RchiOBHm_Chr4g0405701

serine threonine-protein kinase

Basic Information

Type: gene
Biological Identity
rosa_chinensis
4
Physical Location & Seq
Forward (+)
26374205 .. 26378155
3951 bp
Loading structure...
UTR
Exon/CDS
Intron
PRQ37717

Sequence Viewer

Length: 1584 bp
ATGCCTTTTGTGGGCCATTCTCCACTTACAGTCAGGAATGCTTGCGTTTCTGATTATTCTGGTGGGTGTTCTAGAAACACTACTTTGAATTGTGGGAACGAGACGAAAGAAAACCAGTTTATACAAATACCTAGCAAGCCATTGTCTGCATCTCAACAACTTCAGCAAGTAGGAAGTATTAAGGATTGTGAATCAATCTGTTTAAGAAAGTGTGATTGCTCTACTTATGGTTATGGAATTACTATTGGATGTGAAATCTGGACAGGAGATCTCTTGGCAGATGACAAAAGAAGAACTTTACACATCAGAGTACCGGCAGCTGAATCAAAGAATCCTAGTTTTAAGCGGTCCTTAGTGATTGGACTAGTCTCAGCAATAACAGGATTGCTTACAATAATTTGTAGCTATTTCATGTGGAAGAAAACTACAAGGAAGAGAAGGAAGCGTCAGAGGAAACATGTTGAGATCATAAGTAATGTTGGTAGAAAGAACGACACAGAACTACCAGTGTTTGACTTGAAGAGTATATTAGCTGCTACTGAAAACTTTTCTACAGCAAATAAACTGGGTGAGGGAGGATTTGGCCCTGTTTGTAAGGGTATTTTGCCTGGAAATCGAAAGGTGGCCATAAAAATGCTGTCAAAGAAGTCTGGTCAAGGGCATCTAGAGTTCATGAATGAGTTGAAGCTGATAGCCAAGCTCCAACATACCAATCTTGTTAGGCTTTTGGGTTGTTGTATTGAAGAGGAGGAAAGGATGTTGATCTATGAGTACATGCCTAATAGAAGTTTGGACAAATTTTTGTTTGATCCATCTAAAAGAACAAAATTGGATTGGGATAGATGCTTTAGAATTATAGAAGGCATTGGTCAAGGACTACTTTATATCCACAAGTACTCGAGATTGAAAATCATACATAGGGACCTAAAAGCAAGTAATATTTTGTTGGATGAAGCAATGAATCCTAAAATCTCAGACTTTGGAATGGCAAGGATATTCGTCATGAATCAAACTGAAGCAAATACCAATAGGGTTGTTGGGACATACGGTTTTATGTCACCTGAATATGCACGCTATGGCCATTTCTCTGAGAAATCGGATGTATTTAGTTTCGGAGTACTACTGTTGGAGATTATAAGTGGAAAGAAGAATGCTGCTCTCAATCGTTATGAAAATTCACTTACTCTTGCTGGATGGGTATGGAAACTGTGGAAAAAGGGTAGGGGAATGGAGGTAATTGACGAATCAGTAAGGGAAACATGTCGACCTCATGAAGCTTTAAGGTGTATCCATGTAGGTCTTCTGTGTGTTCAAGAAGCCCCAGCTGATCGACCAACAATGTTTTCTGTAGTTCATATGTTGCAAGCCGACAAAAGTACAACACCTCCACCCTCCAAAGAACCTGCATTTTCAACACACAAGAATTCCAGTGCCGCTAGTTCTTCTTCTCAAACATCTAGCATTTTTTCCACCAATCAATTGACCGTTCGTATATATCATAGAAGGTCGATAGAGGTTCAATGTCAGTCGATGATAAATTTTCCTACAATATATACCTTGGCCTTTAATCAATTAAATGCTTGA

Protein Analysis

527

Amino Acids

59.38

Weight (kDa)

9.38

Isoelectric Point (pI)

44.7

Instability Index
Protein Domains (Pfam)
Domain Name Pfam ID Position E-value Description
PAN_2 PF08276 34 - 92 8.4e-09 PAN-like domain
Pkinase PF00069 184 - 390 2.9e-46 Protein kinase domain
PK_Tyr_Ser-Thr PF07714 185 - 453 3.5e-51 Protein tyrosine and serine/threonine kinase
Hydrophobicity Profile (Kyte-Doolittle)
AI Structure Prediction Report
Calculating structure properties...

