Basic Information

Type: gene

Biological Identity

Species

rosa_samantha

Chromosome

Chr1B

Orthogroup ID

OG0000685

Physical Location & Seq

Genomic Coordinates Forward (+)

914847 .. 917374

Gene Structure 2528 bp

Loading structure...

UTR

Exon/CDS

Intron

Transcript / Protein ID

Rh1BG007800.1

Sequence Viewer

CDS Protein Promoter (2k)

Length: 1104 bp

ATGAAGCTGTGGCAAGACTTCATCAGCTTGGAAAGGATACTGTTGTTGATGCCGGGATATTTGATGGGTCTTTACTCTTTAGGCATCATTTCTGTGGTATCTGCATTGAAGGTTTTGAGTATCGGTGGAAAATTGGGAGAACTAAAGCGGCCGGTTGAGGTAATAGCATTTAGTGATGAAGAAGGAGTAAGGTTTCAGTCTACATTCATAGGAAGTGCTGCTATAGCTGGAATTTTACCAGTTTCAGCACTGCAAATAACTGATAAAAGTGGTGTGACAATTCAAGATGCTCTTAAGAAAAACTCCATTGAAGTTACAAAGGAGCACTTGCAGCAGCTCAGATATGACCCAAAGTCAGTTTGGGGTTATGTTAAGATTCACATTGAACAAGGGCCTGTACTGGAATGGGTTGGTTTTCCTCTAGGAGTTGTTAAAGGCATAGCTGGACAGACAAGACTAAAGGTTACAATGAGAGGTTCCCAGGGGCATGCTGGAACAGTTCCAATGTCCATGCGTCATGACCCTATGGCTGCTGCTGCAGAAGCCATTGTATTGTTAGAAAGCCTATGTAAGCATCCTCAAGATTTTCTGTCTTTTGATGGTCAATGCAAAAGTTACTCATTAGAATCACTTTCAACTTCACTCGTTTGTACCGTTGGTGAGATATCGACTTGGCCAAGTGCAAGTAATGTTATTCCAGGACAGGTAACATTCACAGTAGATTTACGTGCAATTGATGACATGGGACGTGAAGCTGTTGTATGTGAATTATCCAACCGATTGTATCAAATATGTGATAGGCGTTCAGTTTCGTGCACAATTGATCGTAAGCATGATGCAAATGCAGTCATTTGCGATTCTGAGCTGACTTCGAAGCTGAAGTCAGCAGCTTATCTTGGGCTCAAGAGAATGACAGGTTCTGTTCAGGATGAAGTACCTGTATTAATGAGTGGAGCAGGACATGATGCAATCGCTCTGTCTCATTTAACTAAGGTGGGTATGCTTTTTGTCCGCTGTCGAGGAGGCATAAGTCACTTCCCTGAAGAACATGTATTGGATGATATGATGTTTGGGCATCTGGTTTGGCAATCCTGGCATTCATAG

Functional Annotation

Gene Ontology

Molecular Function
Biological Process
Cellular Component

GO:0000255 GO:0000256 GO:0003674 GO:0003824 GO:0005575 GO:0005622 GO:0005623 GO:0005737 GO:0005783 GO:0006139 GO:0006144 GO:0006145 GO:0006725 GO:0006807 GO:0008150 GO:0008152 GO:0009056 GO:0009112 GO:0009987 GO:0010135 GO:0010136 GO:0012505 GO:0016787 GO:0016810 GO:0016813 GO:0017144 GO:0019439 GO:0034641 GO:0042737 GO:0043226 GO:0043227 GO:0043229 GO:0043231 GO:0043603 GO:0043605 GO:0044237 GO:0044238 GO:0044248 GO:0044270 GO:0044281 GO:0044424 GO:0044444 GO:0044464 GO:0046113 GO:0046483 GO:0046700 GO:0047652 GO:0055086 GO:0071704 GO:0072521 GO:0072523 GO:1901360 GO:1901361 GO:1901564 GO:1901565 GO:1901575

KEGG Pathways

Metabolic & Signaling

ko00230 ko01120 map00230 map01120

Pfam Domains

Protein Families

M20_dimer Peptidase_M20 Peptidase_M28

Protein Analysis

367

Amino Acids

40.13

Weight (kDa)

6.46

Isoelectric Point (pI)

37.66

Instability Index

Protein Domains (Pfam)

Domain Name	Pfam ID	Position	E-value	Description
Peptidase_M20	PF01546	31 - 354	4.4e-10	Peptidase family M20/M25/M40
M20_dimer	PF07687	149 - 261	1.2e-07	Peptidase dimerisation domain

Hydrophobicity Profile (Kyte-Doolittle)

AI Structure Prediction Report

Calculating structure properties...

