FvH4_6g12550

Belongs to the UDP-glycosyltransferase family

Design Primers CRISPR Design

Basic Information

Type: gene
Biological Identity
fragaria_vesca
Fvb6
OG0000638
Physical Location & Seq
Forward (+)
7594780 .. 7596366
1587 bp
Loading structure...
UTR
Exon/CDS
Intron
FvH4_6g12550.t1

Sequence Viewer

Length: 894 bp
ATGGAGAAATCTCAATCTCACGTAGTGATCTTTCCATTTCCACTACAAGGCCACATCAAACCATTGCTATGCTTAGCAGAGCTTCTCTGCCATGCAGGCCTTCATGTCACCTTCGTCAACACCCGCCACAACCACAACCGCTTAGCCAACCTCCCTGCCCTCTCCGCCCGCTTCCCCACCCTCAATTTCGAGTCCATCTCGGACGGTCTCCCCCTTGAGCACCCGCGTACTTTGGGCAGCGAGTTGTTGATGGCGTTGAAGACGTCGATTAAGCCTCATTTTAAGGAGCTGCTCCAGTCACCCCTTAAGGCTAACGGTGTTGCGCTACCGGCGCCAACATGTATTATTACAGATGGACTTGTGACTTTTGCCTTTGATGTTGCAGAGGAGTTGGGGTTACCTATACTTGGTTATAATGTTCCATGTACTCGTTACATGTGGACTTGTCTTTGTCTTCCCAATCTCATAGAACAAGGCCAGCTTCCTTTCCCAGATGACGACATGAATGTGGAAATTACCGGCGTCCCGGGAATGGAAGGCCTTCTGCGACGTGTAGATTTACCTGGTTTTTGCAGAGTAAAACAGTCGAACCATCCTGGCCTTCAATTTGCCATCCGCGAAACTCAGACACAAAAGCGAGCTTCGGCTCTAATCCTCGACACCATTTACGAGCTTGACGCTCCATGTCTGTCTCATATGGCCAACATGTTCCCTAAAATCTACACTCTCGGCCCTCTCCACTCTCTCCTACATTCTCAAATCGGCGACGTCTCGCGATCCTTAGCATCCTACGGCGTGGCTCGTGATAGTGTTGTGAAAGGTGGATCTTCAAACCATAATTTGGAGATGCTAATTCAAGACATTAGGAATATGCATGCAAGACCACAGCAGTGA
Functional Annotation
Gene Ontology
Molecular Function
Biological Process
Cellular Component
No ontology terms assigned.
KEGG Pathways
Metabolic & Signaling
No pathways identified.
Pfam Domains
Protein Families
UDPGT

Protein Analysis

298

Amino Acids

32.85

Weight (kDa)

6.69

Isoelectric Point (pI)

55.49

Instability Index
Protein Domains (Pfam)
No Pfam domains detected for this protein.
Hydrophobicity Profile (Kyte-Doolittle)
AI Structure Prediction Report
Calculating structure properties...

Genomic Context

Gene Family Tree

Species Filter
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Publication-ready
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Orthologous Genes (Group: OG0000638)