Genomic Context

Gene Family Tree

Species Filter
Loading...
Style Settings
Image
Tree File
Tip: Beautify your tree with professional tools

Download the Full Tree (.nwk) file, then upload it to any of the following tools to customize colors, fonts, annotations, clades, and branch support.

Publication-ready

Orthologous Genes (Group: OG0000564)

Species Orthologous Gene IDs
fragaria_vesca FvH4_6g44064 FvH4_6g44064 FvH4_6g44100 FvH4_6g44101 FvH4_6g51830
malus_domestica MD09G1094400.v1.1 MD09G1094800.v1.1 MD09G1095200.v1.1 MD09G1095900.v1.1 MD09G1096000.v1.1 MD09G1096200.v1.1 MD17G1083900.v1.1 MD17G1088500.v1.1
prunus_persica Prupe.3G224900_v2.0.a1 Prupe.3G224900_v2.0.a1 Prupe.3G224900_v2.0.a1 Prupe.3G224900_v2.0.a1 Prupe.3G224900_v2.0.a1 Prupe.3G224900_v2.0.a1 Prupe.3G225100_v2.0.a1 Prupe.3G225100_v2.0.a1 Prupe.3G225100_v2.0.a1 Prupe.3G225200_v2.0.a1 Prupe.3G225200_v2.0.a1
pyrus_communis pycom09g01920 pycom09g01930 pycom09g01940 pycom09g01950 pycom09g02000 pycom09g02050 pycom09g02070 pycom17g08540
rosa_chinensis RchiOBHm_Chr1g0318601 RchiOBHm_Chr1g0318761 RchiOBHm_Chr1g0342541 RchiOBHm_Chr2g0160651 RchiOBHm_Chr2g0160831 RchiOBHm_Chr4g0405661 RchiOBHm_Chr4g0405691 RchiOBHm_Chr4g0405701 RchiOBHm_Chr7g0238591
rosa_laevigata RLG00000002292 RLG00000008826 RLG00000008828 RLG00000008829 RLG00000021243 RLG00000021245 RLG00000030522
rosa_multiflora Rmu_co8432573.1_g000001 Rmu_sc0001200.1_g000041 Rmu_sc0001200.1_g000043 Rmu_sc0002545.1_g000017 Rmu_sc0002773.1_g000017 Rmu_sc0012550.1_g000008
rosa_roxburghii Rroxscaffold_2G00089500 Rroxscaffold_5G00350130 Rroxscaffold_5G00350180
rosa_rugosa Rorug01G0022300 Rorug04G0061600
rosa_samantha Rh1AG035300 Rh1CG034300 Rh1DG036100 Rh1DG149100 Rh2AG550900 Rh2BG564200 Rh2BG565500 Rh2CG535300 Rh2DG574400 Rh4AG133100 Rh4AG133200 Rh4AG133300 Rh4BG128200 Rh4CG141000 Rh4DG127700 Rh4DG127900
rosa_wichuraiana Rw2G045540 Rw2G045650 Rw4G010790 Rw4G010800 Rw4G010810 Rw4G010830 Rw7G027620 Rw7G027630