Genomic Context

Gene Family Tree

Species Filter

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Align Labels (Cladogram)

Image

Tree File

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Publication-ready

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Orthologous Genes (Group: OG0000685)

Species	Orthologous Gene IDs
arabidopsis_thaliana	AT4G20070 AT4G20070
fragaria_vesca	FvH4_3g12850 FvH4_3g12850 FvH4_3g12850
malus_domestica	MD10G1228200.v1.1
prunus_persica	Prupe.4G116700_v2.0.a1 Prupe.4G116700_v2.0.a1 Prupe.4G116700_v2.0.a1 Prupe.4G116700_v2.0.a1 Prupe.4G116700_v2.0.a1 Prupe.4G116700_v2.0.a1 Prupe.4G116700_v2.0.a1 Prupe.4G116700_v2.0.a1
pyrus_communis	pycom10g19170
rosa_chinensis	RchiOBHm_Chr1g0315031 RchiOBHm_Chr1g0315061 RchiOBHm_Chr1g0315091 RchiOBHm_Chr1g0315141 RchiOBHm_Chr1g0315151 RchiOBHm_Chr1g0315201 RchiOBHm_Chr1g0315231 RchiOBHm_Chr1g0315241 RchiOBHm_Chr1g0315311 RchiOBHm_Chr1g0315381 RchiOBHm_Chr1g0315441 RchiOBHm_Chr1g0315471 RchiOBHm_Chr1g0315531 RchiOBHm_Chr1g0315541 RchiOBHm_Chr1g0315581 RchiOBHm_Chr1g0315591 RchiOBHm_Chr1g0315611 RchiOBHm_Chr5g0020791 RchiOBHm_Chr5g0020891 RchiOBHm_Chr5g0021441 RchiOBHm_Chr5g0021511 RchiOBHm_Chr5g0021521 RchiOBHm_Chr5g0029451 RchiOBHm_Chr5g0029471
rosa_laevigata	RLG00000030744 RLG00000030748 RLG00000032563 RLG00000032611
rosa_multiflora	Rmu_sc0000576.1_g000021 Rmu_sc0002052.1_g000003 Rmu_sc0004180.1_g000017 Rmu_sc0004180.1_g000030 Rmu_sc0004180.1_g000035 Rmu_sc0023070.1_g000002 Rmu_ssc0000244.1_g000002
rosa_roxburghii	Rroxscaffold_1G00057130 Rroxscaffold_1G00057510 Rroxscaffold_4G00331580
rosa_rugosa	Rorug01G0004300 Rorug05G0063000 Rorug05G0067300
rosa_samantha	Rh1AG015000 Rh1AG015600 Rh1BG007400 Rh1BG007800 Rh1CG013200 Rh5AG152900 Rh5BG152700 Rh5BG155800 Rh5CG164200 Rh5CG164700 Rh5CG170000 Rh5CG227100
rosa_wichuraiana	Rw1G000740 Rw1G000780 Rw1G000810 Rw1G000840 Rw1G000870 Rw1G000900 Rw5G013560