Species Orthologous Gene IDs
fragaria_vesca FvH4_5g19301 FvH4_6g12520 FvH4_6g12520 FvH4_6g12550 FvH4_6g12560 FvH4_6g12580
malus_domestica MD04G1140700.v1.1 MD04G1141300.v1.1 MD04G1141400.v1.1 MD04G1141500.v1.1 MD12G1156200.v1.1
prunus_persica Prupe.6G265900_v2.0.a1 Prupe.6G266600_v2.0.a1 Prupe.6G266700_v2.0.a1 Prupe.6G266900_v2.0.a1 Prupe.6G267000_v2.0.a1
pyrus_communis pycom04g12860
rosa_chinensis RchiOBHm_Chr2g0160211 RchiOBHm_Chr3g0464331 RchiOBHm_Chr3g0464341 RchiOBHm_Chr3g0464361 RchiOBHm_Chr3g0464371 RchiOBHm_Chr3g0464421 RchiOBHm_Chr3g0464441
rosa_laevigata RLG00000001199 RLG00000007070 RLG00000015964 RLG00000020699 RLG00000024697 RLG00000024699 RLG00000024701 RLG00000024702 RLG00000024703 RLG00000024705 RLG00000027329 RLG00000034971 RLG00000035066
rosa_multiflora Rmu_sc0003074.1_g000014 Rmu_sc0003074.1_g000020 Rmu_sc0003074.1_g000024 Rmu_sc0003074.1_g000025 Rmu_sc0003074.1_g000027 Rmu_sc0003074.1_g000029 Rmu_sc0039307.1_g000001
rosa_roxburghii Rroxscaffold_1G00025190 Rroxscaffold_2G00078980 Rroxscaffold_6G00415680 Rroxscaffold_6G00415710 Rroxscaffold_6G00415740 Rroxscaffold_6G00415770 Rroxscaffold_6G00415800
rosa_rugosa Rorug03G0071500 Rorug03G0071700 Rorug03G0071800 Rorug03G0072300
rosa_samantha Rh3BG133600 Rh3BG133700 Rh3BG133800 Rh3BG133900 Rh3BG134100 Rh3BG134200 Rh3CG135900 Rh3CG136100 Rh3CG136200 Rh3CG136300 Rh3CG136600 Rh3DG135100 Rh3DG135200 Rh3DG135300 Rh3DG135400 Rh3DG135600 Rh4BG384400 Rh5CG268300
rosa_wichuraiana Rw3G010950 Rw3G010960 Rw3G010970 Rw3G010980 Rw3G012210