Restriction Enzyme Sites

1 / 10
Enzyme Recognition Site Cut Count Positions (bp)
AanI TTATAA 1 cut(s) 1136
Acc36I ACCTGC 1 cut(s) 1411
AccI GTMKAC 1 cut(s) 1264
AciI CCGC 2 cut(s) 346, 1434
AclWI GGATC 1 cut(s) 803
AcoI YGGCCR 2 cut(s) 624, 1078
AcsI RAATTY 4 cut(s) 797, 1174, 1423, 1537
AcuI CTGAAG 2 cut(s) 146, 1035
AfaI GTAC 5 cut(s) 312, 773, 896, 1119, 1378
AfiI CCNNNNNNNGG 1 cut(s) 11
AflIII ACRYGT 2 cut(s) 457, 1259
AgsI TTSAA 8 cut(s) 88, 520, 685, 743, 907, 1313, 1413, 1520
AhlI ACTAGT 1 cut(s) 364
AjnI CCWGG 1 cut(s) 607
AjuI GAANNNNNNNTTGG 2 cut(s) 1327, 1359
AluBI AGCT 7 cut(s) 320, 405, 533, 688, 700, 1277, 1325
AluI AGCT 7 cut(s) 320, 405, 533, 688, 700, 1277, 1325
Alw26I GTCTC 2 cut(s) 95, 373
AlwI GGATC 1 cut(s) 803
Ama87I CYCGRG 1 cut(s) 898
AoxI GGCC 5 cut(s) 13, 583, 624, 1078, 1560
ApeKI GCWGC 3 cut(s) 317, 533, 1154
ApoI RAATTY 4 cut(s) 797, 1174, 1423, 1537
AspS9I GGNCC 4 cut(s) 13, 348, 584, 922
AsuHPI GGTGA 2 cut(s) 581, 1050
AvaI CYCGRG 1 cut(s) 898
AvaII GGWCC 2 cut(s) 348, 922
BalI TGGCCA 2 cut(s) 626, 1080
BbsI GAAGAC 1 cut(s) 1292
BbvI GCAGC 3 cut(s) 329, 520, 1141
BccI CCATC 2 cut(s) 820, 1188
BciT130I CCWGG 1 cut(s) 609
BciVI GTATCC 1 cut(s) 1298
BcoDI GTCTC 2 cut(s) 95, 373
BcuI ACTAGT 1 cut(s) 364
BfaI CTAG 7 cut(s) 72, 132, 336, 365, 665, 1437, 1458
BfmI CTRYAG 2 cut(s) 552, 1347
BfuAI ACCTGC 1 cut(s) 1411
BfuI GTATCC 1 cut(s) 1298
BglII AGATCT 1 cut(s) 268
BisI GCNGC 4 cut(s) 318, 534, 1155, 1434
BlsI GCNGC 4 cut(s) 319, 535, 1156, 1435
BmcAI AGTACT 2 cut(s) 896, 1119
Bme1390I CCNGG 1 cut(s) 609
Bme18I GGWCC 2 cut(s) 348, 922
BmeT110I CYCGRG 1 cut(s) 898
BmgT120I GGNCC 4 cut(s) 13, 348, 584, 922
BmiI GGNNCC 1 cut(s) 923
BmrFI CCNGG 1 cut(s) 609
BmrI ACTGGG 1 cut(s) 575
BmsI GCATC 3 cut(s) 158, 670, 833
BmuI ACTGGG 1 cut(s) 575
BpiI GAAGAC 1 cut(s) 1292
BsaBI GATNNNNATC 1 cut(s) 761
BsaJI CCNNGG 1 cut(s) 1557
BsaXI ACNNNNNCTCC 2 cut(s) 1369, 1399
Bsc4I CCNNNNNNNGG 1 cut(s) 11
Bse118I RCCGGY 1 cut(s) 313
Bse1I ACTGG 4 cut(s) 115, 506, 570, 1428
Bse3DI GCAATG 1 cut(s) 963
Bse8I GATNNNNATC 1 cut(s) 761
BseBI CCWGG 1 cut(s) 609
BseDI CCNNGG 1 cut(s) 1557
BseGI GGATG 5 cut(s) 254, 762, 955, 1105, 1199
BseJI GATNNNNATC 1 cut(s) 761
BseLI CCNNNNNNNGG 1 cut(s) 11
BseMI GCAATG 1 cut(s) 963
BseMII CTCAG 3 cut(s) 384, 987, 1080
BseNI ACTGG 4 cut(s) 115, 506, 570, 1428
BseRI GAGGAG 1 cut(s) 761
BseXI GCAGC 3 cut(s) 329, 520, 1141