Restriction Enzyme Sites

1 / 10

Enzyme	Recognition Site	Cut Count	Positions (bp)
AccI	GTMKAC	1 cut(s)	200
AciI	CCGC	2 cut(s)	148, 1012
AcoI	YGGCCR	2 cut(s)	149, 674
AcsI	RAATTY	1 cut(s)	231
AcuI	CTGAAG	2 cut(s)	899, 1062
AfaI	GTAC	3 cut(s)	399, 652, 936
AflII	CTTAAG	1 cut(s)	293
AflIII	ACRYGT	1 cut(s)	1048
AgsI	TTSAA	5 cut(s)	109, 284, 311, 386, 636
AhdI	GACNNNNNGTC	1 cut(s)	352
AjiI	CACGTC	1 cut(s)	749
AjnI	CCWGG	3 cut(s)	480, 697, 1091
AluBI	AGCT	9 cut(s)	7, 27, 227, 337, 443, 755, 865, 877, 890
AluI	AGCT	9 cut(s)	7, 27, 227, 337, 443, 755, 865, 877, 890
Alw21I	GWGCWC	2 cut(s)	327, 818
Alw26I	GTCTC	1 cut(s)	984
Alw44I	GTGCAC	1 cut(s)	814
AlwNI	CAGNNNCTG	1 cut(s)	920
AoxI	GGCC	3 cut(s)	149, 392, 674
ApaLI	GTGCAC	1 cut(s)	814
ApeKI	GCWGC	7 cut(s)	218, 331, 334, 530, 533, 536, 887
ApoI	RAATTY	1 cut(s)	231
ArsI	GACNNNNNNTTYG	2 cut(s)	866, 898
AseI	ATTAAT	1 cut(s)	944
AspS9I	GGNCC	1 cut(s)	392
AsuC2I	CCSGG	1 cut(s)	54
AsuHPI	GGTGA	1 cut(s)	671
AsuII	TTCGAA	1 cut(s)	872
BaeGI	GKGCMC	1 cut(s)	818
BaeI	ACNNNNGTAYC	2 cut(s)	29, 62
BalI	TGGCCA	1 cut(s)	676
BanII	GRGCYC	1 cut(s)	903
Bbv12I	GWGCWC	2 cut(s)	327, 818
BbvI	GCAGC	7 cut(s)	205, 343, 346, 517, 520, 523, 899
BccI	CCATC	2 cut(s)	58, 593
BciT130I	CCWGG	3 cut(s)	482, 699, 1093
BciVI	GTATCC	1 cut(s)	30
BcnI	CCSGG	1 cut(s)	54
BcoDI	GTCTC	1 cut(s)	984
BfaI	CTAG	1 cut(s)	422
BfmI	CTRYAG	2 cut(s)	222, 537
BfrI	CTTAAG	1 cut(s)	293
BfuI	GTATCC	1 cut(s)	30
BisI	GCNGC	8 cut(s)	149, 219, 332, 335, 531, 534, 537, 888
BlsI	GCNGC	8 cut(s)	150, 220, 333, 336, 532, 535, 538, 889
Bme1390I	CCNGG	4 cut(s)	54, 482, 699, 1093
BmeRI	GACNNNNNGTC	1 cut(s)	352
BmgBI	CACGTC	1 cut(s)	749
BmgT120I	GGNCC	1 cut(s)	392
BmiI	GGNNCC	1 cut(s)	478
BmrFI	CCNGG	4 cut(s)	54, 482, 699, 1093
BmsI	GCATC	7 cut(s)	39, 93, 277, 583, 826, 955, 1084
Bpu14I	TTCGAA	1 cut(s)	872
BpuEI	CTTGAG	2 cut(s)	564, 887
BpuMI	CCSGG	1 cut(s)	54
BsaAI	YACGTR	1 cut(s)	728
BsaJI	CCNNGG	2 cut(s)	480, 481
Bse118I	RCCGGY	1 cut(s)	151
Bse1I	ACTGG	2 cut(s)	239, 405
BseBI	CCWGG	3 cut(s)	482, 699, 1093
BseDI	CCNNGG	2 cut(s)	480, 481
BseGI	GGATG	3 cut(s)	574, 934, 1063
BseMII	CTCAG	2 cut(s)	352, 852
BseNI	ACTGG	2 cut(s)	239, 405
BseRI	GAGGAG	1 cut(s)	1035
BseSI	GKGCMC	1 cut(s)	818
BseX3I	CGGCCG	1 cut(s)	149
BseXI	GCAGC	7 cut(s)	205, 343, 346, 517, 520, 523, 899