Restriction Enzyme Sites

1 / 10
Enzyme Recognition Site Cut Count Positions (bp)
AanI TTATAA 1 cut(s) 414
AatII GACGTC 2 cut(s) 266, 771
AccB1I GGYRCC 1 cut(s) 331
AccB7I CCANNNNNTGG 1 cut(s) 841
AccII CGCG 3 cut(s) 226, 618, 775
AciI CCGC 6 cut(s) 124, 139, 165, 169, 224, 616
AclWI GGATC 2 cut(s) 771, 832
AcoI YGGCCR 1 cut(s) 699
AcyI GRCGYC 4 cut(s) 263, 332, 522, 768
AdeI CACNNNGTG 1 cut(s) 25
AfaI GTAC 2 cut(s) 229, 427
AfiI CCNNNNNNNGG 4 cut(s) 47, 407, 532, 841
AflII CTTAAG 1 cut(s) 305
AflIII ACRYGT 4 cut(s) 338, 435, 550, 705
AgsI TTSAA 4 cut(s) 259, 605, 831, 857
AjiI CACGTC 1 cut(s) 551
AjnI CCWGG 2 cut(s) 562, 595
AleI CACNNNNGTG 1 cut(s) 889
AluBI AGCT 5 cut(s) 82, 289, 481, 641, 673
AluI AGCT 5 cut(s) 82, 289, 481, 641, 673
Alw21I GWGCWC 1 cut(s) 222
Alw26I GTCTC 3 cut(s) 212, 696, 775
AlwI GGATC 2 cut(s) 771, 832
Ama87I CYCGRG 1 cut(s) 526
AoxI GGCC 7 cut(s) 49, 97, 475, 538, 598, 699, 730
ApeKI GCWGC 2 cut(s) 237, 289
Asp700I GAANNNNTTC 1 cut(s) 540
AspLEI GCGC 2 cut(s) 325, 334
AspS9I GGNCC 1 cut(s) 731
AsuC2I CCSGG 2 cut(s) 527, 528
AsuHPI GGTGA 2 cut(s) 100, 291
AvaI CYCGRG 1 cut(s) 526
BalI TGGCCA 1 cut(s) 701
BanI GGYRCC 1 cut(s) 331
BauI CACGAG 1 cut(s) 801
BbsI GAAGAC 2 cut(s) 266, 446
Bbv12I GWGCWC 1 cut(s) 222
BbvI GCAGC 2 cut(s) 249, 276
BccI CCATC 5 cut(s) 203, 244, 347, 600, 620
BceAI ACGGC 1 cut(s) 808
BciT130I CCWGG 2 cut(s) 564, 597
BcnI CCSGG 2 cut(s) 527, 528
BcoDI GTCTC 3 cut(s) 212, 696, 775
BfoI RGCGCY 1 cut(s) 335
BfrI CTTAAG 1 cut(s) 305
BglI GCCNNNNNGGC 1 cut(s) 96
BisI GCNGC 2 cut(s) 238, 290
BlpI GCTNAGC 2 cut(s) 73, 142
BlsI GCNGC 2 cut(s) 239, 291
Bme1390I CCNGG 4 cut(s) 527, 528, 564, 597
BmeT110I CYCGRG 1 cut(s) 526
BmgBI CACGTC 1 cut(s) 551
BmgT120I GGNCC 1 cut(s) 731
BmiI GGNNCC 1 cut(s) 333
BmrFI CCNGG 4 cut(s) 527, 528, 564, 597
BmsI GCATC 2 cut(s) 794, 837
BpiI GAAGAC 2 cut(s) 266, 446
BplI GAGNNNNNCTC 2 cut(s) 182, 214
BpmI CTGGAG 1 cut(s) 278
Bpu10I CCTNAGC 1 cut(s) 781
Bpu1102I GCTNAGC 2 cut(s) 73, 142
BpuEI CTTGAG 1 cut(s) 236
BpuMI CCSGG 2 cut(s) 527, 528
BsaAI YACGTR 1 cut(s) 22
BsaHI GRCGYC 4 cut(s) 263, 332, 522, 768
BsaI GGTCTC 1 cut(s) 212
BsaJI CCNNGG 1 cut(s) 526
BsaXI ACNNNNNCTCC 2 cut(s) 380, 410
Bsc4I CCNNNNNNNGG 4 cut(s) 47, 407, 532, 841
Bse118I RCCGGY 2 cut(s) 328, 518
Bse1I ACTGG 1 cut(s) 295
Bse3DI GCAATG 1 cut(s) 62
BseBI CCWGG 2 cut(s) 564, 597
BseDI CCNNGG 1 cut(s) 526
BseGI GGATG 3 cut(s) 592, 612, 785
BseLI CCNNNNNNNGG 4 cut(s) 47, 407, 532, 841
BseMI GCAATG 1 cut(s) 62
BseMII CTCAG 1 cut(s) 638
BseNI ACTGG 1 cut(s) 295
BseRI GAGGAG 1 cut(s) 401
BseXI GCAGC 2 cut(s) 249, 276