BseYI CCCAGC 1 cut(s) 1321
BshFI GGCC 5 cut(s) 15, 585, 626, 1080, 1562
BsiHKCI CYCGRG 1 cut(s) 898
BsiSI CCGG 1 cut(s) 314
BslFI GGGAC 2 cut(s) 935, 1054
BslI CCNNNNNNNGG 1 cut(s) 11
BsmAI GTCTC 2 cut(s) 95, 373
BsmBI CGTCTC 1 cut(s) 95
BsmFI GGGAC 2 cut(s) 935, 1054
BsmI GAATGC 2 cut(s) 43, 1156
BsnI GGCC 5 cut(s) 15, 585, 626, 1080, 1562
BsoBI CYCGRG 1 cut(s) 898
Bsp143I GATC 5 cut(s) 268, 465, 762, 808, 1327
BspACI CCGC 2 cut(s) 346, 1434
BspANI GGCC 5 cut(s) 15, 585, 626, 1080, 1562
BspCNI CTCAG 3 cut(s) 383, 986, 1081
BspHI TCATGA 3 cut(s) 672, 1002, 1270
BspLI GGNNCC 1 cut(s) 923
BspMI ACCTGC 1 cut(s) 1411
BspPI GGATC 1 cut(s) 803
BsrDI GCAATG 1 cut(s) 963
BsrFI RCCGGY 1 cut(s) 313
BsrI ACTGG 4 cut(s) 115, 506, 570, 1428
BssAI RCCGGY 1 cut(s) 313
BssECI CCNNGG 1 cut(s) 1557
BssMI GATC 5 cut(s) 268, 465, 762, 808, 1327
BssT1I CCWWGG 1 cut(s) 1557
Bst2UI CCWGG 1 cut(s) 609
Bst4CI ACNGT 5 cut(s) 31, 1049, 1125, 1209, 1486
Bst6I CTCTTC 3 cut(s) 428, 515, 738
BstC8I GCNNGC 4 cut(s) 43, 137, 1072, 1365
BstDEI CTNAG 4 cut(s) 352, 370, 973, 1089
BstF5I GGATG 5 cut(s) 254, 762, 955, 1105, 1199
BstKTI GATC 5 cut(s) 271, 468, 765, 811, 1330
BstMAI GTCTC 2 cut(s) 95, 373
BstMBI GATC 5 cut(s) 268, 465, 762, 808, 1327
BstNI CCWGG 1 cut(s) 609
BstNSI RCATGY 3 cut(s) 461, 778, 1263
BstSCI CCNGG 1 cut(s) 607
BstSFI CTRYAG 2 cut(s) 552, 1347
BstV1I GCAGC 3 cut(s) 329, 520, 1141
BstV2I GAAGAC 1 cut(s) 1292
BstX2I RGATCY 1 cut(s) 268
BstYI RGATCY 1 cut(s) 268
BsuI GTATCC 1 cut(s) 1298
BsuRI GGCC 5 cut(s) 15, 585, 626, 1080, 1562
BtsCI GGATG 5 cut(s) 254, 762, 955, 1105, 1199
BtsIMutI CAGTG 2 cut(s) 513, 1435
BveI ACCTGC 1 cut(s) 1411
Cac8I GCNNGC 4 cut(s) 43, 137, 1072, 1365
CciI TCATGA 3 cut(s) 672, 1002, 1270
Cfr10I RCCGGY 1 cut(s) 313
Cfr13I GGNCC 4 cut(s) 13, 348, 584, 922
CseI GACGC 1 cut(s) 434
Csp6I GTAC 5 cut(s) 311, 772, 895, 1118, 1377
CviAII CATG 8 cut(s) 412, 458, 673, 775, 1003, 1260, 1271, 1292
CviQI GTAC 5 cut(s) 311, 772, 895, 1118, 1377
DdeI CTNAG 4 cut(s) 352, 370, 973, 1089
DpnI GATC 5 cut(s) 270, 467, 764, 810, 1329
DpnII GATC 5 cut(s) 268, 465, 762, 808, 1327
EaeI YGGCCR 2 cut(s) 624, 1078
Eam1104I CTCTTC 3 cut(s) 428, 515, 738
EarI CTCTTC 3 cut(s) 428, 515, 738
Eco130I CCWWGG 1 cut(s) 1557
Eco47I GGWCC 2 cut(s) 348, 922
Eco57I CTGAAG 2 cut(s) 146, 1035
Eco88I CYCGRG 1 cut(s) 898
EcoO109I RGGNCCY 1 cut(s) 922
