cut(s)	1179, 1211
FaqI	GGGAC	1 cut(s)	142
FatI	CATG	5 cut(s)	86, 414, 708, 1019, 1143
FauNDI	CATATG	2 cut(s)	316, 940
FblI	GTMKAC	1 cut(s)	388
Fnu4HI	GCNGC	9 cut(s)	226, 951, 954, 957, 1026, 1226, 1254, 1284, 1466
Fsp4HI	GCNGC	9 cut(s)	226, 951, 954, 957, 1026, 1226, 1254, 1284, 1466
FspBI	CTAG	9 cut(s)	222, 620, 738, 983, 1154, 1175, 1209, 1310, 1425
GlaI	GCGC	1 cut(s)	583
GluI	GCNGC	9 cut(s)	226, 951, 954, 957, 1026, 1226, 1254, 1284, 1466
GsuI	CTGGAG	2 cut(s)	270, 893
HaeIII	GGCC	3 cut(s)	284, 309, 1323
HapII	CCGG	3 cut(s)	258, 858, 875
HhaI	GCGC	1 cut(s)	584
Hin1II	CATG	5 cut(s)	90, 418, 712, 1023, 1147
Hin6I	GCGC	1 cut(s)	582
HinP1I	GCGC	1 cut(s)	582
HinfI	GANTC	3 cut(s)	252, 771, 1335
HpaII	CCGG	3 cut(s)	258, 858, 875
HphI	GGTGA	2 cut(s)	218, 247
Hpy166II	GTNNAC	2 cut(s)	290, 389
Hpy188I	TCNGA	1 cut(s)	336
Hpy8I	GTNNAC	2 cut(s)	290, 389
HpyAV	CCTTC	4 cut(s)	459, 810, 995, 1311
HpyCH4III	ACNGT	2 cut(s)	120, 925
HpyCH4V	TGCA	9 cut(s)	512, 731, 806, 956, 1040, 1186, 1253, 1357, 1468
HpyF10VI	GCNNNNNNNGC	8 cut(s)	320, 503, 850, 956, 1013, 1022, 1292, 1415
HpyF3I	CTNAG	3 cut(s)	324, 379, 572
Hsp92II	CATG	5 cut(s)	90, 418, 712, 1023, 1147
HspAI	GCGC	1 cut(s)	582
Kpn2I	TCCGGA	1 cut(s)	857
Kzo9I	GATC	2 cut(s)	150, 445
LmnI	GCTCC	3 cut(s)	320, 986, 1153
Lsp1109I	GCAGC	9 cut(s)	237, 937, 940, 968, 1012, 1237, 1265, 1270, 1452
LweI	GCATC	1 cut(s)	831
MaeI	CTAG	9 cut(s)	222, 620, 738, 983, 1154, 1175, 1209, 1310, 1425
MaeIII	GTNAC	8 cut(s)	122, 197, 253, 919, 992, 1060, 1135, 1378
MalI	GATC	2 cut(s)	152, 447
MboI	GATC	2 cut(s)	150, 445
MboII	GAAGA	6 cut(s)	50, 116, 124, 509, 711, 775
MfeI	CAATTG	1 cut(s)	1278
MflI	RGATCY	1 cut(s)	445
MhlI	GDGCHC	1 cut(s)	1009
MlsI	TGGCCA	2 cut(s)	284, 309
MluNI	TGGCCA	2 cut(s)	284, 309
MlyI	GAGTC	1 cut(s)	261
MmeI	TCCRAC	2 cut(s)	628, 999
Mox20I	TGGCCA	2 cut(s)	284, 309
MroI	TCCGGA	1 cut(s)	857
MscI	TGGCCA	2 cut(s)	284, 309
MseI	TTAA	2 cut(s)	554, 627
MslI	CAYNNNNRTG	2 cut(s)	204, 1142
Msp20I	TGGCCA	2 cut(s)	284, 309
MspI	CCGG	3 cut(s)	258, 858, 875
MunI	CAATTG	1 cut(s)	1278
MvnI	CGCG	1 cut(s)	584