Bsh1236I CGCG 3 cut(s) 226, 618, 775
BshFI GGCC 7 cut(s) 51, 99, 477, 540, 600, 701, 732
BshNI GGYRCC 1 cut(s) 331
BsiHKAI GWGCWC 1 cut(s) 222
BsiHKCI CYCGRG 1 cut(s) 526
BsiSI CCGG 3 cut(s) 329, 519, 527
BslFI GGGAC 1 cut(s) 509
BslI CCNNNNNNNGG 4 cut(s) 47, 407, 532, 841
BsmAI GTCTC 3 cut(s) 212, 696, 775
BsmBI CGTCTC 1 cut(s) 775
BsmFI GGGAC 1 cut(s) 509
BsnI GGCC 7 cut(s) 51, 99, 477, 540, 600, 701, 732
Bso31I GGTCTC 1 cut(s) 212
BsoBI CYCGRG 1 cut(s) 526
Bsp1286I GDGCHC 1 cut(s) 222
Bsp143I GATC 3 cut(s) 27, 776, 824
Bsp1720I GCTNAGC 2 cut(s) 73, 142
Bsp68I TCGCGA 1 cut(s) 775
BspACI CCGC 6 cut(s) 124, 139, 165, 169, 224, 616
BspANI GGCC 7 cut(s) 51, 99, 477, 540, 600, 701, 732
BspCNI CTCAG 1 cut(s) 637
BspFNI CGCG 3 cut(s) 226, 618, 775
BspLI GGNNCC 1 cut(s) 333
BspPI GGATC 2 cut(s) 771, 832
BspT107I GGYRCC 1 cut(s) 331
BspTI CTTAAG 1 cut(s) 305
BspTNI GGTCTC 1 cut(s) 212
BsrDI GCAATG 1 cut(s) 62
BsrFI RCCGGY 2 cut(s) 328, 518
BsrI ACTGG 1 cut(s) 295
BssAI RCCGGY 2 cut(s) 328, 518
BssECI CCNNGG 1 cut(s) 526
BssMI GATC 3 cut(s) 27, 776, 824
BssNI GRCGYC 4 cut(s) 263, 332, 522, 768
BssSI CACGAG 1 cut(s) 801
Bst2BI CACGAG 1 cut(s) 801
Bst2UI CCWGG 2 cut(s) 564, 597
Bst4CI ACNGT 3 cut(s) 206, 317, 585
BstACI GRCGYC 4 cut(s) 263, 332, 522, 768
BstAFI CTTAAG 1 cut(s) 305
BstBAI YACGTR 1 cut(s) 22
BstC8I GCNNGC 5 cut(s) 97, 169, 479, 639, 876
BstDEI CTNAG 4 cut(s) 73, 142, 624, 781
BstEII GGTNACC 1 cut(s) 396
BstF5I GGATG 3 cut(s) 592, 612, 785
BstFNI CGCG 3 cut(s) 226, 618, 775
BstH2I RGCGCY 1 cut(s) 335
BstHHI GCGC 2 cut(s) 325, 334
BstKTI GATC 3 cut(s) 30, 779, 827
BstMAI GTCTC 3 cut(s) 212, 696, 775
BstMBI GATC 3 cut(s) 27, 776, 824
BstMWI GCNNNNNNNGC 4 cut(s) 96, 164, 329, 331
BstNI CCWGG 2 cut(s) 564, 597
BstNSI RCATGY 4 cut(s) 342, 439, 709, 878
BstPI GGTNACC 1 cut(s) 396
BstSCI CCNGG 4 cut(s) 525, 526, 562, 595
BstUI CGCG 3 cut(s) 226, 618, 775
BstV1I GCAGC 2 cut(s) 249, 276
BstV2I GAAGAC 2 cut(s) 266, 446
BstX2I RGATCY 1 cut(s) 824
BstYI RGATCY 1 cut(s) 824
BsuRI GGCC 7 cut(s) 51, 99, 477, 540, 600, 701, 732
BtrI CACGTC 1 cut(s) 551
BtsCI GGATG 3 cut(s) 592, 612, 785
BtuMI TCGCGA 1 cut(s) 775
Cac8I GCNNGC 5 cut(s) 97, 169, 479, 639, 876
CfoI GCGC 2 cut(s) 325, 334
Cfr10I RCCGGY 2 cut(s) 328, 518
Cfr13I GGNCC 1 cut(s) 731
Cfr9I CCCGGG 1 cut(s) 526
CseI GACGC 2 cut(s) 511, 686
CsiI ACCWGGT 1 cut(s) 562
Csp6I GTAC 2 cut(s) 228, 426
CviAII CATG 9 cut(s) 92, 104, 339, 423, 436, 502, 684, 706, 875
CviQI GTAC 2 cut(s) 228, 426
DdeI CTNAG 4 cut(s) 73, 142, 624, 781
DinI GGCGCC 1 cut(s) 333
DpnI GATC 3 cut(s) 29, 778, 826
DpnII GATC 3 cut(s) 27, 776, 824
DraIII CACNNNGTG 1 cut(s) 25