Bsh1285I	CGRYCG	1 cut(s)	152
BshFI	GGCC	3 cut(s)	151, 394, 676
BsiEI	CGRYCG	1 cut(s)	152
BsiHKAI	GWGCWC	2 cut(s)	327, 818
BsiSI	CCGG	2 cut(s)	53, 152
BslFI	GGGAC	1 cut(s)	759
BsmAI	GTCTC	1 cut(s)	984
BsmFI	GGGAC	1 cut(s)	759
BsmI	GAATGC	1 cut(s)	1096
BsnI	GGCC	3 cut(s)	151, 394, 676
Bsp119I	TTCGAA	1 cut(s)	872
Bsp1286I	GDGCHC	3 cut(s)	327, 818, 903
Bsp143I	GATC	1 cut(s)	823
BspACI	CCGC	2 cut(s)	148, 1012
BspANI	GGCC	3 cut(s)	151, 394, 676
BspCNI	CTCAG	2 cut(s)	351, 853
BspHI	TCATGA	1 cut(s)	517
BspLI	GGNNCC	1 cut(s)	478
BspMAI	CTGCAG	1 cut(s)	541
BspT104I	TTCGAA	1 cut(s)	872
BspTI	CTTAAG	1 cut(s)	293
BsrFI	RCCGGY	1 cut(s)	151
BsrI	ACTGG	2 cut(s)	239, 405
BssAI	RCCGGY	1 cut(s)	151
BssECI	CCNNGG	2 cut(s)	480, 481
BssMI	GATC	1 cut(s)	823
Bst2UI	CCWGG	3 cut(s)	482, 699, 1093
Bst4CI	ACNGT	4 cut(s)	42, 499, 655, 718
BstAFI	CTTAAG	1 cut(s)	293
BstBAI	YACGTR	1 cut(s)	728
BstBI	TTCGAA	1 cut(s)	872
BstC8I	GCNNGC	1 cut(s)	489
BstDEI	CTNAG	3 cut(s)	338, 861, 990
BstF5I	GGATG	3 cut(s)	574, 934, 1063
BstKTI	GATC	1 cut(s)	826
BstMAI	GTCTC	1 cut(s)	984
BstMBI	GATC	1 cut(s)	823
BstMCI	CGRYCG	1 cut(s)	152
BstMWI	GCNNNNNNNGC	5 cut(s)	224, 331, 536, 542, 1093
BstNI	CCWGG	3 cut(s)	482, 699, 1093
BstNSI	RCATGY	2 cut(s)	491, 1052
BstSCI	CCNGG	4 cut(s)	52, 480, 697, 1091
BstSFI	CTRYAG	2 cut(s)	222, 537
BstSLI	GKGCMC	1 cut(s)	818
BstV1I	GCAGC	7 cut(s)	205, 343, 346, 517, 520, 523, 899
BstZI	CGGCCG	1 cut(s)	149
BsuI	GTATCC	1 cut(s)	30
BsuRI	GGCC	3 cut(s)	151, 394, 676
BtrI	CACGTC	1 cut(s)	749
BtsCI	GGATG	3 cut(s)	574, 934, 1063
BtsI	GCAGTG	1 cut(s)	248
BtsIMutI	CAGTG	1 cut(s)	248
Cac8I	GCNNGC	1 cut(s)	489
CaiI	CAGNNNCTG	1 cut(s)	920
CciI	TCATGA	1 cut(s)	517
Cfr10I	RCCGGY	1 cut(s)	151
Cfr13I	GGNCC	1 cut(s)	392
CseI	GACGC	1 cut(s)	503
Csp6I	GTAC	3 cut(s)	398, 651, 935
CviAII	CATG	7 cut(s)	488, 511, 518, 742, 833, 962, 1049
CviQI	GTAC	3 cut(s)	398, 651, 935
DdeI	CTNAG	3 cut(s)	338, 861, 990
DpnI	GATC	1 cut(s)	825
DpnII	GATC	1 cut(s)	823
DriI	GACNNNNNGTC	1 cut(s)	352
EaeI	YGGCCR	2 cut(s)	149, 674
EagI	CGGCCG	1 cut(s)	149
Eam1105I	GACNNNNNGTC	1 cut(s)	352
EclXI	CGGCCG	1 cut(s)	149
Eco24I	GRGCYC	1 cut(s)	903
Eco32I	GATATC	1 cut(s)	666
Eco52I	CGGCCG	1 cut(s)	149
Eco57I	CTGAAG	2 cut(s)	899, 1062
EcoO109I	RGGNCCY	1 cut(s)	392