EcoRI GAATTC 1 cut(s) 1423
EcoRII CCWGG 1 cut(s) 607
EcoT14I CCWWGG 1 cut(s) 1557
ErhI CCWWGG 1 cut(s) 1557
Esp3I CGTCTC 1 cut(s) 95
FaeI CATG 8 cut(s) 415, 461, 676, 778, 1006, 1263, 1274, 1295
FalI AAGNNNNNCTT 6 cut(s) 280, 312, 335, 367, 864, 896
FaqI GGGAC 2 cut(s) 935, 1054
FatI CATG 8 cut(s) 411, 457, 672, 774, 1002, 1259, 1270, 1291
FauNDI CATATG 1 cut(s) 1356
FblI GTMKAC 1 cut(s) 1264
Fnu4HI GCNGC 4 cut(s) 318, 534, 1155, 1434
FokI GGATG 5 cut(s) 261, 769, 962, 1112, 1206
Fsp4HI GCNGC 4 cut(s) 318, 534, 1155, 1434
FspBI CTAG 7 cut(s) 72, 132, 336, 365, 665, 1437, 1458
GluI GCNGC 4 cut(s) 318, 534, 1155, 1434
GsaI CCCAGC 1 cut(s) 1325
HaeIII GGCC 5 cut(s) 15, 585, 626, 1080, 1562
HapII CCGG 1 cut(s) 314
HgaI GACGC 1 cut(s) 434
Hin1II CATG 8 cut(s) 415, 461, 676, 778, 1006, 1263, 1274, 1295
HincII GTYRAC 1 cut(s) 1265
HindII GTYRAC 1 cut(s) 1265
HindIII AAGCTT 1 cut(s) 1275
HinfI GANTC 6 cut(s) 191, 323, 331, 961, 1006, 1244
HpaII CCGG 1 cut(s) 314
HphI GGTGA 2 cut(s) 581, 1050
Hpy166II GTNNAC 1 cut(s) 1265
Hpy188I TCNGA 7 cut(s) 52, 308, 450, 976, 1090, 1099, 1115
Hpy188III TCNNGA 9 cut(s) 34, 72, 259, 665, 673, 900, 1003, 1271, 1313
Hpy8I GTNNAC 1 cut(s) 1265
HpyAV CCTTC 3 cut(s) 432, 854, 1497
HpyCH4III ACNGT 5 cut(s) 31, 1049, 1125, 1209, 1486
HpyCH4V TGCA 4 cut(s) 149, 1070, 1363, 1406
HpyF3I CTNAG 4 cut(s) 352, 370, 973, 1089
Hsp92II CATG 8 cut(s) 415, 461, 676, 778, 1006, 1263, 1274, 1295
Kzo9I GATC 5 cut(s) 268, 465, 762, 808, 1327
LmnI GCTCC 1 cut(s) 705
Lsp1109I GCAGC 3 cut(s) 329, 520, 1141
LweI GCATC 3 cut(s) 158, 670, 833
MaeI CTAG 7 cut(s) 72, 132, 336, 365, 665, 1437, 1458
MaeIII GTNAC 1 cut(s) 1056
MalI GATC 5 cut(s) 270, 467, 764, 810, 1329
MboI GATC 5 cut(s) 268, 465, 762, 808, 1327
MboII GAAGA 9 cut(s) 303, 430, 445, 532, 755, 1159, 1292, 1434, 1437
MfeI CAATTG 1 cut(s) 1478
MflI RGATCY 1 cut(s) 268
MlsI TGGCCA 2 cut(s) 626, 1080
MluNI TGGCCA 2 cut(s) 626, 1080
MmeI TCCRAC 3 cut(s) 727, 927, 1107
Mox20I TGGCCA 2 cut(s) 626, 1080
MscI TGGCCA 2 cut(s) 626, 1080
MseI TTAA 6 cut(s) 180, 203, 342, 1280, 1566, 1574
MslI CAYNNNNRTG 1 cut(s) 632
Msp20I TGGCCA 2 cut(s) 626, 1080
MspA1I CMGCKG 2 cut(s) 320, 1325
MspI CCGG 1 cut(s) 314
MspR9I CCNGG 1 cut(s) 609
MunI CAATTG 1 cut(s) 1478
Mva1269I GAATGC 2 cut(s) 43, 1156
MvaI CCWGG 1 cut(s) 609
NdeI CATATG 1 cut(s) 1356
NdeII GATC 5 cut(s) 268, 465, 762, 808, 1327