EaeI YGGCCR 1 cut(s) 699
EciI GGCGGA 1 cut(s) 154
Eco147I AGGCCT 2 cut(s) 99, 540
Eco31I GGTCTC 1 cut(s) 212
Eco88I CYCGRG 1 cut(s) 526
Eco91I GGTNACC 1 cut(s) 396
EcoO65I GGTNACC 1 cut(s) 396
EcoRII CCWGG 2 cut(s) 562, 595
EcoT22I ATGCAT 1 cut(s) 876
EgeI GGCGCC 1 cut(s) 333
EheI GGCGCC 1 cut(s) 333
Esp3I CGTCTC 1 cut(s) 775
FaeI CATG 9 cut(s) 95, 107, 342, 426, 439, 505, 687, 709, 878
FalI AAGNNNNNCTT 2 cut(s) 465, 497
FaqI GGGAC 1 cut(s) 509
FatI CATG 9 cut(s) 91, 103, 338, 422, 435, 501, 683, 705, 874
FauI CCCGC 3 cut(s) 131, 176, 231
FauNDI CATATG 1 cut(s) 696
Fnu4HI GCNGC 2 cut(s) 238, 290
FokI GGATG 3 cut(s) 579, 599, 772
Fsp4HI GCNGC 2 cut(s) 238, 290
GlaI GCGC 2 cut(s) 324, 333
GluI GCNGC 2 cut(s) 238, 290
GsuI CTGGAG 1 cut(s) 278
HaeII RGCGCY 1 cut(s) 335
HaeIII GGCC 7 cut(s) 51, 99, 477, 540, 600, 701, 732
HapII CCGG 3 cut(s) 329, 519, 527
HgaI GACGC 2 cut(s) 511, 686
HhaI GCGC 2 cut(s) 325, 334
Hin1I GRCGYC 4 cut(s) 263, 332, 522, 768
Hin1II CATG 9 cut(s) 95, 107, 342, 426, 439, 505, 687, 709, 878
Hin6I GCGC 2 cut(s) 323, 332
HinP1I GCGC 2 cut(s) 323, 332
HincII GTYRAC 1 cut(s) 118
HindII GTYRAC 1 cut(s) 118
HinfI GANTC 1 cut(s) 191
HpaII CCGG 3 cut(s) 329, 519, 527
HphI GGTGA 2 cut(s) 100, 291
Hpy166II GTNNAC 2 cut(s) 118, 441
Hpy188I TCNGA 2 cut(s) 202, 627
Hpy188III TCNNGA 3 cut(s) 774, 803, 857
Hpy8I GTNNAC 2 cut(s) 118, 441
Hpy99I CGWCG 3 cut(s) 268, 552, 770
HpyAV CCTTC 5 cut(s) 110, 121, 530, 551, 611
HpyCH4III ACNGT 3 cut(s) 206, 317, 585
HpyCH4IV ACGT 4 cut(s) 21, 263, 550, 768
HpyCH4V TGCA 5 cut(s) 95, 383, 573, 874, 878
HpyF10VI GCNNNNNNNGC 4 cut(s) 96, 164, 329, 331
HpyF3I CTNAG 4 cut(s) 73, 142, 624, 781
HpySE526I ACGT 4 cut(s) 21, 263, 550, 768
Hsp92I GRCGYC 4 cut(s) 263, 332, 522, 768
Hsp92II CATG 9 cut(s) 95, 107, 342, 426, 439, 505, 687, 709, 878
HspAI GCGC 2 cut(s) 323, 332
KasI GGCGCC 1 cut(s) 331
Kzo9I GATC 3 cut(s) 27, 776, 824
LmnI GCTCC 3 cut(s) 286, 297, 685
Lsp1109I GCAGC 2 cut(s) 249, 276
LweI GCATC 2 cut(s) 794, 837
MabI ACCWGGT 1 cut(s) 562
MaeII ACGT 4 cut(s) 21, 263, 550, 768
MaeIII GTNAC 5 cut(s) 106, 297, 361, 396, 431
MalI GATC 3 cut(s) 29, 778, 826
MboI GATC 3 cut(s) 27, 776, 824
MboII GAAGA 3 cut(s) 271, 446, 819
MflI RGATCY 1 cut(s) 824
MhlI GDGCHC 1 cut(s) 222
MlsI TGGCCA 1 cut(s) 701
MluCI AATT 5 cut(s) 184, 513, 605, 838, 852
MluNI TGGCCA 1 cut(s) 701
Mly113I GGCGCC 1 cut(s) 332
MlyI GAGTC 1 cut(s) 200
MnlI CCTC 7 cut(s) 161, 170, 191, 285, 379, 665, 744
Mox20I TGGCCA 1 cut(s) 701
Mph1103I ATGCAT 1 cut(s) 876
MroXI GAANNNNTTC 1 cut(s) 540
MscI TGGCCA 1 cut(s) 701
MseI TTAA 3 cut(s) 270, 282, 306