EcoRII	CCWGG	3 cut(s)	480, 697, 1091
EcoRV	GATATC	1 cut(s)	666
EcoT38I	GRGCYC	1 cut(s)	903
FaeI	CATG	7 cut(s)	491, 514, 521, 745, 836, 965, 1052
FalI	AAGNNNNNCTT	4 cut(s)	276, 308, 311, 343
FaqI	GGGAC	1 cut(s)	759
FatI	CATG	7 cut(s)	487, 510, 517, 741, 832, 961, 1048
FblI	GTMKAC	1 cut(s)	200
Fnu4HI	GCNGC	8 cut(s)	149, 219, 332, 335, 531, 534, 537, 888
FokI	GGATG	3 cut(s)	561, 941, 1070
FriOI	GRGCYC	1 cut(s)	903
Fsp4HI	GCNGC	8 cut(s)	149, 219, 332, 335, 531, 534, 537, 888
FspBI	CTAG	1 cut(s)	422
GluI	GCNGC	8 cut(s)	149, 219, 332, 335, 531, 534, 537, 888
HaeIII	GGCC	3 cut(s)	151, 394, 676
HapII	CCGG	2 cut(s)	53, 152
HgaI	GACGC	1 cut(s)	503
Hin1II	CATG	7 cut(s)	491, 514, 521, 745, 836, 965, 1052
HinfI	GANTC	3 cut(s)	376, 626, 857
HpaII	CCGG	2 cut(s)	53, 152
HphI	GGTGA	1 cut(s)	671
Hpy166II	GTNNAC	2 cut(s)	201, 816
Hpy188I	TCNGA	2 cut(s)	341, 862
Hpy188III	TCNNGA	5 cut(s)	284, 518, 581, 904, 926
Hpy8I	GTNNAC	2 cut(s)	201, 816
HpyAV	CCTTC	2 cut(s)	103, 176
HpyCH4III	ACNGT	4 cut(s)	42, 499, 655, 718
HpyCH4IV	ACGT	2 cut(s)	727, 748
HpyF10VI	GCNNNNNNNGC	5 cut(s)	224, 331, 536, 542, 1093
HpyF3I	CTNAG	3 cut(s)	338, 861, 990
HpySE526I	ACGT	2 cut(s)	727, 748
Hsp92II	CATG	7 cut(s)	491, 514, 521, 745, 836, 965, 1052
Kzo9I	GATC	1 cut(s)	823
LmnI	GCTCC	2 cut(s)	322, 953
Lsp1109I	GCAGC	7 cut(s)	205, 343, 346, 517, 520, 523, 899
LweI	GCATC	7 cut(s)	39, 93, 277, 583, 826, 955, 1084
MaeI	CTAG	1 cut(s)	422
MaeII	ACGT	2 cut(s)	727, 748
MaeIII	GTNAC	6 cut(s)	274, 313, 463, 614, 706, 1031
MalI	GATC	1 cut(s)	825
MboI	GATC	1 cut(s)	823
MboII	GAAGA	2 cut(s)	191, 1055
MfeI	CAATTG	2 cut(s)	732, 819
MhlI	GDGCHC	3 cut(s)	327, 818, 903
MlsI	TGGCCA	1 cut(s)	676
MluCI	AATT	6 cut(s)	131, 231, 279, 732, 767, 819
MluNI	TGGCCA	1 cut(s)	676
MmeI	TCCRAC	1 cut(s)	798
MnlI	CCTC	6 cut(s)	151, 429, 467, 588, 1013, 1016
Mox20I	TGGCCA	1 cut(s)	676
MscI	TGGCCA	1 cut(s)	676
MseI	TTAA	5 cut(s)	294, 372, 432, 944, 986
MslI	CAYNNNNRTG	1 cut(s)	92
Msp20I	TGGCCA	1 cut(s)	676
MspA1I	CMGCKG	1 cut(s)	1014
MspCI	CTTAAG	1 cut(s)	293
MspI	CCGG	2 cut(s)	53, 152
MspR9I	CCNGG	4 cut(s)	54, 482, 699, 1093
MunI	CAATTG	2 cut(s)	732, 819
Mva1269I	GAATGC	1 cut(s)	1096
MvaI	CCWGG	3 cut(s)	482, 699, 1093
MwoI	GCNNNNNNNGC	5 cut(s)	224, 331, 536, 542, 1093
NciI	CCSGG	1 cut(s)	54