NlaIII CATG 8 cut(s) 415, 461, 676, 778, 1006, 1263, 1274, 1295
NlaIV GGNNCC 1 cut(s) 923
NmuCI GTSAC 1 cut(s) 1056
NspI RCATGY 3 cut(s) 461, 778, 1263
PaeR7I CTCGAG 1 cut(s) 898
PagI TCATGA 3 cut(s) 672, 1002, 1270
PciI ACATGT 2 cut(s) 457, 1259
PctI GAATGC 2 cut(s) 43, 1156
PfeI GAWTC 6 cut(s) 191, 323, 331, 961, 1006, 1244
PkrI GCNGC 4 cut(s) 319, 535, 1156, 1435
PpuMI RGGWCCY 1 cut(s) 922
PscI ACATGT 2 cut(s) 457, 1259
PsiI TTATAA 1 cut(s) 1136
Psp5II RGGWCCY 1 cut(s) 922
Psp6I CCWGG 1 cut(s) 607
PspFI CCCAGC 1 cut(s) 1321
PspGI CCWGG 1 cut(s) 607
PspN4I GGNNCC 1 cut(s) 923
PspPI GGNCC 4 cut(s) 13, 348, 584, 922
PspPPI RGGWCCY 1 cut(s) 922
PsuI RGATCY 1 cut(s) 268
PvuII CAGCTG 2 cut(s) 320, 1325
RsaI GTAC 5 cut(s) 312, 773, 896, 1119, 1378
RsaNI GTAC 5 cut(s) 311, 772, 895, 1118, 1377
RseI CAYNNNNRTG 1 cut(s) 632
SalI GTCGAC 1 cut(s) 1263
SaqAI TTAA 6 cut(s) 180, 203, 342, 1280, 1566, 1574
SatI GCNGC 4 cut(s) 318, 534, 1155, 1434
Sau3AI GATC 5 cut(s) 268, 465, 762, 808, 1327
Sau96I GGNCC 4 cut(s) 13, 348, 584, 922
ScaI AGTACT 2 cut(s) 896, 1119
ScrFI CCNGG 1 cut(s) 609
SfaNI GCATC 3 cut(s) 158, 670, 833
SfcI CTRYAG 2 cut(s) 552, 1347
Sfr274I CTCGAG 1 cut(s) 898
SinI GGWCC 2 cut(s) 348, 922
SlaI CTCGAG 1 cut(s) 898
SmiMI CAYNNNNRTG 1 cut(s) 632
SmlI CTYRAG 1 cut(s) 898
SmoI CTYRAG 1 cut(s) 898
SpeI ACTAGT 1 cut(s) 364
SsiI CCGC 2 cut(s) 346, 1434
SspI AATATT 1 cut(s) 940
SspMI CTAG 7 cut(s) 72, 132, 336, 365, 665, 1437, 1458
StyD4I CCNGG 1 cut(s) 607
StyI CCWWGG 1 cut(s) 1557
TaaI ACNGT 5 cut(s) 31, 1049, 1125, 1209, 1486
TaqI TCGA 6 cut(s) 616, 899, 1264, 1330, 1508, 1529
TatI WGTACW 4 cut(s) 771, 894, 1117, 1376
TauI GCSGC 1 cut(s) 1436
TfiI GAWTC 6 cut(s) 191, 323, 331, 961, 1006, 1244
Tru1I TTAA 6 cut(s) 180, 203, 342, 1280, 1566, 1574
Tru9I TTAA 6 cut(s) 180, 203, 342, 1280, 1566, 1574
TscAI CASTG 2 cut(s) 513, 1435
TseFI GTSAC 1 cut(s) 1056
TseI GCWGC 3 cut(s) 317, 533, 1154
Tsp45I GTSAC 1 cut(s) 1056
TspDTI ATGAA 9 cut(s) 400, 661, 689, 966, 974, 1019, 1185, 1287, 1343
TspRI CASTG 2 cut(s) 513, 1435
VpaK11BI GGWCC 2 cut(s) 348, 922
XapI RAATTY 4 cut(s) 797, 1174, 1423, 1537
XbaI TCTAGA 2 cut(s) 71, 664
XceI RCATGY 3 cut(s) 461, 778, 1263
XhoI CTCGAG 1 cut(s) 898
XmiI GTMKAC 1 cut(s) 1264
XspI CTAG 7 cut(s) 72, 132, 336, 365, 665, 1437, 1458
ZrmI AGTACT 2 cut(s) 896, 1119
Using CommOnly database (standard laboratory enzymes). Scanned on CDS sequence.