MslI CAYNNNNRTG 3 cut(s) 67, 506, 889
Msp20I TGGCCA 1 cut(s) 701
MspCI CTTAAG 1 cut(s) 305
MspI CCGG 3 cut(s) 329, 519, 527
MspR9I CCNGG 4 cut(s) 527, 528, 564, 597
MvaI CCWGG 2 cut(s) 564, 597
MvnI CGCG 3 cut(s) 226, 618, 775
MwoI GCNNNNNNNGC 4 cut(s) 96, 164, 329, 331
NarI GGCGCC 1 cut(s) 332
NciI CCSGG 2 cut(s) 527, 528
NdeI CATATG 1 cut(s) 696
NdeII GATC 3 cut(s) 27, 776, 824
NlaIII CATG 9 cut(s) 95, 107, 342, 426, 439, 505, 687, 709, 878
NlaIV GGNNCC 1 cut(s) 333
NmeAIII GCCGAG 1 cut(s) 708
NmuCI GTSAC 3 cut(s) 106, 297, 361
NruI TCGCGA 1 cut(s) 775
NsiI ATGCAT 1 cut(s) 876
NspI RCATGY 4 cut(s) 342, 439, 709, 878
OliI CACNNNNGTG 1 cut(s) 889
PaeI GCATGC 1 cut(s) 878
PceI AGGCCT 2 cut(s) 99, 540
PciI ACATGT 3 cut(s) 338, 435, 705
PdmI GAANNNNTTC 1 cut(s) 540
PflMI CCANNNNNTGG 1 cut(s) 841
PkrI GCNGC 2 cut(s) 239, 291
PleI GAGTC 1 cut(s) 199
PluTI GGCGCC 1 cut(s) 335
PpsI GAGTC 1 cut(s) 199
Ppu21I YACGTR 1 cut(s) 22
PscI ACATGT 3 cut(s) 338, 435, 705
PsiI TTATAA 1 cut(s) 414
Psp6I CCWGG 2 cut(s) 562, 595
PspEI GGTNACC 1 cut(s) 396
PspGI CCWGG 2 cut(s) 562, 595
PspN4I GGNNCC 1 cut(s) 333
PspPI GGNCC 1 cut(s) 731
PsuI RGATCY 1 cut(s) 824
RruI TCGCGA 1 cut(s) 775
RsaI GTAC 2 cut(s) 229, 427
RsaNI GTAC 2 cut(s) 228, 426
RseI CAYNNNNRTG 3 cut(s) 67, 506, 889
SaqAI TTAA 3 cut(s) 270, 282, 306
SatI GCNGC 2 cut(s) 238, 290
Sau3AI GATC 3 cut(s) 27, 776, 824
Sau96I GGNCC 1 cut(s) 731
SchI GAGTC 1 cut(s) 200
ScrFI CCNGG 4 cut(s) 527, 528, 564, 597
SduI GDGCHC 1 cut(s) 222
SexAI ACCWGGT 1 cut(s) 562
SfaNI GCATC 2 cut(s) 794, 837
SfoI GGCGCC 1 cut(s) 333
SmaI CCCGGG 1 cut(s) 528
SmiMI CAYNNNNRTG 3 cut(s) 67, 506, 889
SmlI CTYRAG 2 cut(s) 215, 305
SmoI CTYRAG 2 cut(s) 215, 305
SphI GCATGC 1 cut(s) 878
Sse9I AATT 5 cut(s) 184, 513, 605, 838, 852
SseBI AGGCCT 2 cut(s) 99, 540
SsiI CCGC 6 cut(s) 124, 139, 165, 169, 224, 616
SspDI GGCGCC 1 cut(s) 331
StuI AGGCCT 2 cut(s) 99, 540
StyD4I CCNGG 4 cut(s) 525, 526, 562, 595
TaaI ACNGT 3 cut(s) 206, 317, 585
TaiI ACGT 4 cut(s) 24, 266, 553, 771
TaqI TCGA 4 cut(s) 189, 266, 587, 657
TasI AATT 5 cut(s) 184, 513, 605, 838, 852
TatI WGTACW 1 cut(s) 425
Tru1I TTAA 3 cut(s) 270, 282, 306
Tru9I TTAA 3 cut(s) 270, 282, 306
TseFI GTSAC 3 cut(s) 106, 297, 361
TseI GCWGC 2 cut(s) 237, 289
Tsp45I GTSAC 3 cut(s) 106, 297, 361
TspDTI ATGAA 2 cut(s) 92, 518
TspMI CCCGGG 1 cut(s) 526
Van91I CCANNNNNTGG 1 cut(s) 841
Vha464I CTTAAG 1 cut(s) 305
XceI RCATGY 4 cut(s) 342, 439, 709, 878
XmaI CCCGGG 1 cut(s) 526
XmnI GAANNNNTTC 1 cut(s) 540
ZraI GACGTC 2 cut(s) 264, 769
Zsp2I ATGCAT 1 cut(s) 876
Using CommOnly database (standard laboratory enzymes). Scanned on CDS sequence.