NdeII	GATC	1 cut(s)	823
NlaIII	CATG	7 cut(s)	491, 514, 521, 745, 836, 965, 1052
NlaIV	GGNNCC	1 cut(s)	478
NmuCI	GTSAC	2 cut(s)	274, 1031
NspI	RCATGY	2 cut(s)	491, 1052
NspV	TTCGAA	1 cut(s)	872
PaeI	GCATGC	1 cut(s)	491
PagI	TCATGA	1 cut(s)	517
PasI	CCCWGGG	1 cut(s)	481
PciI	ACATGT	1 cut(s)	1048
PcsI	WCGNNNNNNNCGW	1 cut(s)	651
PctI	GAATGC	1 cut(s)	1096
PfeI	GAWTC	3 cut(s)	376, 626, 857
PfoI	TCCNGGA	1 cut(s)	697
PkrI	GCNGC	8 cut(s)	150, 220, 333, 336, 532, 535, 538, 889
Ppu21I	YACGTR	1 cut(s)	728
PscI	ACATGT	1 cut(s)	1048
PshBI	ATTAAT	1 cut(s)	944
Psp6I	CCWGG	3 cut(s)	480, 697, 1091
PspGI	CCWGG	3 cut(s)	480, 697, 1091
PspN4I	GGNNCC	1 cut(s)	478
PspPI	GGNCC	1 cut(s)	392
PstI	CTGCAG	1 cut(s)	541
PstNI	CAGNNNCTG	1 cut(s)	920
RsaI	GTAC	3 cut(s)	399, 652, 936
RsaNI	GTAC	3 cut(s)	398, 651, 935
RseI	CAYNNNNRTG	1 cut(s)	92
SaqAI	TTAA	5 cut(s)	294, 372, 432, 944, 986
SatI	GCNGC	8 cut(s)	149, 219, 332, 335, 531, 534, 537, 888
Sau3AI	GATC	1 cut(s)	823
Sau96I	GGNCC	1 cut(s)	392
ScrFI	CCNGG	4 cut(s)	54, 482, 699, 1093
SduI	GDGCHC	3 cut(s)	327, 818, 903
SfaNI	GCATC	7 cut(s)	39, 93, 277, 583, 826, 955, 1084
SfcI	CTRYAG	2 cut(s)	222, 537
SfuI	TTCGAA	1 cut(s)	872
SmiMI	CAYNNNNRTG	1 cut(s)	92
SmlI	CTYRAG	3 cut(s)	293, 579, 902
SmoI	CTYRAG	3 cut(s)	293, 579, 902
SphI	GCATGC	1 cut(s)	491
Sse9I	AATT	6 cut(s)	131, 231, 279, 732, 767, 819
SsiI	CCGC	2 cut(s)	148, 1012
SspMI	CTAG	1 cut(s)	422
StyD4I	CCNGG	4 cut(s)	52, 480, 697, 1091
TaaI	ACNGT	4 cut(s)	42, 499, 655, 718
TaiI	ACGT	2 cut(s)	730, 751
TaqI	TCGA	3 cut(s)	668, 872, 1018
TasI	AATT	6 cut(s)	131, 231, 279, 732, 767, 819
TatI	WGTACW	1 cut(s)	397
TauI	GCSGC	1 cut(s)	151
TfiI	GAWTC	3 cut(s)	376, 626, 857
Tru1I	TTAA	5 cut(s)	294, 372, 432, 944, 986
Tru9I	TTAA	5 cut(s)	294, 372, 432, 944, 986
TscAI	CASTG	1 cut(s)	255
TseFI	GTSAC	2 cut(s)	274, 1031
TseI	GCWGC	7 cut(s)	218, 331, 334, 530, 533, 536, 887
Tsp45I	GTSAC	2 cut(s)	274, 1031
TspDTI	ATGAA	6 cut(s)	10, 17, 192, 196, 945, 1089
TspRI	CASTG	1 cut(s)	255
Vha464I	CTTAAG	1 cut(s)	293
VneI	GTGCAC	1 cut(s)	814
VspI	ATTAAT	1 cut(s)	944
XapI	RAATTY	1 cut(s)	231
XceI	RCATGY	2 cut(s)	491, 1052
XcmI	CCANNNNNNNNNTGG	2 cut(s)	357, 488
XmiI	GTMKAC	1 cut(s)	200
XspI	CTAG	1 cut(s)	422

Using CommOnly database (standard laboratory enzymes). Scanned on CDS sequence.