MwoI	GCNNNNNNNGC	8 cut(s)	320, 503, 850, 956, 1013, 1022, 1292, 1415
NcoI	CCATGG	1 cut(s)	1143
NdeI	CATATG	2 cut(s)	316, 940
NdeII	GATC	2 cut(s)	150, 445
NlaIII	CATG	5 cut(s)	90, 418, 712, 1023, 1147
NlaIV	GGNNCC	2 cut(s)	1149, 1322
NmuCI	GTSAC	5 cut(s)	197, 253, 919, 1135, 1378
NspI	RCATGY	1 cut(s)	1023
OliI	CACNNNNGTG	1 cut(s)	204
PciI	ACATGT	1 cut(s)	1019
PfeI	GAWTC	2 cut(s)	771, 1335
PkrI	GCNGC	9 cut(s)	227, 952, 955, 958, 1027, 1227, 1255, 1285, 1467
PleI	GAGTC	1 cut(s)	260
PpsI	GAGTC	1 cut(s)	260
PpuMI	RGGWCCY	1 cut(s)	134
PscI	ACATGT	1 cut(s)	1019
Psp5II	RGGWCCY	1 cut(s)	134
PspEI	GGTNACC	1 cut(s)	992
PspN4I	GGNNCC	2 cut(s)	1149, 1322
PspPI	GGNCC	2 cut(s)	134, 1321
PspPPI	RGGWCCY	1 cut(s)	134
PstI	CTGCAG	1 cut(s)	958
PsuI	RGATCY	1 cut(s)	445
RsaI	GTAC	2 cut(s)	73, 1390
RsaNI	GTAC	2 cut(s)	72, 1389
RseI	CAYNNNNRTG	2 cut(s)	204, 1142
SaqAI	TTAA	2 cut(s)	554, 627
SatI	GCNGC	9 cut(s)	226, 951, 954, 957, 1026, 1226, 1254, 1284, 1466
Sau3AI	GATC	2 cut(s)	150, 445
Sau96I	GGNCC	2 cut(s)	134, 1321
ScaI	AGTACT	1 cut(s)	73
SchI	GAGTC	1 cut(s)	261
SduI	GDGCHC	1 cut(s)	1009
SfaNI	GCATC	1 cut(s)	831
SfcI	CTRYAG	2 cut(s)	954, 1056
SinI	GGWCC	1 cut(s)	134
SmiMI	CAYNNNNRTG	2 cut(s)	204, 1142
SmlI	CTYRAG	1 cut(s)	821
SmoI	CTYRAG	1 cut(s)	821
SspMI	CTAG	9 cut(s)	222, 620, 738, 983, 1154, 1175, 1209, 1310, 1425
StyI	CCWWGG	1 cut(s)	1143
TaaI	ACNGT	2 cut(s)	120, 925
TaqI	TCGA	2 cut(s)	182, 1449
TatI	WGTACW	2 cut(s)	71, 1388
TfiI	GAWTC	2 cut(s)	771, 1335
Tru1I	TTAA	2 cut(s)	554, 627
Tru9I	TTAA	2 cut(s)	554, 627
TscAI	CASTG	4 cut(s)	125, 253, 292, 364
TseFI	GTSAC	5 cut(s)	197, 253, 919, 1135, 1378
TseI	GCWGC	9 cut(s)	225, 950, 953, 956, 1025, 1225, 1253, 1283, 1465
Tsp45I	GTSAC	5 cut(s)	197, 253, 919, 1135, 1378
TspDTI	ATGAA	2 cut(s)	663, 725
TspRI	CASTG	4 cut(s)	125, 253, 292, 364
VpaK11BI	GGWCC	1 cut(s)	134
XapI	RAATTY	3 cut(s)	473, 861, 1272
XceI	RCATGY	1 cut(s)	1023
XmiI	GTMKAC	1 cut(s)	388
XspI	CTAG	9 cut(s)	222, 620, 738, 983, 1154, 1175, 1209, 1310, 1425
ZrmI	AGTACT	1 cut(s)	73

Using CommOnly database (standard laboratory enzymes). Scanned on CDS sequence.